Timing of Lateglacial vegetation dynamics and respective palaeoenvironmental conditions in southern Estonia: evidence from the sediment record of Lake Nakri

This paper investigates a detailed well‐dated Lateglacial floristic colonisation in the eastern Baltic area, ca. 14 000–9000 cal. a BP, using palynological, macrofossil, loss‐on‐ignition, and ¹⁴C data. During 14 000–13 400 cal. a BP, primarily treeless pioneer tundra vegetation existed. Tree birch (Betula sect. Albae) macro‐remains and a high tree pollen accumulation rate indicate the presence of forest‐tundra with birch and possibly pine (Pinus sylvestris L.) trees during 13 400–12 850 cal. a BP. Palaeobotanical data indicate that the colonisation and development of forested areas were very rapid, arising within a period of less than 50 years. Thus far, there are no indications of conifer macrofossils in Estonia to support the presence of coniferous forests in the Lateglacial period. Signs of Greenland Interstadial 1b cooling during 13 100 cal. a BP are distinguishable. Biostratigraphic evidence indicates that the vegetation was again mostly treeless tundra during the final colder episode of the Lateglacial period associated with Greenland Stadial 1, approximately 12 850–11 650 cal. a BP. This was followed by onset of the Holocene vegetation, with the expansion of boreal forests, in response to rapid climatic warming. Copyright © 2011 John Wiley & Sons, Ltd.     

Holocene floristic diversity and richness in northeast Norway revealed by sedimentary ancient DNA (sedaDNA) and pollen

We present a Holocene record of floristic diversity and environmental change for the central Varanger Peninsula, Finnmark, based on ancient DNA extracted from the sediments of a small lake (sedaDNA). The record covers the period c. 10 700 to 3300 cal. a BP and is complemented by pollen data. Measures of species richness, sample evenness and beta diversity were calculated based on sedaDNA sampling intervals and 1000‐year time windows. We identified 101 vascular plant and 17 bryophyte taxa, a high proportion (86%) of which are still growing within the region today. The high species richness (>60 taxa) observed in the Early Holocene, including representatives from all important plant functional groups, shows that modern shrub‐tundra communities, and much of their species complement, were in place as early as c. 10 700 cal. a BP. We infer that postglacial colonization of the area occurred prior to the full Holocene, during the Pleistocene‐Holocene transition, Younger Dryas stadial or earlier. Abundant DNA of the extra‐limital aquatic plant Callitriche hermaphroditica suggests it expanded its range northward between c. 10 200 and 9600 cal. a BP, when summers were warmer than present. High values of Pinus DNA occur throughout the record, but we cannot say with certainty if they represent prior local presence; however, pollen influx values >500 grains cm^?2 a^?1 between c. 8000 and 7300 cal. a BP strongly suggest the presence of pine woodland during this period. As the site lies beyond the modern tree limit of pine, it is likely that this expansion also reflects a response to warmer Early Holocene summers.     

Climate and environment in southwest Sweden 15.5–11.3 cal. ka BP

Lake sedimentary records that allow documentation of the distinct climatic and environmental shifts during the early part of the Last Termination are scarce for northern Europe. This multi‐proxy study of the sediments of Atteköpsmosse, southwest Sweden, therefore fills an important gap and provides detailed information regarding past hydroclimatic conditions and local environmental responses to climatic shifts. Lake infilling started c. 15.5 cal. ka BP, but low aquatic productivity, cold summer lake water temperatures, unstable catchments, and scarce herb and shrub vegetation prevailed until c. 14.7–14.5 cal. ka BP. Inflow of warmer air masses and higher July air temperatures favoured a rise in aquatic productivity and lake water summer temperatures, and the establishment of a diverse herb, shrub and dwarf shrub vegetation, which also included tree birch c. 14.5 cal. ka BP. Freshening of the moisture source region c. 13.7–13.6 cal. ka BP does not seem to have had a large impact on the ancient lake and its catchment, as lake aquatic productivity increased further and lake water summer temperatures and minimum mean July air temperatures remained around 12–14 °C. In contrast, further freshening of the moisture source region c. 13 cal. ka BP triggered a decrease in lake productivity, drier conditions and lower lake water summer temperatures. Macroscopic finds of tree Betula and Pinus sylvestris at 13–12.8 cal. ka BP demonstrate the presence of these trees in the lake's catchment. The transition into the Holocene (11.6–11.5 cal. ka BP) is marked by a change in chironomid assemblages and by a rise in lake water summer temperatures and aquatic productivity. These changes were followed by the re‐establishment of a diverse aquatic and terrestrial vegetation, including tree birch and Pinus sylvestris at 11.4 cal. ka BP.     

Climatic influence on peatland formation and lateral expansion in sub‐arctic Fennoscandia

Weckström, J., Seppä, H. & Korhola, A. 2010: Climatic influence on peatland formation and lateral expansion in sub‐arctic Fennoscandia. Boreas, Vol. 39, pp. 761–769. 10.1111/j.1502‐3885.2010.00168.x. ISSN 0300‐9843. The initiation and lateral expansion patterns of five small sub‐arctic peatlands in the Fennoscandian tree‐line region were studied by 21 accelerator mass spectrometry (AMS) ¹⁴C‐dated basal‐peat samples representing three to six dates per site. The radiocarbon dates were converted to calendar years and are based on the median probability. When combined with earlier basal‐peat dates from the region, four distinctive periods can be observed in the cumulative record of the dates. The early Holocene, from c. 10 000 to 8000 cal. yr BP, was characterized by the fast initiation and rapid expansion of peatlands, whereas at 8000–4000 cal. yr BP lateral expansion was modest. The most intensive period of peatland expansion occurred at the beginning of the late Holocene at c. 4000 to 3000 cal. yr BP, after which it slowed down towards the present. All these periods are in rough agreement with the main Holocene climatic periods in the area, namely the relatively warm and moist early Holocene, the warm and dry Holocene thermal maximum (HTM) at 8000–4000 cal. yr BP, and the start of the cooler and moister trend (neoglacial cooling) from c. 4000 cal. yr BP to the present, indicating a broad‐scale climatic control on the lateral growth of sub‐arctic peatlands in Fennoscandia. In order to study the lateral expansion of peatlands and to evaluate their Holocene succession patterns, more studies based on multiple dates from the same peatland are needed.     

Midge‐inferred Holocene effective moisture fluctuations in a subarctic lake,northern Lapland

Luoto, T. P. & Sarmaja‐Korjonen, K. 2011: Midge‐inferred Holocene effective moisture fluctuations in a subarctic lake, northern Lapland. Boreas, 10.1111/j.1502‐3885.2011.00217.x. ISSN 0300‐9483. We examined fossil midge (Diptera: Chironomidae) assemblages from Lake Várddoaijávri, northern Finland to track Holocene effective moisture variability. Application of a midge‐based water‐depth calibration model showed that the early Holocene was characterized by a high water level compared with the Holocene average, but the inferred values decreased at c. 8000 cal. a BP and increased again towards c. 6000 cal. a BP. The inferred water level decreased at c. 5500 cal. a BP, but increased again towards c. 4000 cal. a BP. Between 4000 and 3000 cal. a BP the lake experienced two rapid events of lower water level. A relatively high water level detected at c. 3000 cal. a BP was followed by a lowering towards c. 2000 cal. a BP. The time period from c. 2000 cal. a BP onwards was characterized by a general rise in lake level towards the present. Overall, the present reconstruction shows a close correspondence in its trends to previous lake‐level records in the region. Two common core taxa, Paratanytarsus and Corynocera ambigua, did not correlate significantly with water depth in the calibration data, creating a potential error source for the present lake‐level reconstruction. However, statistical analysis showed a clear community response to long‐term lake‐level changes, and therefore the major trends in Holocene effective moisture patterns were revealed. The present palaeoclimatic information can also serve as valuable background data when assessing the effects of the present climate change.     

Vegetation responses to climatic variability in the Swiss Southern Alps during the Misox event at the early–mid Holocene transition

Sedimentary pollen, charcoal and plant macrofossil analyses with high resolution and precision suggest a strong shift in vegetation composition during the early to mid‐Holocene transition in the upper mountain belt. At Piano mire (1439 m above sea level (a.s.l.), Ticino, Switzerland) forests were dominated by Abies alba during the early Holocene (prior to ca. 8000 cal. a BP). Abrupt collapses of A. alba at ca. 7800–7400 cal. a BP enabled the expansion of the light‐demanding pioneer Betula. Afterwards A. alba populations regained their previous abundance in the forests. Within the dating uncertainties of our record we assume that a unique combination of wet and cold years between 8400 and 7500 cal. a BP led to repeated lethal disadvantages for Abies. Our record of Abies oscillations is in good biostratigraphic agreement with the record that has been used to define the Misox cold event (Pian di Signano, 1540 m a.s.l.), which has been previously correlated with the 8200 cal. a BP event. Given the age estimates of the Abies collapses in our well‐dated record, our results suggest that additional efforts are needed to understand the linkage between the Misox and the 8200 cal. a BP event. They imply a high sensitivity of mountain vegetation far below the tree line (～800 m) to Holocene climatic changes of about 2°C in annual air temperature. Copyright © 2010 John Wiley & Sons, Ltd.     

Vegetation and climate changes in northwestern Russia during the Lateglacial and Holocene inferred from the Lake Ladoga pollen record

The new pollen record from the upper 12.75 m of a sediment core obtained in Lake Ladoga documents regional vegetation and climate changes in northwestern Russia over the last 13.9 cal. ka. The Lateglacial chronostratigraphy is based on varve chronology, while the Holocene stratigraphy is based on AMS ¹⁴C and OSL dates, supported by comparison with regional pollen records. During the Lateglacial (c. 13.9–11.2 cal. ka BP), the Lake Ladoga region experienced several climatic fluctuations as reflected in vegetation changes. Shrub and grass communities dominated between c. 13.9 and 13.2 cal. ka BP. The increase in Picea pollen at c. 13.2 cal. ka BP probably reflects the appearance of spruce in the southern Ladoga region at the beginning of the Allerød interstadial. After c. 12.6 cal. ka BP, the Younger Dryas cooling caused a significant decrease in spruce and increase in Artemisia with other herbs, indicative of tundra‐ and steppe‐like vegetation. A sharp transition from tundra‐steppe habitats to sparse birch forests characterizes the onset of Holocene warming c. 11.2 cal. ka BP. Pine forests dominated in the region from c. 9.0 to 8.1 cal. ka BP. The most favourable climatic conditions for deciduous broad‐leaved taxa existed between c. 8.1 and 5.5 cal. ka BP. Alder experiences an abrupt increase in the local vegetation c. 7.8 cal. ka BP. The decrease in tree pollen taxa (especially Picea) and the increase in herbs (mainly Poaceae) probably reflect human activity during the last 2.2 cal. ka. Pine forests have dominated the region since that time. Secale and other Cerealia pollen as well as ruderal herbs are permanently recorded since c. 0.8 cal. ka BP.     

Holocene vegetation change in northernmost Fennoscandia and the impact on prehistoric foragers 12 000–2000 cal. a BP – A review

While single pollen records are widely used in reconstructing the environment for nearby prehistoric settlements, they are less helpful when addressing large‐scale issues of variation in human settlement patterns. In order to assess the impact of vegetation change on regional prehistoric settlement and subsistence patterns in an ecotone sensitive area, we inferred the general change in main vegetation types based on palaeobotanical investigations from across northernmost Fennoscandia. Tundra vegetation was predominant during the Lateglacial and earliest parts of the Holocene. Maritime birch forests rich in ferns started to expand c. 11 000 cal. a BP and became dominant from 10 000 cal. a BP. Pine expanded from the NE of the investigation area and pine‐birch forest dominated in the inland around 8000 cal. a BP. A gradual degeneration of forest towards more open birch woodland started c. 6000 cal. a BP with the most marked change around 3500 cal. a BP. Along the northern outer coast, this eventually led to open heathland. Comparison with the archaeological setting suggests a general correlation between low forest cover and extensive mobility patterns, while widespread and varied forest cover appear to have led to a more sedentary way of life. The background for this is arguably that the forested landscapes hosted a larger diversity of resources within a shorter foraging distance, while areas and periods with low forest cover required longer travels to obtain the desired prey and materials.     

The Lateglacial and Holocene in Central Europe: a multi‐proxy environmental record from the Bohemian Forest,Czech Republic

The Hercynian mountain ranges were islands of mountain glaciation and alpine tundra in a Central European ice‐free corridor during the Late Pleistocene. Today they are notable areas of glacial landforms, alpine‐forest free areas, peatlands and woodlands. However, our knowledge of the Lateglacial and early Holocene environmental changes in this region is limited. We present a new multi‐proxy reconstruction of a mid‐altitude environment in the Bohemian Forest spanning this period. A core (5.2 m length) in the ?erné Lake cirque (1028 m a.s.l.) was subjected to lithological, geochemical, pollen and macrofossil analysis supplemented by two optically stimulated luminescence (OSL) and 10 AMS radiocarbon dates. We determined the impact of regional and supraregional climate changes on the environment. The two most significant changes in sedimentation during the Lateglacial (17.6 and 15.8–15.5 cal. ka BP) were synchronous with regional glacial chronostratigraphy. Unlike Central European mountain ranges, in the Bohemian Forest the Younger Dryas was not coincident with glacier re‐advance, but was a dry, cold episode with low lake levels, which prevailed until the early Preboreal. Plant macrofossils indicate local establishment of Betula nana and Betula pendula/pubescens at 15.4–13.4 cal. ka BP. Comparison with Holocene records from Central Europe shows a similar immigration history of vegetation at mid and higher altitudes. The tree line exceeded an altitude of ~1000 m a.s.l. around 10.5 cal. ka BP and coincided with rapid geochemical changes in the sediment. The 8.2 ka BP event did not have any response in the sedimentary record, but corresponded to stabilization of the Picea abies population and expansion of Fagus. Fagus colonized the Bohemian Forest earlier than other Hercynian mid‐mountains, but never predominated in the composition of the forest at higher elevations. Abies alba was the last tree species that immigrated to the study area.     

Holocene vegetation and fire dynamics in the supra‐mediterranean belt of the Nebrodi Mountains (Sicily,Italy)

High‐resolution pollen, macrofossil and charcoal data, combined with accelerator mass spectrometry ¹⁴C dating and multivariate analysis, were used to reconstruct Holocene vegetation and fire dynamics at Urio Quattrocchi, a small lake in the supra‐mediterranean belt in the Nebrodi Mountains of Sicily (Italy). The data suggest that after 10 000 cal a BP increasing moisture availability supported closed forests with deciduous (Quercus cerris, Fagus sylvatica and Fraxinus spp.) and evergreen (Quercus ilex) species. Species‐rich closed forest persisted until 6850 cal a BP, when Neolithic activities caused a forest decline and affected plant diversity. Secondary forest with abundant Ilex aquifolium recovered between 6650 and 6000 cal a BP, indicating moist conditions. From 5000 cal a BP, agriculture and pastoralism led to the currently fragmented landscape with sparse deciduous forests (Quercus cerris). The study suggests that evergreen broadleaved species were more important at elevations above 1000 m a.s.l. before ca. 5000 cal a BP than subsequently, which might reflect less human impact or warmer‐than‐today climatic conditions between 10 000 and 5000 cal a BP. Despite land use since Neolithic times, deciduous supra‐mediterranean forests were never completely displaced from the Nebrodi Mountains, because of favourable moist conditions that persisted throughout the Holocene. Reconstructed vegetation dynamics document the absence of any pronounced mid‐ or late‐Holocene ‘aridification’ trend at the site, an issue which is controversially debated in Italy and the Mediterranean region. Copyright © 2012 John Wiley & Sons, Ltd.     

Holocene precipitation and temperature variations in the East Asian monsoonal margin from pollen data from Hulun Lake in northeastern Inner Mongolia,China

Wen, R. L., Xiao, J. L., Chang, Z. G., Zhai, D. Y., Xu, Q. H., Li, Y. C. & Itoh, S. 2009: Holocene precipitation and temperature variations in the East Asian monsoonal margin from pollen data from Hulun Lake in northeastern Inner Mongolia, China. Boreas, 10.1111/j.1502‐3885.2009.00125.x. ISSN 0300‐9483. Quantitative palaeoclimatic reconstruction with the weighted averaging partial least squares method was applied to the pollen profile from Hulun Lake in northeastern Inner Mongolia. The data provide a detailed history of variations in precipitation and temperature over the northeastern margin of the East Asian summer monsoon during the Holocene. A warm and dry climate prevailed over the lake region until c. 8000 cal. BP. During the period c. 8000–4400 cal. BP, precipitation increased markedly and temperature gradually declined. The interval between c. 4400 and 3350 cal. BP was marked by extremely dry and relatively cold conditions. Precipitation recovered from c. 3350 to 1000 cal. BP, with temperatures rising c. 3350–2050 cal. BP and dropping c. 2050–1000 cal. BP. During the last 500 years, the climate of the lake region displayed a general trend of warming and wetting. While Holocene temperature variations in the mid‐high latitude monsoonal margin were controlled by changes in summer solar radiation in the Northern Hemisphere, they could also be related to the strength of the East Asian summer monsoon. The lack of precipitation during the early Holocene could be attributed to the weakened summer monsoon resulting from the existence of remnant ice sheets in the Northern Hemisphere. Changes in the monsoonal precipitation during the middle to late Holocene would have been associated with the ocean–atmosphere interacting processes occurring in the western tropical Pacific.     

The Holocene expansion of grassland in northern Europe reconstructed from molluscan assemblages

From the synthesis of the malacological data collected from 12 sites in the large flood‐plain of the Seine basin, three main environmental stages have been reconstructed. During the first half of the Holocene, forest environments are prevalent (Seine 1). As early as c. 6.5 cal. ka BP, the first evidence of woodland clearance is observed (Seine 2) and, from c. 3.4 cal. ka BP, the lowlands were largely cleared of trees and are dominated by grassland (Seine 3). This three‐stage development of environmental conditions is consistent with the environmental developments reconstructed from molluscan successions in England, Germany, Luxembourg, Switzerland, Poland, the Czech Republic and Slovakia. Our results highlight anthropogenic disturbance as the key factor in the openness of the Holocene landscape and pinpoint the period between c. 3.6 and c. 2.8 cal. ka BP as a transitional phase of this large‐scale environmental change.     

Late Younger Dryas and early Holocene palaeoenvironments in the Skagerrak,eastern North Atlantic: a multiproxy study

Erbs‐Hansen, D. R., Knudsen, K. L., Gary, A. C., Jansen, E., Gyllencreutz, R., Scao, V. & Lambeck, K. 2011: Late Younger Dryas and early Holocene palaeoenvironments in the Skagerrak, eastern North Atlantic: a multiproxy study. Boreas, 10.1111/j.1502‐3885.2011.00205.x. ISSN 0300‐9843 A high‐resolution study of palaeoenvironmental changes through the late Younger Dryas and early Holocene in the Skagerrak, the eastern North Atlantic, is based on multiproxy analyses of core MD99‐2286 combined with palaeowater depth modelling for the area. The late Younger Dryas was characterized by a cold ice‐distal benthic foraminiferal fauna. After the transition to the Preboreal (c. 11 650 cal. a BP) this fauna was replaced by a Cassidulina neoteretis‐dominated fauna, indicating the influence of chilled Atlantic Water at the sea floor. Persisting relatively cold bottom‐water conditions until c. 10 300 cal. a BP are presumably a result of an outflow of glacial meltwater from the Baltic area across south‐central Sweden, which led to a strong stratification of the water column at MD99‐2286, as also indicated by C. neoteretis. A short‐term peak in the C/N ratio at c. 10 200 cal. a BP is suggested to indicate input of terrestrial material, which may represent the drainage of an ice‐dammed lake in southern Norway, the Glomma event. After the last drainage route across south‐central Sweden closed, c. 10 300 cal. a BP, the meltwater influence diminished, and the Skagerrak resembled a fjord with a stable inflow of waters from the North Atlantic through the Norwegian Trench and a gradual increase in boreal species. Full interglacial conditions were established at the sea floor from c. 9250 cal. a BP. Subsequent warm stable conditions were interrupted by a short‐term cooling around 8300–8200 cal. a BP, representing the 8.2 ka event.     

Holocene hydrological variability of Lake Ladoga,northwest Russia,as inferred from diatom oxygen isotopes

This article presents a new comprehensive assessment of the Holocene hydrological variability of Lake Ladoga, northwest Russia. The reconstruction is based on oxygen isotopes of lacustrine diatom silica (δ¹⁸O_diatom) preserved in sediment core Co 1309, and is complemented by a diatom assemblage analysis and a survey of modern isotope hydrology. The data indicate that Lake Ladoga has existed as a freshwater reservoir since at least 10.8 cal. ka BP. The δ¹⁸O_diatom values range from +29.8 to +35.0‰, and relatively higher δ¹⁸O_diatom values around +34.7‰ between c. 7.1 and 5.7 cal. ka BP are considered to reflect the Holocene Thermal Maximum. A continuous depletion in δ¹⁸O_diatom since c. 6.1 cal. ka BP accelerates after c. 4 cal. ka BP, indicating Middle to Late Holocene cooling that culminates during the interval 0.8–0.2 cal. ka BP, corresponding to the Little Ice Age. Lake‐level rises result in lower δ¹⁸O_diatom values, whereas lower lake levels cause higher δ¹⁸O_diatom values. The diatom isotope record gives an indication for a rather early opening of the Neva River outflow at c. 4.4–4.0 cal. ka BP. Generally, overall high δ¹⁸O_diatom values around +33.5‰ characterize a persistent evaporative lake system throughout the Holocene. As the Lake Ladoga δ¹⁸O_diatom record is roughly in line with the 60°N summer insolation, a linkage to broader‐scale climate change is likely.     

8000 years of vegetation history in the northern Iberian Peninsula inferred from the palaeoenvironmental study of the Zalama ombrotrophic bog (Basque‐Cantabrian Mountains,Spain)

This paper focuses on pollen, spores, non‐pollen palynomorphs (NPPs) and certain geochemical elements from the ombrotrophic blanket bog of Zalama (Basque‐Cantabrian Mountains, northern Iberian Peninsula), with the support of a robust chronology based on 17 AMS ¹⁴C dates. The main results related to the last 8000 years show that, during the early middle Holocene, pines and deciduous forests were the most extensive tree formations. At the beginning of the succession, pines reach 44%, showing regional presence, whereas after 7600 cal. a BP, deciduous forests were particularly abundant. From c. 6500 cal. a BP the pollen diagram constructed from our samples shows the first anthropogenic evidence, linked with the new economic practices related to the Neolithic of the Basque‐Cantabrian Mountains. From 3300 cal. a BP the expansion of Fagus sylvatica is particularly clear, and has since then become one of the dominant forest species in this region. We also discuss the Holocene evolution of other noteworthy plant communities in southwestern Europe, such as Taxus baccata, Juglans and shrublands.     

Late Quaternary history of the Kap Mackenzie area,northeast Greenland

Wagner, B., Bennike, O., Cremer, H. & Klug, M. 2010: Late Quaternary history of the Kap Mackenzie area, northeast Greenland. Boreas, Vol. 39, pp. 492–504. 10.1111/j.1502‐3885.2010.00148.x. ISSN 0300‐9483. The Kap Mackenzie area on the outer coast of northeast Greenland was glaciated during the last glacial stage, and pre‐Holocene shell material was brought to the area. Dating of marine shells indicates that deglaciation occurred in the earliest Holocene, before 10 800 cal. a BP. The marine limit is around 53 m a.s.l. In the wake of the deglaciation, a glaciomarine fauna characterized the area, but after c. one millennium a more species‐rich marine fauna took over. This fauna included Mytilus edulis and Mysella sovaliki, which do not live in the region at present; the latter is new to the Holocene fauna of northeast Greenland. The oldest M. edulis sample is dated to c. 9500 cal. a BP, which is the earliest date for the species from the region and indicates that the Holocene thermal maximum began earlier in the region than previously documented. This is supported by driftwood dated to c. 9650 cal. a BP, which is the earliest driftwood date so far from northeastern Greenland and implies that the coastal area was at least partly free of sea ice in summer. As indicated by former studies, the Storegga tsunami hit the Kap Mackenzie area at c. 8100 cal. a BP. Loon Lake, at 18 m a.s.l., was isolated from the sea at c. 6200 cal. a BP, which is distinctly later than expected from existing relative sea‐level curves for the region.     

Vegetation and climate c. 12 300–9000 cal. yr BP at Andøya,NW Norway

Owing to proximity of the North Atlantic Stream and the shelf, the Andøya biota are assumed to have responded rapidly to climatic changes taking place after the Weichselian glaciation. Palynological, macrofossil, loss‐on‐ignition, tephra and ¹⁴C data from three sites at the northern part of the island of Andøya were studied. The period 12 300–11 950 cal. yr BP was characterized by polar desert vegetation, and 11 950–11 050 cal. yr BP by a moisture‐demanding predominantly low‐arctic Oxyria vegetation. During the period 11 050–10 650 cal. yr BP, there was a climatic amelioration towards a sub‐arctic climate and heaths dominated by Empetrum. After 10 650 cal. yr BP the Oxyria vegetation disappeared. As early as about 10 800 cal. yr BP the bryozoan Cristatella mucedo indicated a climate sufficient for Betula woodland. However, tree birch did not establish until 10 420–10 250 cal. yr BP, indicating a time‐lag for the formation of Betula ecotypes adapted to the oceanic climate of Andøya. From about 10 150 to 9400 cal. yr BP the summers were dry and warm. There was a change towards moister, though comparatively warm, climatic conditions about 9400 cal. yr BP. The present data are compared with evidence from marine sediments and the deglaciation history in the region. It is suggested that during most of the period 11 500–10 250 cal. yr BP a similar situation as in present southern Greenland existed, with birch woodland in the inner fjords near the ice sheet and low‐arctic heath vegetation along the outer coast.     

Palaeoecological evidence of changes in vegetation and climate during the Holocene in the pre‐Polar Urals,northeast European Russia

This study investigated Holocene tree‐line history and climatic change in the pre‐Polar Urals, northeast European Russia. A sediment core from Mezhgornoe Lake situated at the present‐day alpine tree‐line was studied for pollen, plant macrofossils, Cladocera and diatoms. A peat section from Vangyr Mire in the nearby mixed mountain taiga zone was analysed for pollen. The results suggest that the study area experienced a climatic optimum in the early Holocene and that summer temperatures were at least 2°C warmer than today. Tree birch immigrated to the Mezhgornoe Lake area at the onset of the Holocene. Mixed spruce forests followed at ca. 9500–9000 ¹⁴C yr BP. Climate was moist and the water level of Mezhgornoe Lake rose rapidly. The hypsithermal phase lasted until ca. 5500–4500 ¹⁴C yr BP, after which the mixed forest withdrew from the Mezhgornoe catchment as a result of the climate cooling. The gradual altitudinal downward shift of vegetation zones resulted in the present situation, with larch forming the tree‐line. Copyright © 2003 John Wiley & Sons, Ltd.     

Changes of treelines and alpine vegetation in relation to post‐glacial climate dynamics in northern Fennoscandia based on pollen and chironomid records

Palaeoclimatic records derived from a variety of independent proxies provide evidence of post‐glacial changes of temperature and soil moisture in northern Fennoscandia. We use pollen percentage, pollen influx, stomatal and chironomid records from Toskaljavri, a high‐altitude lake in northern Finland, to assess how treelines and alpine vegetation there have responded to these climate changes. The evidence suggests that the cool, moist climate of the early Holocene supported birch forest in the area 9600 cal. yr BP onwards and that a rise of temperature triggered the immigration of pine at 8300 cal. yr BP. At 6100–4000 cal. yr BP altitudinal treeline in the area was formed by pine, in contrast to the modern situation where mountain birch reaches a higher elevation. Alpine vegetation also demonstrates clear changes. Plant communities typical of dry, oligotrophic heaths of northern Fennoscandia expanded during the dry climatic period at 7000–4000 cal. yr BP and decreased in response to cooler and moister conditions after 4000 cal. yr BP. Alpine plant communities favouring moist sites show an inverse pattern, expanding after a change towards moister climate after 4000 cal. yr BP. In a redundancy analysis (RDA), a statistically significant proportion of the variability in the total chironomid assemblages was captured by changes in the pollen types reflecting alpine vegetation typical of moist sites. Although chironomid community changes appeared to follow the major patterns in the alpine vegetation succession, the present study does not support a direct link between the changing treeline position and chironomid stratigraphy. Rather, the data indicate that the terrestrial and aquatic environments have each responded directly to the same ultimate cause, namely changing Holocene climate. Copyright © 2002 John Wiley & Sons, Ltd.     

Fire and drought as drivers of early Holocene tree line changes in the Peruvian Andes

Three pollen and charcoal records from three lakes lying at 3400 m elevation in southern Peru provided a record of landscape change spanning the last ca.18 000 cal. a BP. The tree line lay close to the site between ca. 16 000 and 12 000 cal. a BP, with Polylepis woodlands growing near the lakes. Progressively drying conditions led to increased fire after 12 000 cal. a BP, coinciding with a decline in Polylepis cover and Andean forest relicts as puna grasslands expanded. A strong decrease in the rate of sediment deposition between ca. 12 000 and ca. 4400 cal. a BP was interpreted to indicate the presence of sedimentary hiatuses. With the return of wet conditions after 4400 cal. a BP, forests did not reassemble around the lakes. Instead, fire‐maintained grasslands dominated the landscape. Humans probably induced the intensified fire activity during the late Holocene and thereby deflected local successions. The modern fragmented landscape, with Polylepsis woodlands existing in fire‐resistant pockets above the general limit of the Andean tree line, resulted from the intensification of human land use practices during the last 4400 cal. a BP. Copyright © 2011 John Wiley & Sons, Ltd.