Late-Glacial Vegetation and Climate Change in Western Oregon

Pollen records from two sites in western Oregon provide information on late-glacial variations in vegetation and climate and on the extent and character of Younger Dryas cooling in the Pacific Northwest. A subalpine forest was present at Little Lake, central Coast Range, between 15,700 and 14,850 cal yr B.P. A warm period between 14,850 and 14,500 cal yr B.P. is suggested by an increase inPseudotsugapollen and charcoal. The recurrence of subalpine forest at 14,500 cal yr B.P. implies a return to cool conditions. Another warming trend is evidenced by the reestablishment ofPseudotsugaforest at 14,250 cal yr B.P. Increased haploxylonPinuspollen between 12,400 and 11,000 cal yr B.P. indicates cooler winters than before. After 11,000 cal yr B.P. warm dry conditions are implied by the expansion ofPseudotsuga.A subalpine parkland occupied Gordon Lake, western Cascade Range, until 14,500 cal yr B.P., when it was replaced during a warming trend by a montane forest. A rise inPinuspollen from 12,800 to 11,000 cal yr B.P. suggests increased summer aridity.Pseudotsugadominated the vegetation after 11,000 cal yr B.P. Other records from the Pacific Northwest show an expansion ofPinusfrom ca. 13,000 to 11,000 cal yr B.P. This expansion may be a response either to submillennial climate changes of Younger Dryas age or to millennial-scale climatic variations.     

Late glacial and postglacial pollen and plant macrofossils from Lake West Okoboji, Northwestern Iowa

Pollen and plant macrofossils preserved in lake sediment from Lake West Okoboji, Dickinson County, Iowa, indicate how the vegetation of that area changed during the late glacial and postglacial. A closed coniferous forest, dominated by spruce and larch trees, produced the Picea-Larix pollen assemblage zone. Fir trees were a minor constituent of this forest; pine trees were probably absent. Black ash trees increased in abundance at Lake West Okoboji and by 13,500 yr ago were an important constituent of the forest. The sediment accumulation rate and the pollen influx were low throughout this time. Birch and alder pollen peaked in abundance approximately 11,800 yr ago. Pollen influx increased rapidly as birch and alder replaced coniferous trees on the uplands. A deciduous forest, containing abundant oak and elm trees, replaced the birch-alder-coniferous forest. This forest inhabited northwestern Iowa from approximately 11,000 to 9000 yr B.P. Nonarboreal species became prevalent between approximately 9000 and 7700 yr B.P. as prairie began to replace deciduous forest on the uplands. Charred remains of Amorpha canescens and other upland species attest to the presence of prairie fires as an aid in establishing prairie and destroying the forest. The pollen influx declined. The warmest, driest part of the postglacial occurred in northwestern Iowa from approximately 7700 to 3200 yr ago. Lake level fell 9 to 10 m, and prairie extended to the edge of the lake. Wet-ground weeds inhabited areas near lake level which were alternately flooded, then dry. Pollen influx was approximately 100 grains/cm²/yr during the driest time in this dry interval.Deciduous trees, particularly oaks, returned after approximately 3200 yr B.P. Prairie continued to occupy the uplands but trees were more common in the lowlying wet areas. Settlement by Europeans in northwestern Iowa about 1865 is marked by an increase in weed pollen. Macrofossil deposition changed in 1910 in response to the stabilization of lake level.     

Palynological evidence for early Holocene aridity in the southern Sierra Nevada, California

Sediments of Balsam Meadow have produced a 11,000-yr pollen record from the southern Sierra Nevada of California. The Balsam Meadow diagram is divided into three zones. (1) The Artemisia zone (11,000–7000 yr B.P.) is characterized by percentages of sagebrush (Artemisia) and other nonarboreal pollen higher than can be found in the modern local vegetation. Vegetation during this interval was probably similar to the modern vegetation on the east slope of the Sierra Nevada and the climate was drier than that of today. (2) Pinus pollen exceeded 80% from 7000 to 3000 yr B.P. in the Pinus zone. The climate was moister than during the Artemisia zone. (3) Fir (Abies, Cupressaceae, and oak (Quercus) percentages increased after 3000 yr B.P. in the Abies zone as the modern vegetation at the site developed and the present cool-moist climatic regime was established. Decreased fire frequency after 1200 yr B.P. is reflected in decreased abundance of macroscopic charcoal and increased concentration of Abies magnifica and Pinus murrayana needles.     

Late quaternary pollen records from the Kobuk and Noatak river drainages, northwestern Alaska

Pollen diagrams from Joe and Niliq Lakes date to ca. 28,000 and 14,000 yr B.P., respectively. Mesic shurb tundra grew near Joe Lake ca. 28,000 to 26,000 yr B.P. with local Populus populations prior to ca. 27,000 yr B.P. Shrub communities decreased as climate changed with the onset of Itkillik II glaciation (25,000 to 11,500 yr B.P.), and graminoid-dominated tundra characterized vegetation ca. 18,500 to 13,500 yr B.P. Herb tundra was replaced by shrub Betula tundra near both sites ca. 13,500 yr B.P. with local expansion of Populus ca. 11,000 to 10,000 yr B.P. and Alnus ca. 9000 yr B.P. Mixed Picea glauca/P. mariana woodland was established near Joe Lake ca. 6000 yr B.P. These pollen records when combined with others from northern Alaska and northwestern Canada indicate (1) mesic tundra was more common in northwestern Alaska than in northeastern Alaska or northwestern Canada during the Duvanny Yar glacial interval (25,000 to 14,000 yr B.P.); (2) with deglaciation, shrub Betula expanded rapidly in northwestern Alaska but slowly in areas farther east; (3) an early postglacial thermal maximum occurred in northwestern Alaska but had only limited effect on vegetation; and (4) pollen patterns in northern Alaska and northwestern Canada suggest regional differences in late Quaternary climates.     

Late-Quaternary vegetation history at White Pond on the inner Coastal Plain of South Carolina

At White Pond near Columbia, South Carolina, a pollen assemblage of Pinus banksiana (jack pine), Picea (spruce), and herbs is dated between 19,100 and 12,800 ¹⁴C yr B.P. Plants of sandhill habitats are more prominent than at other sites of similar age, and pollen of deciduous trees is infrequent. The vegetation was probably a mosaic of pine and spruce stands with prairies and sand-dune vegetation. The climate may have been like that of the eastern boreal forest today. ¹⁴C dates of 12,800 and 9500 yr B.P. bracket a time when Quercus (oak), Carya (hickory), Fagus (beech), and Ostrya-Carpinus (ironwood) dominated the vegetation. It is estimated that beech and hickory made up at least 25% of the forest trees. Conifers were rare or absent. The environment is interpreted as hickory-rich mesic deciduous forest with a climate similar to but slightly warmer than that of the northern hardwoods region of western New York State. After 9500 yr B.P. oak and pine forest dominated the landscape, with pine becoming the most important tree genus in the later Holocene.     

Palynological Evidence for Climatic Change and Human Activity during the Holocene on Minorca (Balearic Islands)

Four pollen diagrams from Minorca (Balearic Islands) have been correlated with other previously studied sequences from Majorca and Minorca to define a Holocene landscape sequence for the region from 8000 yr B.P. to the present. The lower part of the pollen diagrams reflects a climatic phase with more rain and less-marked seasonality than today. Significant quantities ofCorylus, Buxus,and mesophilous taxa are found. In the middle part, between 5000 and 4000 yr B.P., a strong change is recorded in composition and structure of the vegetational landscape, with vegetation appearing that was adapted to Mediterranean conditions. This episode coincided with the first human colonization of the island and also with a widespread climatic change in the western Mediterranean region. The change in taxa was complex and some sclerophyllous taxa suchOleaplayed an important role in the transformation of the landscape physiognomy from the mid-Holocene until the present. Although human activities have removed much of the Mediterranean vegetation on the Balearic Islands, it seems clear that the changes have been brought about, in part, by increasing dryness.     

A late quaternary pollen record from the Transvaal bushveld, South Africa

Pollen spectra from cores of organic spring deposits from the Transvaal provide evidence for the climatic evolution of the province during the last 35,000 yr B.P. or more. The past climatic phases are derived from palynological reconstructions of past vegetation types by comparison of fossil pollen data with modern surface pollen spectra from various localities. Evidence is provided for an early moist, cool phase with relatively mesic bushveld and expanded montane forest in the central Transvaal, followed by a drier period with drier bushveld which probably lasted until approximately 25,000 yr B.P. During the next phase, which at the latest ended about 11,000 yr B.P., the temperatures were probably 5°–6°C cooler than at present. At that time bushveld vegetation in the central Transvaal was replaced by open grassland with macchia elements. Climatic amelioration came and semiarid savanna returned to the plains, at first gradually and then developing into a warm Kalahari thornveld-type vegetation. After 6000 yr B.P. it apparently became slightly wetter and a more broad-leafed bushveld developed. About 4000 yr B.P. it again became cooler and slightly wetter and the bushveld vegetation on the central and northern plains was comparable to present open upland types. After 2000 yr B.P. conditions gradually became warner until about 1000 yr B.P., when the modern climate of the central Transvaal bushveld originated.     

Quaternary vegetation history of the Mississippi embayment

Nonconnah Creek, located in the loess-mantled Blufflands along the eastern wall of the Lower Mississippi Alluvial Valley in Tennessee displays a sedimentary sequence representing the Altonian Substage through the Woodfordian Substage of the Wisconsinan Stage. The site has a biostratigraphic record for the Altonian and Farmdalian Substages that documents warm-temperate upland oak-pine forest, prairie, and bottomland forest. At 23,000 yr B.P., white spruce and larch migrated into the Nonconnah Creek watershed and along braided-stream surfaces in the Mississippi Valley as far as southeastern Louisiana. The pollen and plant-macrofossil record from Nonconnah Creek provides the first documentation of a full-glacial locality in eastern North America for beech, yellow poplar, oak, history, black walnut, and other mesic deciduous forest taxa. During the full and late glacial, the Mississippi Valley was a barrier to the migration of pine species, while the adjacent Blufflands provided a refuge for mesic deciduous forest taxa. Regional climatic amelioration, beginning about 16,500 yr B.P., is reflected by increases in pollen percentages of cooltemperate deciduous trees at Nonconnah Creek. The demise of spruce and jack pine occurred 12,500 yr B.P. between 34° and 37° N in eastern North America in response to postglacial warming.     

Late Pleistocene piñon pines in the Chihuahuan Desert

Examination of late Pleistocene packrat middens from the northern and central Chihuahuan Desert disclosed macrofossils of Colorado piñon (Pinus edulis) and Texas piñon (P. remota). Radiocarbon dating indicates that Texas piñon was widespread in Trans-Pecos Texas and northeastern Mexico between 30,000 and 11,000 yr B.P. Today it is found in small refugia east of its former range. In the late Pleistocene Colorado piñon occurred at lower elevations on the northern edge of the Chihuahuan Desert. Both species occurred in the Hueco Mountains, near El Paso, Texas. No clear evidence was found of the presence of Mexican piñon (P. cembroides), though today it is abundant in the Davis and Chisos Mountains. A paleoclimate is postulated that had the following characteristics: increased winter precipitation from Pacific frontal sources, reduced summer temperatures and precipitation, and milder winter temperatures due to a reduced frequency of Arctic airmass incursion. Winter precipitation appears to have decreased from north to south, while winter temperatures, and, possibly, summer precipitation, increased from north to south. During the late Pleistocene, the northern Chihuahuan Desert was dominated by woodlands of piñon pines, junipers, and oaks. The desert-scrub communities that characterize the area today are a Holocene phenomenon.     

A late Wisconsin and Holocene vegetation history from the central brooks range: Implications for Alaskan palaeoecology

Five pollen diagrams reveal late Wisconsin and Holocene vegetation changes in the Walker Lake/Alatna Valley region of the central Brooks Range, approximately 100 km west of the area studied by D. A. Livingstone (1955, Ecology36, 587–600). New insights into the vegetation history of this region are provided by calculations of pollen influx and by the use of linear discriminant analysis to separate Picea glauca and P. mariana pollen. Three major pollen zones are identified: (1) a basal herb zone, characterized by high percentages of Cyperaceae, Gramineae, Salix, and Artemisia, and low total pollen influx; (2) a shrub Betula zone with increased total pollen influx and very high percentages of Betula pollen, predominantly in the size range of B. nana and B. glandulosa; and (3) and Alnus zone dominated by Alnus pollen. Lakes currently within the boreal forest or near tree line show relatively high percentages of Picea pollen in the Alnus zone. Several striking vegetation changes occurred between ca. 10,000 and 7000 yr B.P. Between ca. 11,000 and 10,000 yr B.P., Populus balsamifera pollen percentages as great as 30% indicate that this species was present at low-elevation sites near Walker Lake. These populations declined abruptly ca. 10,000 yr ago and have never regained prominence. About 8500 yr B.P., Picea glauca pollen reached 10–15%, indicating the arrival of P. glauca in or near the study area. P. glauca populations evidently decreased ca. 8000 yr ago, when Picea pollen percentages and influx fell to low values. About 7000 yr B.P., Alnus pollen percentages and influx rose sharply as alder shrubs became established widely. Picea once more expanded ca. 5000 yr ago, but these populations were dominated by P. mariana rather than P. glauca, which increased slowly at this time and may still be advancing northward. Some vegetation changes have been remarkably synchronous over wide areas of interior Alaska, and probably reflect responses of in situ vegetation to environmental changes, but others may reflect the lagged responses of species migrating into new areas.     

Expansion of Magellanic Moorland during the late Pleistocene: Palynological evidence from northern Isla de Chiloé, Chile

The late Quaternary vegetation of northern Isla de Chiloé is inferred from palynological analysis of a section in the Río Negro drainage (42°03′S, 73°50′W). At ca. 30,500 yr B.P., maxima of Astelia and Donatia occurred, suggesting wetland development. From that time until ca. 27,000 yr B.P., steppe indicators such as Compositae/Gramineae dominated, suggesting drier conditions. After 27,000 yr B.P., the moorland shrub Dacrydium gradually increased, reaching a maximum by 18,000 yr B.P. At this time Astelia increased again, suggesting development of cushion bog during cold and wet conditions. The glacial-postglacial transition is characterized by a marked change from peaty sediments to clays, a decrease in the cushion bog flora, and the prevalence of Gramineae/ Compositae and swamp taxa. This vegetation prevailed until ca.7000 yr B.P. when forest taxa became dominant. The floristic pattern inferred from the pollen spectra of the Rio Negro section suggests that the late Pleistocene vegetation of Chiloé resembled modern Magellanic Moorland vegetation (52°–56°lat S). Based on climatic conditions presently associated with Magellanic Moorland, its occurrence in Chiloé at low elevations during the late Pleistocene implies a decrease in average temperature of at least 4°C and an increase in annual precipitation of at least 1500 mm.     

Late quaternary vegetational change in the Kotzebue sound area, northwestern Alaska

Two sediment cores from Kaiyak and Squirrel lakes in northwestern Alaska yielded pollen records that date to ca. 39,000 and 27,000 yr B.P., respectively. Between 39,000 and 14,000 yr B.P., the vegetation around these lakes was dominated by Gramineae and Cyperaceae with some Salix and possibly Betula nana/glandulosa forming a local, shrub component of the vegetation. Betula pollen percentages increased about 14,000 yr B.P., indicating the presence of a birchdominated shrub tundra. Alnus pollen appeared at both sites between 9000 and 8000yr B.P., and Picea pollen (mostly P. mariana) arrived at Squirrel Lake about 5000 yr B.P. The current foresttundra mosaic around Squirrel Lake was established at this time, whereas shrub tundra existed near Kaiyak Lake throughout the Holocene. When compared to other pollen records from north-western North America, these cores (1) represent a meadow component of lowland. Beringian tundra between 39,000 and 14,000 yr B.P., (2) demonstrate an early Holocene arrival of Alnus in northwestern Alaska that predates most other Alnus horizons in northern Alaska or northwestern Canada, and (3) show an east-to-west migration of Picea across northern Alaska from 9000 to 5000 yr B.P.     

Paleoecology of an early Holocene faunal and floral assemblage from the Dows Local Biota of north-central Iowa

The faunas and floras from the Dows Local Biota provide an opportunity to compare Holocene taxa without a cultural bias. The Dows Local Biota is located in a large depression on the back side (north) of the Altamont I Moraine complex within the Des Moines Lobe. The Dows Silt Fauna/Flora ( = DSF; ca. 9380 ± 130 yr B.P.), one horizon of the Dows Local Biota, was collected for plant macrofossils, mollusks, and micromammals. DSF terrestrial gastropods are upland mesic forest dwellers although one species, Strobilops affinis, is characteristic of more xeric forests and may represent open woods. The aquatic gastropods reflect both permanent and periodic waters. DSF micromammals prefer an open, mesic, deciduous forest. The micromammal sympatry is restricted to a small area within the tension zone and deciduous forest belt of west-central Wisconsin. DSF plants are characteristic of upland forests, moist meadowlands or disturbed areas, and aquatic habitats. The DSF plant sympatry is large but restricted to the conifer-hardwood and deciduous forests along the Great Lakes-New England regions. Quantitative climatic data for the combined DSF sympatries suggest that Dows (ca. 9380 yr B.P.) was cooler than at present, and is nearly identical to that achieved by pollen analyses at the Cherokee Sewer-Lake West Okoboji sites (ca. 9000 yr B.P.) in northwest Iowa. Based on common habitat interpretations and sympatries, about 9380 yr B.P. north-central Iowa was cooler and moister than at present and was occupied by an open deciduous forest.     

Late Wisconsin vegetational and climatic history at Kylen Lake, northeastern Minnesota

Kylen Lake, located within the Toimi drumlin field, is critically positioned in relation to Late Wisconsin glacial advances, for it lies between the areas covered by the Superior and St. Louis glacial lobes between 12,000 and 16,000 yr B.P. The pollen and plant-macrofossil record suggests the presence of open species-rich “tundra barrens” from 13,600 to 15,850 yr B.P. Small changes in percentages of Artemisia pollen between 14,300 and 13,600 yr B.P. appear to be artifacts of pollen-percentage data. Shrub-tundra with dwarf birch, willow, and Rhododendron lapponicum developed between 13,600 and 12,000 yr B.P. Black and white spruce and tamarack then expanded to form a vegetation not dissimilar to that of the modern forest-tundra ecotone of northern Canada. At 10,700 B.P. spruce and jack pine increased to form a mosaic dominated by jack pine and white spruce on dry sites and black spruce, tamarack, and deciduous trees such as elm and ash on moist fertile sites. At 9250 yr B.P. red pine and paper birch became dominant to form a vegetation that may have resembled the dry northern forests of Wisconsin today. The diagram terminates at 8410 ± 85 yr B.P. Climatic interpretation of this vegetational succession suggests a progressive increase in temperature since 14,300 yr B.P. This unidirectional trend in climate contrasts with the glacial history of the area. Hypotheses are presented to explain this lack of correspondence between pollen stratigraphy and glacial history. The preferred hypothesis is that the ice-margin fluctuations were controlled primarily by changes in winter snow accumulation in the source area of the glacier, whereas the vegetation and hence the pollen stratigraphy were controlled by climatic changes in front of the ice margin.     

Late quaternary vegetation of southern Isla Grande de Chiloé, Chile

Late-glacial-Holocene forest history of southern Isla Chiloé (latitude 43°10′ S) was reconstructed on the basis of pollen analysis in three profiles (Laguna Soledad, Laguna Chaiguata, Puerto Carmen). Prior to 12,500 yr B.P. pollen records are dominated by plant taxa characteristic of open habitats (Zone I). From 12,500 yr B.P. to the present, tree species predominate in the pollen records (Zones II–V). Between 12,500 and 9500 yr B.P. ombrophyllous taxa (Nothofagus, Podocarpus nubigena. Myrtaceae, Fitzroya/Pilgerodendron, and Drimys) are frequent in all pollen diagrams, suggesting a wetter and colder climate than the present. Between 9000 and 5500 yr B.P. Valdivian forest elements, such as Nothofagus dombeyi type, Weinmannia, and Eucryphia/Caldcluvia, dominate, indicating a period of drier and warmer climate. From 5500 yr B.P. onward, the expansion of mixed North Patagonian-Subantarctic forest elements and the increased frequence of Tepualia suggest increased rainfall and temperatures oscillating around the modern values.The change from open to forest vegetation (ca. 12,500 yr B.P.) probably represents the most pronounced climatic change in the record and can be interpreted as the glacial-postglacial transition in the study area.     

Late-glacial vegetation and climate at the Manis Mastodon site,Olympic Peninsula,Washington

As the late Wisconsin Cordilleran Ice Sheet retreated, sediment accumulated in shallow depressions at the Manis Mastodon Archaeological site on the Olympic Peninsula, near Sequim, Washington. Pollen, plant macrofossils, and bones of mastodon, caribou, and bison occur within the lower 47 cm of these deposits. The fossil pollen and seed assemblages indicate persistence for 1000 yr (11,000–12,000 yr B.P.) of an herb-and-shrub-dominated landscape at a time when forest species appear elsewhere in Washington and in adjacent British Columbia.At present, Sequim is near the northern coastal limits of both Cactaceae and Ceratophyllum. Mean annual precipitation is 42.7 cm and summer temperatures average 15°–16°C in July. The absence of coniferous trees and the presence of cactus and Ceratophyllum in late-glacial sediments are explained by a regional climate that was drier and at least as warm as today. These conditions persisted in the rain shadow of the Olympic Mountains until at least 11,000 yr B.P.     

Reconstruction of climate and vegetation changes of Lake Bayanchagan (Inner Mongolia): Holocene variability of the East Asian monsoon

A high-resolution pollen and Pediastrum record, spanning 12,500 yr, is presented for Lake Bayanchagan (115.21°E, 41.65°N, and 1355 m a.s.l.), southern Inner Mongolia. Individual pollen taxa (PT-MAT) and the PFT affinity scores (PFT-MAT) were used for quantitative climatic reconstruction from pollen and algal data. Both techniques indicate that a cold and dry climate, similar to that of today, prevailed before 10,500 cal yr B.P. The wettest climate occurred between 10,500 and 6500 cal yr B.P., at which time annual precipitation was up to 30–60% higher than today. The early Holocene increases in temperature and precipitation occurred simultaneously, but mid-Holocene cooling started at approximately 8000 cal yr B.P., 1500 yr earlier than the drying. Vegetation reconstruction was based on the objective assignment of pollen taxa to the plant functional type. The results suggest that this region was dominated by steppe vegetation throughout the Holocene, except for the period 9200 to 6700 cal yr B.P., when forest patches were relatively common. Inner Mongolia is situated at the limit of the present East Asian monsoon and patterns of vegetation and climate changes in that region during the Holocene probably reflect fluctuations in the monsoon's response to solar insolation variations. The early to middle Holocene monsoon undoubtedly extended to more northern latitudes than at present.     

Biogeographic implications of a packrat midden sequence from the Sacramento Mountains, south-central New Mexico

Thirteen packrat (Neotoma spp.) and two porcupine (Erethizon dorsatum) middens from 1555 to 1690 m elevation from the Sacramento Mountains, New Mexico, provide an 18,000-yr vegetation record in the northern Chiuahuan Desert. The vegetation sequence is a mesic, Wisconsin fullglacial (18,000–16,000 yr B.P.) pinyon-juniper-oak woodland; a xeric, early Holocene (ca. 11,000–8000 yr B.P.) juniper-oak woodland; a middle Holocene (ca. 8000-4000 yr B.P.) desert-grassland; and a late Holocene (ca. 4000 yr B.P. to present) Chihuahuan desertscrub. The frequency of spring freezes and summer droughts in the late Wisconsin probably set the northern limits of Pinus edulis and Juniperus monosperma at about 34°N, or 6° south of today's limit. Rising summer tempratures in the early Holocene eliminated pinyon and other mesic woodland plants from the desert lowlands and allowed the woodland to move upslope. At this time pinyon-juniper woodland and pine forest dominated by Pinus ponderosa probably began their spectacular Holocene expansions to the north. Continued warming in the middle Holocene led to very warm summers with strong monsoons, relatively dry, cold winters, and widespread desert-grasslands. Desertscrub communities in the northern Chihuahuan Desert did not develop until the late Holocene when the biseasonal rainfall shifted slightly back toward the winter, catastrophic winter freezes decreased, and droughts in all seasons increased. The creosote bush desertscrub corridor across the Continental Divide between the Chihuahuan and Sonoran deserts was probably connected for the first time since the last interglaciation.     

Postglacial Vegetation and Climate of the Cascade Range, Central Oregon

Pollen data from two sites provide information on the postglacial vegetation and climate history of the Cascade Range. Indian Prairie in the western Cascade Range was colonized by subalpine forests of Pinus, Picea, and Tsuga and open meadows prior to ca. 12,400 ¹⁴C yr B.P. The treeline lay 500 to 1000 m below its modern elevation and conditions were cooler than at present. From ca. 12,400 to ca. 9950 ¹⁴C yr B.P. Abies became important and the forest resembled that presently found at middle elevations in the western Cascade Range. The pollen record implies a rise in treeline and warmer conditions than before. From ca. 10,000 to 4000-4500 ¹⁴C yr B.P., conditions that were warmer and effectively drier than today led to the establishment of a closed forest composed of Pseudotsuga , Abies, and, at lower elevations, Quercus and Corylus . During this period, Gold Lake Bog in the High Cascades was surrounded by closed forest of Pinus and Abies. The early-Holocene pollen assemblages at both Indian Prairie and Gold Lake Bog lack modern analogues, and it is likely that greater-than-present summer radiation fostered unique climatic conditions and vegetation associations at middle and high elevations. In the late Holocene, beginning ca. 4000-4500 ¹⁴C yr B.P., cooler and more humid conditions prevailed and the modern vegetation was established. A comparison of these sites with others in the Pacific Northwest suggests that major patterns of vegetational change at individual sites were a response to large-scale changes in the climate system that affected the entire region.     

A 35,000 Year Vegetation and Climate History from Potato Lake, Mogollon Rim, Arizona

A new record from Potato Lake, central Arizona, details vegetation and climate changes since the mid-Wisconsin for the southern Colorado Plateau. Recovery of a longer record, discrimination of pine pollen to species groups, and identification of macrofossil remains extend Whiteside's (1965) original study. During the mid-Wisconsin (ca. 35,000-21,000 yr B.P.) a mixed forest of Engelmann spruce (Picea engelmannii) and other conifers grew at the site, suggesting a minimum elevational vegetation depression of ca. 460 m. Summer temperatures were as much as 5°C cooler than today. During the late Wisconsin (ca. 21,000-10,400 yr B.P.), even-cooler temperatures (7°C colder than today; ca. 800 m depression) allowed Engelmann spruce alone to predominate. Warming by ca. 10,400 yr B.P. led to the establishment of the modern ponderosa pine (Pinus ponderosa) forest. Thus, the mid-Wisconsin was not warm enough to support ponderosa pine forests in regions where the species predominates today. Climatic estimates presented here are consistent with other lines of evidence suggesting a cool and/or wet mid-Wisconsin, and a cold and/or wet late-Wisconsin climate for much of the Southwest. Potato Lake was almost completely dry during the mid-Holocene, but lake levels increased to near modern conditions by ca. 3000 yr B.P.