Distribution patterns of the ghost crab Ocypode cursor on sandy beaches of a tropical island in the Cabo Verde archipelago,Eastern Central Atlantic

The spatial distribution of the ghost crab Ocypode cursor was determined for beaches on eastern Boa Vista Island, Cabo Verde Archipelago. The main objectives were to analyse the across-shore distribution by means of burrow counts and to identify preferential zones and spatial segregation. Six beaches were investigated using 20 across-shore beach transects. It was demonstrated that ghost crabs occur throughout the entire intertidal zone, with higher densities in a 9–13 m band above the waterline. There was an exponential relationship between the carapace length of captured crabs and the diameter of their burrows. Mean burrow diameter was correlated with burrow density and increased linearly with distance from the waterline, although small burrows, representing individuals smaller than the median size, occurred across the beach. Burrow diameter and depth were positively correlated, with larger holes indicating larger crabs and deeper burrows. Deeper burrows were more common in the upper intertidal zone. These results are similar to those described for ghost crab populations elsewhere and contribute to an understanding of the ecology of the species in the local context.     

Synergistic effect of extreme climatic events and urbanization on population density of the ghost crab Ocypode quadrata (Fabricius, 1787)

The combined effects of urbanization and extreme climatic events were examined on the ghost crab Ocypode quadrata, a well‐recognized ecological indicator in sandy beaches. The effect of storm waves was assessed before and after these climate events in beaches with different levels of urbanization in the coast of Brazil, Tropical Atlantic Ocean. The number of burrows was used as a metric for the population size of O. quadrata and the number of humans along the beaches as a proxy for the urbanization level. Significant differences were recorded in this species’ population density according to the human pressure. Our core modelling approach showed that the number of O. quadrata burrows was lower after storm waves in urbanized zones, compared to non‐urbanized ones. The interaction between storm waves and urbanization impacted the number of O. quadrata burrows negatively, and suggest that species’ populations are more vulnerable to the combined effects of extreme climatic events and urbanization. Extreme events are predicted to increase in frequency in the next few decades, and may modify or impair ecosystem functioning in sandy beaches by reducing populations of the O. quadrata, an important beach scavenger, which markedly contribute to the flux of energy in coastal trophic webs.     

青岛潮间带底内蟹类生态及洞穴观察

栖息于潮间带的海生动物中,营底内生活的种类占很大的比例。这类动物以独特的生活方式适应或克服潮间带的各种不利条件而得到生存和繁衍。从报道来看现代海洋底内动物泂穴的研究具有多方面的意义。如某些潮间带底内动物的鉴定,仅以外部形态作依据显然不够,还需以其他生态资料,如粪便、洞穴、卵块等作参考。有些潮间带动物的地理分布范围与其营底内生活有关，它们可以借助洞穴的保护而渡过寒暖不适的气候。更重要的是这方面研究能对古生态学和地史学问題的探讨提供科学依据。有很多种潮间带底内动物洞穴、排出物和爬行痕迹等被作为“生活遗迹”而保存在化石中,因此这类化石的沉积岩层实际上已成为该类动物生存发展年代的记载。但是,在岩层古生态学分析研究中,比较困难的是如何区别古代生物居住、死亡或埋藏的地点,这个同题涉及到对沉积物形成的地点和过程的确定,进一步就可用这些资料来研究古代海岸变迁和海面的升降。就潮间带底内动物的居住和死亡地点(埋藏与死亡地点也可以不同)来说,有的可能一致(如某些不太活动的种类),有的则不一致(如贝类)，作为古生态学依据,各具其长短,贝类无洞穴化石,其贝壳可成为化石,但埋藏与生活地点往往不一致;蟹类的优点是洞穴能成化石，其上并留有爬行痕迹;洞穴化石的埋藏与动物原来生活地点一致(亦有个別情况不一致)。早在地史年代就已有不少虾蟹类遗体及生活迹迹化石被发现:因此，现代潮间带蟹类洞穴的研究不仅对认识或鉴定古代甲売类动物有价值,而且对进一步鉴定种类、寻找演化线索和促进古生态学的发展都具有重要意义。 青岛潮间带在我国北方沿岸潮间带中颇具代表性。潮汐是规则的半日潮。 著者曾于1954-1964年,选以下四个点进行了调查:1,沧口潮间带,位于胶州湾内东北区中部,座东向西，坡度小的泥质和泥沙质滩涂;2.沙子口潮间带,位于青岛以东,为沙质滩涂;3.湛山湾潮间带:位青岛以东近郊,为沙质滩涂;4.大黑澜潮间带,位于胶州湾口,为岩石岸滩。每月进行二次大潮采集,每一采样点取两个定量(50×50厘米)样品和若于定性补充样品。从1963年5月至1964年9月,结合甲壳类生态进行了专题调查,本文就是根据上述调查,在软底质滩涂所获11种蟹类材料总结的,对所见各种蟹类的洞穴均经现场观察、测量并作了详细记录和拍照。对同一种蟹类洞穴的形态在不同地点和时间内所出现的变化也给予了特别的注意。 本文定稿过程中征得了山东海洋学院李嘉泳及本所曾呈奎,刘瑞玉、郑执中、吴宝铃、齐钟彦、马绣同等同志的修改意见,文中插图、照片由王兴虞、宋华中同志给予帮助,在此一并表示感谢。     

Abundance,Tidal Movement,Population Structure and Burrowing Rate of Emerita analoga (Anomura,Hippidae) at a Dissipative and a Reflective Sandy Beach in South Central Chile

Abstract. To evaluate the effects of beach morphodynamics upon the abundance, tidal movement, population structure and burrowing rate of the crab Emerita analoga (Stimpson) (Anomura, Hippidae) we sampled two beaches in south central Chile (ca. 42° S), Mar Brava and Ahui with dissipative and reflective characteristics, res­pectively. The swash zone at the dissipative beach was 5 – 6 times wider than that of the reflective beach. A at the dissipative beach, upwash speeds were higher and the number of effluent line crossings were lower by more than an order of magnitude. To examine the tidal movement of E. analoga, we collected crabs from 5 to 6 tidal levels of each beach every 2 h across 12 h of the tidal cycle. The intertidal distribution of crabs differed between beaches; i. e., at the dissipative beach they were primarily located at the swash zone, while at the reflective beach they were mostly located at the low tide level and shallow subtidal. The change in position of crabs was pronounced across the tidal cycle at the dissipative beach (Mar Brava), with most of the animals remaining in the active swash zone. Body size data were used to construct size frequency distributions for each population. Crabs from the dissipative beach reached larger sizes than those at the reflective beach. Sediments were coarser at the latter versus the former beach. Crabs burrowed at similar rates in the sand from both beaches, a result which supports the idea that E. analoga is a “sediment generalist” capable of burrowing successfully in a wide range of sediment types. This characteristic is likely a key to the broad success of this species on the full range of beach morphodynamic types along the coasts of South and North America.     

Population surveys of toheroa (Mollusca: Eulamellffiranchiata) on Northland Beaches, 1962–67

Surveys made each year between 1962 and 1967 to estimate populations of toheroa (Amphidesma ventricosum Gray) on the three main west coast beaches of Northland, New Zealand, are described. The surveys were carried out as close as possible to the beginning and ending of each gathering season. There were striking changes in the surveyed areas, probably mainly due to natural causes and not as a direct result of gathering. One beach, Muriwai 1963, even showed an improvement in population after the gathering season, but numbers of toheroas on this beach have usually been erratic and small during the surveyed period. The other beaches showed marked declines, which were mostly sudden and unsynchronised between beaches; these declines were too big to be accounted for solely by gathering. A private commercial survey, carried out annually during the gathering season on a leased portion of Dargaville Beach, shows very similar results to the equivalent departmental surveys.     

Population Abundances, Tidal Movement, Burrowing Ability and Oxygen Uptake of Emerita analoga (Stimpson) (Crustacea, Anomura) on a Sandy Beach of South-Central Chile

Abstract. Field sampling and other experiments were carried out during February 2001 to determine whether different morphodynamic characteristics occurring within an intermediate sandy beach of southern Chile (ca. 39°S) convey differences in population abundance, tidal movement, burrowing ability and oxygen uptake of the anomuran crab Emerita analoga (Stimpson, 1857). Crabs were collected along transects extended between the lowest swash levels and the retention zone above the effluent line of the south and north end of the beach. Burrowing times of nearly 70 crabs collected at each study site were measured in saturated sands collected from the lowest swash level of each site. Oxygen uptake of crabs was measured in incubation glass bottles. The intertidal zone of the north end of the beach was wider (56 m) and flatter (1/14) than that of the south end (45 m and 1/9, respectively). In general, the swash zone of the north end was significantly wider than the south end throughout the sampling period. The frequency of swashes and number of swash crossings above the effluent line, plus up‐wash speed, were usually higher at the steeper south end of the beach. The mean population abundance of E. analoga per linear metre of beach was significantly higher at the north end, whereas density per square metre was significantly higher at the south end. No differences were found in biomass figures. Although the highest abundance of crabs at the north end was usually observed at the lowest swash levels, similar population abundances occurred along all the tidal levels sampled at the south end. Burrowing times of crabs collected from both ends of the beach increased significantly with increasing carapace length and body mass. The mean burrowing time of crabs collected at the south end of the beach was shorter than that of those collected at the north end. Oxygen uptake of E. analoga was positively and exponentially correlated with the size of individuals collected from both ends of the beach. Results of ANCOVA showed no significant difference between the regression lines obtained for the oxygen uptake of crabs collected at both ends of the beach. It is concluded that physical features of each end of the beach seem not to differ enough to produce differences in oxygen uptake of E. analoga, or in the biomass, population structure and body size of crabs, within a single beach of south‐central Chile.     

Variations in grain‐size and sorting on two kaikoura beaches

Two exposed, high‐energy beaches on the Kaikoura coast of New Zealand are composed of sand and gravel derived from a greywacke terrain. Both beaches can be classified as mixed beaches although the sediment varies from dominantly gravel at the ends of the beach to dominantly sand at the centre, through transition zones in which sand and gravel are mixed. Sixty‐four surface samples were analysed for grain size; two sediment parameters, mean grain size (M_z) and sorting (σ_I), were calculated.

A striking feature of the cumulative frequency curves is that both unimodai and bimodal distributions include median sizes over the whole range of sampled material, even though bimodal samples display two strong modes in the sand and gravel grades. The general deficiency lof sediment dn the very coarse sand and granule classes (0 to — 2 _F ) noted by numerous authors in many parts of the world is apparent in the poorly‐sorted bimodal samples. However, the best‐sorted samples also occur in these two classes.

Mean grain size of samples ranges from medium sand (1.820) to medium pebbles (—4.7 _F ), and sorting ranges from very well sorted (0.250) to very poorly sorted (2.69 _F ). Mean erain size on the northern beach is significantly greater than on the southern beach, but values of sorting are comparable. The greater mean size on one beach compared with the other is thought to be a function of the grade of material supplied by local rivers; the similarity in sorting presumably reflects the similarity of the processes acting on the two beaches.

Mixed sand‐shingle beaches are relatively rare on a world scale but common in New Zealand. Sediment distributions along the Kaikoura beaches do not reveal a regular decrease in size away from the rivers which supply material to shore at present. Instead, the beaches are differentiated into a number of sediment zones composed of either sand, or mixed sand‐gravel, or gravel. On each beach a gravel zone is located furthest from the river outlets. Sorting generally improves toward the Kaikoura Peninsula. Explanations for these trends are not given. Variations in size and sorting across the two beaches do not show a well developed zonation because of the high level of wave energy which continually mixes the material across the beach.     

Burrowing and burying by the crab Macrophthalmus hirtipes

Macrophthalmus hirtipes (Jacquinot, 1853) excavates burrows under water in soft, muddy sand that remains waterlogged at low tide. The crabs are immersed for at least 4 h in every tide. The burrows are usually single, blind‐ending, slightly curved passages; their openings do not appear to be deliberately closed by the crabs. Excavation is effected by using one set of walking legs as a scoop while the other set provides sideways traction.

Burrow openings are readily obscured by water movement, so that apparent burrow density does not give a true indication of the number of crabs in an area, but only indicates how many have been active recently; the true density may be found by sieving.

The burrowing activity of M. hirtipes is compared with that of other ocypodid crabs, and the functions of burrows are discussed. A distinction is made between excavating a burrow and rapid burying, because the two behaviours are very different, and must involve different adaptations and selection pressures.     

Population Dynamics of Pinnixa patagoniensis Rathbun, 1918 Brachyura: Pinnotheridae a Symbiotic Crab of Sergio mirim Thalassinidea: Callianassidae in Cassino Beach, Southern Brazil

Abstract. Pinnotherid crabs are typically found associated with many benthic invertebrates. On sandy beaches of southern Brazil, Pinnixa patagoniensis R athbun , 1918 inhabits burrows of the ghost shrimp Sergio mirim R odrigues , 1971. The population dynamics of this crab were studied from monthly samples taken between March 1989 and November 1991 near the Patos Lagoon outlet, southern Brazil ca. 32°13'S; 52°15'W. There was no correlation between the number and size of the crabs and the size of their respective hosts. Mean densities and incidence reached 3.1 indiv. opening^-1 range: 0 to 29 and 83.9%. respectively. Sex-ratios were not significantly different from 1:1 for the different size classes and sampling dates. Ovigerous females were found between October and March, showing an average fecundity of 1529 eggs per female. The annual recruitment pattern was bimodal, with a minor peak in summer and a fall peak that accounted for almost 70% of the population maintenance. Males and females both reached a maximum carapace width of 11.5 mm and had a life span of nearly two years as estimated from fitted von B ertalanffy growth curve parameters.     

Fish assemblage structure on sandy beaches with different anthropogenic influences and proximity of spawning grounds

Fish assemblages in an insular (preserved) and a continental (disturbed) sandy beach were compared to assess any changes that could be attributed to anthropogenic influences and/or the proximity of the fish spawning grounds. We expected that the closer geographical position to the spawning grounds and the small amount of anthropogenic disturbance on the insular beach would be likely to provide more suitable conditions for early fish development compared with the continental beach. A total of 192 samples (96 in each beach) were taken, yielding 68 fish species, mostly young‐of‐the‐year. Fish assemblage structure differed significantly between the two beaches. Moreover, the insular beach had higher number of species, number of individuals and biomass compared with the continental beach. The commercially important Clupeiformes Harengula clupeola, Anchoa tricolor and Anchoa januaria, Perciformes Micropogonias furnieri and Mugiliformes Mugil liza were typical species on the insular beach, partitioning the seasonal use of the beach. On the other hand, a few abundant non‐commercial species, mainly the Atheriniformes Atherinella brasiliensis and the Perciformes Eucinostomus argenteus and Diapterus rhombeus, occurred all year round at the continental beach. The high fish richness and abundance and the more conspicuous species turnover across seasons on the insular beach are probable indications of more complex and dynamic organization of the communities favored by better geographical position and less anthropogenic disturbance in the area.     

Impact of vehicles on benthic macrofauna on a subtropical sand beach

Our study aimed to identify possible relationships between the structure of the benthic macrofauna and the intense use of vehicles in Cassino Beach. Physical changes in the sediment were identified, with high compaction in the sector most affected by the vehicles, especially during the summer. The benthic macrofauna was significantly affected by vehicles, with lower densities observed in the most impacted sector, mainly in the summer, unlike the control sector, where higher densities were observed during the pre-summer and summer. We concluded that the vehicle traffic changes the physical environment of the beach and consequently affects the benthic community in the face of this disturbance. We conclude that the benthic macrofauna is highly impacted by vehicle traffic at Cassino Beach, mainly in the period with the highest traffic, and alternatives are needed for this impact, helping to recover this ecosystem.     

Are strandline meiofaunal assemblages affected by a once-only mechanical beach cleaning? Experimental findings

The increasing usage of sandy beaches as recreational resources has forced regional authorities of many tourist countries to remove all litter of fabricated origin and natural wrack from the beach. Consequently, a variety of heavy equipment has been developed during the last decades and is now used almost daily at many beaches. A field experiment, following a 'before-after-control-impact' (BACI) design, was conducted at the strandline of De Panne (Belgium) to investigate the impacts of mechanical beach cleaning on the strandline-associated meiofaunal assemblages, focussing on the free-living nematodes. Natural strandline assemblages were exposed to a one-off 5 cm deep mechanical beach cleaning and observed for 24 h. Differences between cleaned plots and those from control plots in terms of decreased percentage of organic matter, decreased total abundance and changed community structure were noticed from immediately after the experimental cleaning onwards and recovered to initial values after the following high water. Any impacts due to cleaning on species richness, Pielou's evenness and taxonomic diversity were shown to be minor in relation to the daily changes. Recolonization in the cleaned sediments is assumed to occur from the underlying sediments initiated by the elevated water table during the rising tide.     

Community and Population Responses of the Macroinfauna to Physical Factors over a Range of Exposed Sandy Beaches in South-central Chile

Ten exposed sandy sites covering a range from reflective to dissipative beaches were sampled in south-central Chile to evaluate: (1) spatial changes in species richness, abundance and biomass of the intertidal macroinfauna in response to changes in mean grain size, beach face slope and beach type, and (2) spatial changes in abundance, biomass and body sizes of the most abundant species in response to changes in the physical factors. The number of species, abundance and biomass per beach in general decreased with increasing particle size and beach face slope (steeper beaches) and increased from reflective to dissipative conditions. The best fit for number of species was with Dean's parameter, a measure of beach type, whereas for abundance and biomass the best fits were found with particle size. The isopod Excirolana braziliensis and the anomuran Emerita analoga increased in abundance and biomass towards dissipative conditions, whereas Excirolana hirsuticauda showed the opposite trend in biomass and was significantly larger in beaches with steeper profiles. It is concluded that responses to changes in beach type are more pronounced at community level than within species populations.     

台风季节朱家尖岛砂、砾质岬湾海滩的不同沉积地貌动态变化

岬湾砂、砾质海岸是海岛、海岸带重要的旅游资源, 具有较高的社会经济和生态价值, 长期以来备受关注。本文以浙江朱家尖岛东岸的5个代表性岬湾海滩为例, 基于2019年台风季节早期、中期及晚期测量获得的海滩地形和沉积物数据, 结合海滩近岸的水动力数据, 分析了砂质海滩和砾石海滩这两种不同类型岬湾海滩的沉积地貌动态变化。结果表明, 朱家尖岛东岸5个海滩在台风季节出现了不同的体积变化, 大沙里、东沙、千沙、乌石塘和小乌石塘海滩的单宽体积变化分别为11.93m³·m^-1、-54.41m³·m^-1、-19.75m³·m^-1、2.19m³·m^-1和-1.96m³·m^-1。砾石滩较砂质海滩更为稳定, 无人类活动干扰的砂质海滩在台风季节侵蚀更少、变化更小。台风季节大沙里、东沙、千沙、乌石塘和小乌石塘海滩表层沉积物的平均粒径分别为2.47Փ、2.24Փ、2.64Փ、-5.96Փ和-6.03Փ, 粒径粗化和离岸输运是5个海滩表层沉积物在台风季节的主要表现, 砂质海滩的沉积物粒度特征变化比砾石海滩要大。沉积物粒径、台风强度及台风期间的主要波向与海滩走向之间的关系、海岸工程这3种因素都可能对海滩在台风季节的沉积地貌动态变化产生影响。本文研究结果可为台风季节的海滩管理提供参考。     

The distribution and behaviour of tar balls along the Israeli coast

A fortnightly sampling of tar balls on six beaches along the Israeli coastline between 14 April 1975 and 25 June 1976 showed that the mean content of tar during that period was 3625 g m^?1 of beach front. The northern and central parts of the coast were significantly more polluted than the southern part. Between July 1975 and February 1976 the mean tar quantity decreased continuously from 5635 to 1344 g m^?1.A comparison of tar quantities on the Israeli beaches with those of other beaches in the world showed that the Israeli beaches are more polluted than those of the west Atlantic coast, are as polluted as other beaches on the eastern coast of the Mediterranean Sea, and are less polluted than the beaches of Alexandria, Egypt, and Paphos, Cyprus. It is suggested that the tar content on the beach is related to the degree of oil pollution in the sea. The closer a beach is to an oil shipping lane or an oil dumping site, the heavier it is polluted.During storms, beach tar balls are pushed by the waves to the back of the beach or, in the case of a cliffed coast, are carried along the shore by the longshore current. When the tar balls reach a gap in the cliff (such as an estuary), they are carried inland by the storm waves. There the tar may become buried or dry, shrink and break into small particles which are then dispersed by the wind.     

Assessing the response of sandy-beach macrobenthos to recreation and the ecological status of the beach ecosystem at Liandao,China

Sandy beaches, prime recreational assets that attract more people than other types of shorelines, are the main assets underpinning many coastal economies and developments. However, the intense stressors to the ecology of sandy beaches caused by recreational activities are generally underestimated or ignored. This study was the first to emphasize the ecological characteristics of recreational beaches in China and assess the response of macrobenthos at Liandao Beach, a typical sandy beach in the central part of China. Using the Before/During/After/Control/Impact (BDACI) method, differences in multiple variables (species richness, mean density, diversity) between a heavily recreationally impacted transect (TrH/Impact) and a less-impacted transect (TrL/Control) in different impact periods (Before, During & After) were applied to assess the response of sandy-beach macrobenthos to recreation and tourism. Meanwhile, a comparison between the macrobenthos conditions of the studied beach and reference conditions calculating from historical data of similar beaches with little recreation and tourism disturbances was carried out. Furthermore, the quality control chart approach was used to detect in detail the macrobenthos stable state and yearly response to recreational activities. The results showed that recreation and tourism posed a serious threat to the beach ecosystem; macrobenthos on the studied beach were seriously disturbed by recreational activities, and their variations were highly negatively related to the human recreational level. After monthly recreational impacts, the macrobenthos decreased sharply and recovered with difficulty during 1 year. Over the long-term, the macrobenthos had a sharp decreasing trend, which indicated that the ecological environment of the beach was highly disturbed. Conservation measures for recreational beaches are very challenging because of the economic value and popularity of sandy beaches as recreational sites. However, practicable conservation measures could be implemented that prevent or slow ecological degradation. A number of management and conservation suggestions are put forward, such as the establishment of the ecological carrying capacity of Liandao Beach in terms of direct recreational use, creation of themed exhibitions related to the beach macrobenthos and beach ecosystem and dissemination of this knowledge to visitors, implementation of spatial zoning of beach use, visitor management and environmental education.     

Threats to sandy beach ecosystems: A review

We provide a brief synopsis of the unique physical and ecological attributes of sandy beach ecosystems and review the main anthropogenic pressures acting on the world's single largest type of open shoreline. Threats to beaches arise from a range of stressors which span a spectrum of impact scales from localised effects (e.g. trampling) to a truly global reach (e.g. sea-level rise). These pressures act at multiple temporal and spatial scales, translating into ecological impacts that are manifested across several dimensions in time and space so that today almost every beach on every coastline is threatened by human activities. Press disturbances (whatever the impact source involved) are becoming increasingly common, operating on time scales of years to decades. However, long-term data sets that describe either the natural dynamics of beach systems or the human impacts on beaches are scarce and fragmentary. A top priority is to implement long-term field experiments and monitoring programmes that quantify the dynamics of key ecological attributes on sandy beaches. Because of the inertia associated with global climate change and human population growth, no realistic management scenario will alleviate these threats in the short term. The immediate priority is to avoid further development of coastal areas likely to be directly impacted by retreating shorelines. There is also scope for improvement in experimental design to better distinguish natural variability from anthropogenic impacts. Sea-level rise and other effects of global warming are expected to intensify other anthropogenic pressures, and could cause unprecedented ecological impacts. The definition of the relevant scales of analysis, which will vary according to the magnitude of the impact and the organisational level under analysis, and the recognition of a physical–biological coupling at different scales, should be included in approaches to quantify impacts. Zoning strategies and marine reserves, which have not been widely implemented in sandy beaches, could be a key tool for biodiversity conservation and should also facilitate spillover effects into adjacent beach habitats. Setback and zoning strategies need to be enforced through legislation, and all relevant stakeholders should be included in the design, implementation and institutionalisation of these initiatives. New perspectives for rational management of sandy beaches require paradigm shifts, by including not only basic ecosystem principles, but also incentives for effective governance and sharing of management roles between government and local stakeholders.     

The sandy beach macrofauna of Gulf of Gabès (Tunisia)

This study describes the macrofauna of the three beaches situated in central Gulf of Gabès (Tunisia): Ouderef, Gabès and Zarrat. The Gulf of Gabès has the largest tides in the Mediterranean and the beaches showed a wide intertidal zone. The beaches were sampled once during the spring low tides of June 2005. A transect was extended at each beach, from above the drift line to below the swash line at five sampled levels; at each level six 0.05 m² replicates were taken to a depth of 30 cm and sieved through a 1-mm mesh, and the organisms collected and preserved. The three beaches showed a different physical environment. Sediment type was medium sand at the steeper Ouderef beach, fine sands at Gàbes beach, and very fine sands at the flatter Zarrat beach. The total number of species collected was 31: 12 crustaceans, 10 polychaetes, four molluscs and five insects. The supralittoral and mediolittoral zones were very different. The supralittoral zone was dominated by Talitrus saltator and insects. The most abundant mediolittoral species were the amphipod crustacean Bathyporeia guilliamsoniana at Ouderef beach (23069 ind.m⁻¹), the surf clam Donax trunculus at Gabès beach (60711 ind.m⁻¹) and the spionid polychaete Scolelepis mesnili at Zarrat beach (18345.6 ind.m⁻¹).     

2010—2020及影响因素分析年青岛海滩剖面稳定性

海滩的稳定性及其演变是砂质海滩研究的重要内容。在全球砂质海滩普遍遭受侵蚀的背景下,分析部分稳定海滩的特征和原因可以为海滩维护和科学管理提供重要参考。本文基于近 11 年 （2010—2020 年） 青岛第一、第三和石老人海水浴场共 8 条典型剖面的实地监测高程数据,对海滩滩肩宽度和单宽体积进行计算对比,研究表明,除石老人海水浴场一个剖面变化较大外,其他剖面的滩肩宽度只变化了 0~14 m,是 11 年前的 0%~23.3%;单宽体积变化了 8.5~46.5 m3,是 11 年前的1.0%~9.4%;海滩剖面垂向扰动范围一般在 30~60 cm 范围内。海滩存在局部小尺度淤积或侵蚀,剖面形态总体较为稳定。海平面变化、台风和风暴潮等自然因素是海岸轻微侵蚀或调整的重要影响因素,但对海滩的优化管理 （包括定期监测海滩、适度展开养护、制定完善的管理政策和加强执行力度等人为因素） 可以应对这些海滩变化,并且已成为近年来该研究海滩保持稳定的关键因素。