A New Species of Huaxiapterus （Pterosauria： Tapejaridae） from the Early Cretaceous of Western Liaoning, China

A new species of Huaxiapterus： H. benxiensis sp. nov. is erected based on the new specimen. The diagnostic characters of Huaxiapterus benxiensis are well-developed premaxillary crest and parietal spine, the crest and spine parallel and extending posterodorsally, and a shallow groove present on the dorsal surface of the anterior portion of the mandibular symphysis. The different skull morphologies of Chinese tapejarid pterosaurs indicate that they are much more diverse than the previous thought.     

New Material of Feilongus (Reptilia: Pterosauria) from the Lower Cretaceous Jiufotang Formation of Western Liaoning

Based on a new nearly naturally preserved skull and four cervical vertebrae of the pterosaur Feilongus sp. from the lower Cretaceous Jiufotang Formation of Beipiao, western Liaoning province, northeastern China, the diagnosis of Feilongus is amended. The revised diagnosis notes long, curved, needle-shaped teeth that are confined to the jaw far anterior to the nasoantorbital fenestra; posterior teeth that are slightly smaller than the anterior teeth; cervical vertebrae elongated with a ratio of length to width greater than 5; tooth number of about 78; and two cranial sagittal crests.     

A New Species of Huaxiapterus （Pterosauria： Pterodactyloidea） from the Lower Cretaceous of Western Liaoning, China with Comments on the Systematics of Tapejarid Pterosaurs

A new species of tapejarid pterosaur, Huaxiapterus coroUatus sp. nov. is erected on the basis of a nearly complete skull and postcranial skeleton from the Lower Cretaceous Jiufotang Formation of Liaoning Province, China. Huaxiapterus coroUatus sp. nov. is characterized by a hatchet-shaped rectangular process on the premaxilla, whose short axis is perpendicular to the anterior margin of the premaxillae. Except for this process, other characters of the skull such as the breadth of the snout between the anterior margin of the nasoantorbital fenestra and the anterior margin of the premaxilla are similar to that of Huaxiapterus jii. Huaxiapterus and a second Chinese tapejarid, Sinopterus, share several unique cranial characters in common with Tapejara and these three genera appear to be more closely related to each other than to other azhdarchoids. The Chinese tapejarids （Sinopterus and Huaxiapterus） have relatively elongate skulls and weakly developed cranial crests and seem to be less derived than Tapejara, with its shorter, deeper skull and large cranial crest. Tupuxuarids （Tupuxuara and Thalassodromeus） have often been associated with tapejarids in the family Tapejaridae, but this relationship is controversial because some phylogenetic analyses have supported the pairing of tupuxuarids with Azhdarchidae. We propose that Tapejaridae be restricted to Tapejara, Sinopterus and Huaxiapterus.     

A New Ornithocheirid from the Early Cretaceous of Liaoning Province, China

Based on a nearly complete skeleton with skull from the Early Cretaceous of Liaoning Province, a new ornithocheirid pterosaur: Boreopterus cuiae gen. et sp. nov. is erected. Boreopterus cuiae is different from other pterosaurs preserved with skulls known from the western Liaoning Province and its neighboring areas. Tiffs new oterosaur has more and larger teeth than those in other ornithocheirids. Its anterior nine pairs of teeth are larger than other teeth. The fourth pair of upper and lower teeth are slightly larger than the third pair. Overall, Boreopterus cuiae shows much small range of tooth size variation than Anhanguerapiscator and Coloborhynchus robustus. The new taxon shares with other ornithocherids in having a relatively large size of the third and fourth pairs of teeth.     

New Material of Zhenyuanopterus (Pterosauria) from the Early Cretaceous Yixian Formation of Western Liaoning

A new partial specimen of Zhenyuanopterus from the Lower Cretaceous Yixian Formation of Beipiao city, Liaoning Province is described. This specimen is slightly larger than half the size of the holotype. Although it is not complete, it provides new information on Zhenyuanopterus both in taxonomy and ontogeny： the total number of the caudal vertebrae is 15; the forelimb is more robust than the hindlimb; the growth rate of the humerus and femur is constant, and their ratio is about 1. Compared with the holotype, the scapula and coracoid grow faster than the humerus and the femur. The scapula and coracoid may slow their growth rate after a certain ontogenetic stage. The pectoral girdle, robust forelimb and weak hindlimb of the new material（XHPM1088） together with data from the holotype of Zhenyuanopterus indicate that this taxon spent less time on land than in the sky or other ecological niches such as forests and cliffs.     

New Material of Sinosauropteryx （Theropoda： Compsognathidae） from Western Liaoning, Chinae

An incomplete specimen of Sinosauropteryx prima collected from the Early Cretaceous Yixian Formation of the Dawangzhangzi area in Lingyuan, western Liaoning is depicted. It represents the first systematically described material of this feathered compsognathid outside the Sihetun area in Beipiao. This specimen shows some diagnostic features of Sinosauropteryx prima, such as the very short forelimb in relation to hindlimb with the length ratio of humerus plus radius to femur plus tibia just around 30%, and the long and massive first manual ungual subequal in length to radius. The presence of Sinosauropteryx prima only at Sihetun and Dawangzhangzi supports the suggestion that the fossil-bearing beds in the Sihetun and Dawangzhangzi areas are equivalent to each other within the Yixian Formation.     

Large-Sized Ankylosaur (Dinosauria) from the Lower Cretaceous Jiufotang Formation of Western Liaoning, China

Two incomplete bones referred to ankylosaur scapulocoracoid and humerus are described, which were discovered from the Early Cretaceous Jiufotang Formation of Dachengzi Town, Kazuo County, Liaoning Province. Ankylosaur dinosaurs have ever been found in the Lower Cretaceous Yixian, Fuxin and Sunjiawan Formations of western Liaoning. The occurrence of ankylosaurs within Jiufotang Formation improves our knowledge on the ankylosaur stratigraphic distribution. Based on the length of scapulocoracoid, the new dinosaur is estimated to exceed 6.0 meters long, representing the largest ankylosaur in western Liaoning.     

A Nwe Hadrosauroid Dinosaur from the Mid—Cretaceous of Liaoning,China

A new hadrosauroid dinosaur,Shuangmiaosaurus gilmorei gen.et sp.nov,is descuibed based on a complete left maxilla with articulated premaxilla and lacrimal fragments,and a complete left dentary from the mid-Cretaceous Sunjiawan Formation of Beipiao.Liaoning,northeastern China.Chadistic analysis shows that Shuangmiaosaurus is a basal hadrosauroid,and comprises the sister taxon to Hadrosauridae.In both Shuangmiaosqurus and Hadrosaurids.However,Shuangmiaosaurus does not possess such hadrosaurid synapomorphies as the diamond-shaped maxillary crowns with reduced primary ridges and reduced marginal denticles.     

Pygostyle-like Structure from Beipiaosaurus (Theropoda, Therizinosauroidea) from the Lower Cretaceous Yixian Formation of Liaoning, China

Pygostyle was previously considered as a unique structure of ornithothoracine birds, used to maneuver tail feathers. A similar structure from an oviraptorosaurian dinosaur was considered functionally associated with the rectrices as in birds. We report a pygostyle-like structure from a therizinosauroid dinosaur. The presence of filamentous integuments, but absence of rectrices, on the tail of this therizinosauroid, combined with other lines of evidence, suggests that the initial function of the pygostyle was not related to the rectrices.     

Discovery of a New Pterosaur in Western Liaoning,China

The pterosaur from the Yixian Formation in Beipiao, Liaoning Province, ischaracterized by the medium-sized individual, short tail, presence of the gastralia, strongforelimbs, radius and ulna longer than wing-metacarpal, extremely narrow and elongatemetatarsals, degenerated and small phalanx V of hindlimb and so on. It must be a new genusand species, Eosipterus yangi gen. et sp. nov., in the early groups of the SuborderPterodactyloidea. This is the first record of pterosaurs found in Liaoning Province. The fossilprovides new evidence for further determining the nature and geological time of the JeholFauna and for studying the palaeogeography and palaeoenvironment.     

New Eoenantiornithid Bird from the Early Cretaceous Jiufotang Formation of Western Liaoning, China

1 Introduction Enantiornithes were the most blooming branch among the early birds and had distinct diversities. They flourished in the Early Cretaceous (Kurochkim, 1985; Chiappe, 1991; Sanz and Bonaparte, 1992; Boles, 1995; Elzanowski, 1995), and become extinct at the end of the Late Cretaceous. Lots of the birds have been reported in western Liaoning, China (Sereno and Rao, 1992; Zhou, 1995; Hou, 1996, 1997; Sanz and Chiappe, 1996; Sanz et al., 1997; Hou and Chen, 1999; Hou et al., 1…     

A New Enantiornithine Bird from the Lower Cretaceous Jiufotang Formation in Jinzhou Area,Western Liaoning Province,China

A new Early Cretaceous enantiornithine bird from Liaoning Province of northeastern China, Shengjingornis yangi, gen. et sp. nov., is reported. This new bird possesses the following unique combination of features: a long rostrum, with some teeth in the front; short nasal; slender jugal; Y-shaped furcula, with expanded distal end of the hypocleidum; cake-like sternum, with a low and caudally distributed keel; strut-like and caudally concave coracoid. The derived features of the scapula and the wings suggest a powerful flapping flight capability.     

An Early Cretaceous Avialian Bird, Shenzhouraptor sinensis from Western Liaoning, China

This paper describes an avialian bird from the Early Cretaceous Jiufotang Formation in Yixian county, western Liaoning, China, which is named as Shenzhouraptor sinensis by Ji et al. on July 15, 2002. Shenzhouraptor sinensis is characterized by no teeth in its mouth, the forelimbs longer than the hindlimbs, a long tail with more than 23 caudal vertebrae, U-shaped wishbone, and remiges longer than the total length of ulna and manus. It is certain that the new avialian bird is really capable of powerful flight, representing a missing link between theropod dinosaurs and birds.     

A New Ctenochasmatid Pterosaur from the Early Cretaceous of Liaoning Province

Based on a nearly complete lower jaw from the Early Cretaceous of Liaoning Province, a new ctenochasmatid pterosaur: Liaoxipterus brachyognathus gen, et sp. nov. is erected. Liaoxipterus brachyognathus is different from any known pterosaurs with skulls preserved from western Liaoning Province and its peripheral areas in that the anterior part of the mandibular symphysis is expanded, being widest between the fourth alveolus of each side. Liaoxipterus brachyognathus is assigned to Ctenochasmatidae based on the following characters: the rounded anterior end of the lower jaw is spatulated and dorsoventrally flattened and marked heterodonty in the dentition is absent. It differs from other ctenochasmatid pterosaurs in having relatively small number of teeth. Liaoxipterus is distinguished from some ornithocheirids, which have expanded anterior parts of the mandibular symphyses, such as Anhanguera piscator,Coloborhynchus robustus in which the teeth of the new pterosaur are not as variable.     

辽西朝阳地区下白垩统九佛堂组一新的离龙类

根据发现于辽宁省西部朝阳地区下白垩统九佛堂组的离龙类一具有头骨的不完整骨架，建立Simoedosauridae科中的一新属新种：朝阳辽西龙：Liaoxisaurus chaoyangensis gen．etsp．nov．。朝阳辽西龙的下颌缝合部短，小于下颌长度的20％，牙齿齿槽近似正方形，区别于任何已知的进步的离龙类。它吻部相对较短，占头骨总长的49．8％。     

New Tapejarid Pterosaur from Western Liaoning, China

A new tapejarid pterosaur Huaxiapterus jii gen. et sp. nov. is erected based on a nearly complete skeleton with a skull. The skull morphology of Huaxiapterus jii gen. et sp. nov. appears transitional between the skulls of Sinopterus and Tapejara. The skull morphology （low crest） of Huaxiapterusjii indicates that Huaxiapterus is more closely related to Sinopterus than to Tapejara, which has a high crest. The relatively long and shallow skull of Sinopterus indicates that it is a primitive form. Huaxiapterus is more derived than Sinopterus but more primitive than Tapejara.     

A New Troodontid (Theropoda: Troodontidae) from the Lower Cretaceous Yixian Formation of Western Liaoning, China

A specimen collected from the Lower Cretaceous Yixian Formation of western Liaoning, China, represents a new genus and species of troodontid theropod. The new taxon is named and described on the basis of the holotype and the only known specimen, which comprises an articulated skeleton with the presacral vertebral, shoulder girdle and forelimbs missing as preserved. Diagnostic features of the new species include nasals that are sinusoid in lateral view, absence of a passage connecting the antorbital and maxillary fenestrae, relatively large teeth, plate-like chevrons forming a band along most of the length of the tail, and a long neck between the femoral head and shaft. The temporal constraints of the three paravian groups (Troodontidae, Dromaeosauridae and Aves) combined with the character distributions among the earliest known troodontids indicate a rapid evolution at the base of the Troodontidae.     

New Azhdarchid Pterosaur from the Early Cretaceous of Western Liaoning

A nearly complete skeleton with a lower jaw of pterosaur from the Early Cretaceous of western Liaoning is described and assigned to a new genus, Eoazhdarcho gen. nov.. The new genus is characterized by a relatively small size,the ratio of the length to width of the middle series cervical vertebrae approximately 3.5 and the ratio of humeral length to femoral length approximately 0.96. The humerus of Eoazhdarcho shows great resemblances to that of previously described Azhdarchidae, so it is assigned to the family Azhdarchidae.