Grazing effects on patchy dryland vegetation in northern Patagonia

In this study the spatial patterns and dynamics of vegetation patches along a grazing gradient in the steppe ofLarrea divaricataandStipaspp. in NE Patagonia (Argentina) are described. A general effect of grazing is the reduction of total plant cover resulting from the decrease in cover of perennial grasses (Stipa speciosa, Poa ligularis, Stipa tenuis) and some tall shrubs (Chuquiraga hystrix, Bougainvillea spinosa, Lycium chilense). Dwarf shrubs (Nassauvia fueguianaandJunellia seriphioides) increase their cover under medium and/or high grazing pressures. Plant species are spatially grouped into patches which alternate with areas of bare soil. Eleven types of vegetation patch differing in the dominant plant functional type or species, floristic richness and size were identified with different relative frequency along the grazing gradient. Based on these results, it is postulated that grazing forces the replacement of large patches dominated by tall shrubs with high species richness, byLarrea divaricatapatches or small dwarf shrub patches with low species richness and the extinction of grass patches. This results from: (1) disruption of local balances of species deletions and additions; (2) fragmentation of large patches; and (3) formation of new vegetation patches. These changes lead to differing plant spatial organization and heterogeneity along the grazing gradient which may be described by characteristic arrays of vegetation patches.     

Short-term effects of shrub control on two different plant communities in Argentina

The aim of this work was to study the effect of controlling jarilla (Larreaspp.), nonforage shrubs for livestock, on two plant communities with different forage production in the arid mid-west of Argentina. Total vegetation cover, forage species cover and carrying capacity were determined during three growing seasons, for cleared and uncleared plots. At the end of the study, significant differences were found in theLarrea cuneifoliacommunity, where average forage species cover increased by 156% over the control, and greater carrying capacity (129%) was also obtained from the cleared treatment. No significant differences were found in theLarrea divaricatacommunity. Data lead us to recommend shrub control as an improvement practice only in areas with low forage species cover and poor carrying capacity.     

Aquatic macrophyte distribution in Lake Manzala,Egypt

The macrophyte distribution of Lake Manzala is described in relation to water depth, salinity, dissolved oxygen, pH, Cl^–, NO ₃ ^– and PO ₄ ^3– . Changes in species composition of macrophytes are visualised by means of multivariate analysis. The TWINSPAN classification and CANOCO ordination programs (DCA & CCA) were used to analyse the data set. The classification of 100 stands revealed 8 vegetation groups which indicated eleven dominant communities. These arePhragmites australis, Typha domingensis, Scirpus maritimus, Echinochloa stagnina and Ludwigia stolonifera as emergent hydrophytes;Eichhornia crassipes andAzolla filiculoides as floating hydrophytes. The dominant submerged hydrophytes arePotamogeton pectinatus, Najas armata, Ceratophyllum demersum andRuppia maritima. The northern part of the lake with low depth and relatively high salinity has low species diversity (mainly emergent species). Species diversity increases with decreasing salinity and increasing eutrophication near the mouths of the drains in the western and southern parts of the lake. The recent changes in species distribution can be attributed to the effects of salinity, water depth and drainage water. A checklist of macrophytes in the lake is appended.     

Vegetation and soil characteristics of the wasteland of Valika Chemical Industries near Manghopir, Karachi

The vegetation of the wasteland of Valika chemical industries near Manghopir road, Karachi was studied. Nine plant communities were recognized based on dominant species. In these plant communities the vegetation was disturbed, mostly halophytic and dominated bySuaeda fruticosa,Tamarix indica,Salsola baryosma,Cressa cretica,Atriplex griffithii,Haloxylon recurvum,Indigofera hochstetteri,Prosopis julifloraandChenopodium album.The physico-chemical properties of the soils were also analysed. Soil texture was mostly sandy loam, which exhibited slight variations in the water-holding capacity. The soils contained a sufficient amount of CaCO₃⁻and exhibited mostly alkaline soil pH. The soils of the different plant communities had scarcely any organic matter or inorganic phosphorus. The exchangeable sodium in the soils of many halophytic plant communities was high, with appreciable concentrations of potassium.     

A comparison of classification systems for the conservation of sparsely wooded plains on Melville Island,Northern Australia

A floristic quadrat study of sparsely wooded plains on western Melville Island was conducted and used to define and describe floristic and structural vegetation types and identify their associated environmental correlates. The results were compared with an existing land‐unit survey of the area in order to identify the relative merits of these schemes for conservation planning. All schemes identify a range of broad structural types including Acacia shrubland, Grevillea pteridifolia low woodland, Lophostemon lactifluus low woodland, and Banksia dentata low woodland. However, some of the structural and land‐unit types are composed of two plant communities with distinctive environmental relations. Despite being structurally distinctive, some community types are floristically indistinguishable from surrounding forest dominated by Eucalyptus miniata and E. tetrodonta. The number of species captured by randomly selecting sites from each classificatory type revealed little difference between schemes. All the classificatory approaches were less effective at species capture than a simple iterative procedure. Some of the plant communities seem to be unique and require preservation initiatives in the face of current forestry operations and planned horticultural development.     

Spatial and temporal distribution of tenebrionid species (Coleoptera) in the Negev Highlands, Israel

Tenebrionid species composition and activity level were studied, using pitfall traps, in two sandy (dune and stabilized sand field) and four compact soil (wadi, north- and south-facing slopes, and loess plain) habitats in the Negev Desert Highlands, Israel. Each soil type had a distinctive species composition but habitats within soft types were similar both in species composition and phenologies. The number of species was not correlated with habitat primary production; it was low in sandy habitats (19 spp.) and high in compact soil (33 spp.) habitats. Within the compact soil habitats, large-size species were abundant only in the high plant-cover wadi, and the low plant-cover slopes and plain were dominated by smaller species. We suggest that low plant cover on the slopes and plain exposes large tenebrionids to predation by birds and limits them to the wadi where they are concealed and find refuge in the plant cover.     

Ant communities and livestock grazing in the Great Basin, U.S.A.

The objectives of this study were to determine if metrics for ant species assemblages can be used as indicators of rangeland condition, and to determine the influence of vegetation and ground cover variables, factors often influenced by livestock grazing, on ant communities. The study was conducted in two areas in the Great Basin: a sagebrush-steppe in southeastern Idaho (n=30 sites), and a salt-desert shrub in western Utah (n=27 sites). Sites were selected based on known rangeland condition (i.e. good, fair, poor) associated with livestock grazing. Ant communities differed considerably between the two study areas. Collectively, more ant species occurred at the Idaho sites (30) than at the Utah sites (21), relatively few species (eight) occurred in both areas, species richness was significantly greater at the Idaho sites (mean=12·0 species) than the Utah sites (mean=6·9 species), and Formica spp. were diverse (total of 15 species) at the Idaho sites but rare (one species) at the Utah sites. In Idaho, all species collectively, generalists, and Formica spp. were significantly less abundant on sites in poor condition than that on sites in good or fair condition, whereas in Utah, seed harvesters and Pheidole spp. were significantly more abundant on sites in poor condition than that on sites in good or fair condition. In Idaho, species richness was significantly lower on sites in poor condition. In Idaho, species richness and relative abundances of several ant groups were significantly related to bare patch size and parameters for cover or species richness of several vegetation groups. In contrast to the comparisons involving sites in poor condition, no differences in ant communities in either Idaho or Utah were evident between sites in good and fair condition. Thus, the ant communities responded only to large changes in rangeland condition and to large differences in climatic/edaphic conditions between the two areas. Hence, ant community metrics appear to have limited utility as indicators of rangeland condition in the Great Basin.     

The current status ofAtriplexon the Cape Verde Islands

Extensive areas of low lying coastal plains on Santiago Island, Republic of Cape Verde, are virtually devoid of natural vegetation. Of many introduced trees and shrubs, onlyAtriplexspecies have been found to thrive. Several plantations have been established during the past 15 years by the Cape Verde Government, FAO and other agricultural development groups. Results from these, together with local perceptions, indicate that although growth and survival are good, the primary objective of providing forage is not being met, owing to the unpalatability ofAtriplexto local stock. In recent field trials, severalAtriplexspecies showed good growth, but none were browsed and survival was quite poor, with onlyA. undulata(65%) andA. canescens(100%) exceeding 60% survival. Forage analysis identified high salt content in all species tested, and the highest protein content was found inA. glauca(16%),A. vesicaria(17%) andA. canescens(20%), which also had the lowest salt levels.Atriptex canescenswas identified as worthy of further introduction. An establishedA. halimusplantation that was observed to be browsed was compared to unbrowsed stands in terms of forage, soil and local drinking water analyses. Possible reasons for the unpalatability ofAtriplexto local stock and its resolution are discussed, along with the potential future ofAtriplexon the Cape Verde Islands.     

Small-scale vegetation dynamics in semi-arid steppe in Inner Mongolia

Small-scale vegetation dynamics were followed for ecotones and in uniform stands inArtemisia-dominated steppe vegetation under grazing and when recovering from heavy grazing. Species composition was followed annually for 5 years in 1 m²and 0·25 m²plots for (1) presence–absence; (2) density; and (3) biomass.More rapid vegetation dynamics, in terms of change of type of vegetation and distance moved in DCA species space between sampling occasions, were observed in smaller plots and in early seral stages, where species were few and alpha diversity low. Only the plots recently protected from grazing showed a directional vegetation change; those protected for more than 3 years and those under continued grazing showed, at this scale, non-directional dynamics.     

Acacia communities and species responses to soil and climate gradients in the Sudano-Sahelian zone of West Africa

Abiotic environmental factors have a major impact on the distribution and performance of plant species. In order to assess two major species–environmental relations in Sudano-Sahelian Acacia woodlands, we tested the relationship of soil and climate variables on plant diversity as well as on species responses.The indicator species values clustered in five vegetation units characterized by three to ten diagnostic species with woody species richness means varying from three to seven species per 0.09 ha. The NMS ordination explained 65% of the variation in species composition and revealed that soil properties, annual precipitations and temperature range structured the diversity of Acacia communities. Along the annual precipitations gradient, the response of Acacia polyacantha and Acacia hockii showed maxima in the wettest zone of our study area (more than 850 mm/year) whilst Acacia laeta showed a maximum response in the driest zone (below 500 mm/year). The unimodal response of A. hockii, Acacia gourmaensis and Acacia seyal to the soil available water gradient spanned their central borders, respectively from 13 to 18% (optimum 16%), 11–20% (optimum 15%) and 4–12% (optimum 7.5%).The response of Acacia communities and species to soil and climate gradients, makes them performant afforestation species in specific habitats of the Sudano-Sahelian zone.     

Restoration of palatable grasses: A study case in degraded rangelands of central Argentina

Restoration of palatable grasses on degraded rangelands dominated by unpalatable grasses in central Argentina is limited by low availability of seeds and safe sites for seedling establishment. The objective of our study was to determine how mechanical disturbance of unpalatable grasses (Stipa spp.) in combination with seeding of a palatable grass (Poa ligularis) influenced species composition in a degraded rangeland excluded from livestock grazing. In April 2001 10 blocks were uniformly distributed on a previously burned site dominated by unpalatable grasses, and treatments applied in 8 m×8 m experimental plots. Treatments were ‘disked and seeded’ and control (no disking, no seeding). Perennial plant cover and end-of-season standing crop, at species or species group level, were assessed in December 2002/2003 and in December 2004/2006, respectively. P. ligularis out-competed both tillers of unpalatable grasses that survived mechanical disturbance and seedlings of unpalatable grasses established after mechanical disturbance. The cover and end-of-season standing crop of unpalatable grasses was higher in the control than in the ‘disked and seeded’ treatment, whereas the cover and end-of-season standing crop of P. ligularis was higher in the latter than in the former treatment. Our results suggest that a rapid transition from a state dominated by unpalatable grasses to a state dominated by palatable grasses can be achieved by mechanical disturbance of unpalatable grasses in combination with seeding of palatable grasses in semiarid rangelands of central Argentina.     

Use of large Acacia trees by the cavity dwelling Black-tailed Tree Rat in the Southern Kalahari

Recent extensive harvesting of large, often dead Acacia trees in arid savanna of southern Africa is cause for concern about the conservation status of the arid savanna and its animal community. We mapped vegetation and nests of the Black-tailed Tree Rat Thallomys nigricauda to assess the extent to which the rats depend on particular tree species and on the existence of dead, standing trees. The study was conducted in continuous Acacia woodland on the southern and eastern edge of the Kalahari, South Africa. Trees in which there were tree rat nests were compared with trees of similar size and vigour to identify the characteristics of nest sites. Spatial analysis of tree rat distribution was conducted using Ripley's-L function. We found that T. nigricauda was able to utilize all available tree species, as long as trees were large and old enough so that cavities were existing inside the stem. The spatial distribution of nest trees did not show clumping at the investigated scale, and we therefore reject the notion of the rats forming colonies when inhabiting continuous woodlands. The selection of a particular tree as a nest site was furthermore depending on the close proximity of the major food plant, Acacia mellifera. This may limit the choice of suitable nest sites, since A. mellifera was less likely to grow within a vegetation patch containing a large trees than in patches without large trees.     

Species diversity and regeneration of native species in Pennisetum ciliare (buffelgrass) pastures from the thornscrub of Sonora,México

To examine the influence of buffelgrass land conversion and pasture management on native species diversity and regeneration patterns, we describe community attributes and population structure in four different active pastures in thornscrub vegetation from eastern Sonora, Mexico. We compare a relatively undisturbed thornscrub community with a contiguous five years old active pasture, to identify species able to regenerate under current management practices. Buffelgrass conversion has a significant influence on species diversity and other community attributes. Active pastures have lower species diversity, crown cover and basal area of native species than the studied native thornscrub. An active pasture subjected to heavy grazing and without management of the thorny legume, Acacia cochliacantha, showed the lower species diversity and an almost monospecific stand of this legume. It is likely that the reduction in species diversity is caused by the conversion process, inadequate management and regeneration barriers that limit seedling establishment in active pastures. Our data recorded eight native species (13%) that were able to regenerate in active pastures. However, the great majority of native species were unable to regenerate under pastures. This study shows that under current management, active buffelgrass pastures maintain only a small fraction of native species with regenerating populations in the thornscrub.     

The representation of diatom communities by fossil assemblages in a small acid lake

The representative quality of fossil diatom assemblages in the recent sediment of a lake is compared with its contemporary diatom flora. In April 1986 experimental liming of the catchment of a small acidified lake, Loch Fleet (Galloway, U.K.), produced immediate changes in water quality. Lakewater pH rose from a mean of approximately 4.5 to 6.5, and in the two year period following liming a consistently higher pH was maintained. The marked response of diatom species to changing water quality provided a means of tracing events from living communities to the fossil assemblages. Diatom periphyton and plankton were sampled during a 20 month period and archived material was used to characterise earlier diatom communities. A comparison is made between living diatom communities and diatom assemblages collected by sediment traps and from sediment cores taken during the same period.Following liming, the diatom communities were found to respond within days or weeks to the changes in water quality. There is an initial change from acidobiontic communities, dominated byTabellaria quadriseptata, to dominance by the acidophilous speciesEunotia incisa andPeronia fibula. However, in the epipsammic community the acidobiontic speciesTabellaria binalis fo.elliptica remains abundant after liming. Approximately one year after liming the abundances of species such asAchnanthes minutissima andBrachysira vitrea increase in the epilithon, epiphyton and epibryon, whilst in the epipsammonT. binalis fo.elliptica is replaced by smallEunotia spp. andAchnanthes altaica. During the latter part of 1987 and in 1988, despite a stable pH, fluctuating patterns of species abundances are seen in the epilithon, epiphyton and epibryon whilst the species composition of the epipsammon remains relatively stable. Spring blooms of the planktonic speciesSynedra acus andAsterionella formosa occur during 1988 and 1989 respectively.Sediment trapping, which began in April 1987, records shifts in species composition corresponding with those seen in the epilithon, epiphyton and epibryon and with the blooms of planktonic species. The signal from the smaller, and probably less easily transportable, epipsammic community is not so clearly discernible. Although the fundamental record of the sediment traps is one from living diatom communities, the appearance of taxa extinct during the post-liming period reflects a low, but significant level of sediment resuspension.In contrast to the rapid response of living communities and their record in sediment traps, sediment cores do not begin to reflect changes in diatom composition until about 14 months after the initial liming. The first appearance of circumneutral taxa in significant abundance occurs only approximately 17 months after liming. The delayed reaction of sediment assemblages cannot be attributed principally to a slow rate of transport from the littoral to the profundal zone. Time-averaging processes within the sediment appear to be the main cause of the lag in core response. In contrast, blooms of planktonic species are quickly reflected in the stratigraphy of cores, but indicate that a considerable degree of downward mixing occurs. Comparison of the time trajectories of whole species assemblages in living communities, sediment traps and core surface sediments shows that the direction of change is similar in all three, but that the magnitude of change is attenuated in sediment assemblages.     

Assessment of grazing effect on sheep fescue (Festuca valesiaca) dominated steppe rangelands, in the semi-arid Central Anatolian region of Turkey

In the semi-arid steppe rangelands of Central Turkey, Festuca valesiaca and Thymus sipyleus ssp rosulans have become the dominant species on degraded pastures. We hypothesized that decreases in species richness and abundance are correlated with increasing prevalence of these two species. Therefore, our objectives were to determine whether there are patterns in examined vegetation; how dominant species contribute to these patterns; and how patterns differ between grazed and ungrazed vegetation. We determined that protection from grazing increased species richness. Grazing significantly changed composition through decreasing total plant, forb, grass and F. valesiaca covers, while substantially increasing T. sipyleus cover. Topography, soil and grazing appear to impact the dominance of plant communities where F. valesiaca and T. sipyleus prevail. These two dominant species had a significant effect in shaping vegetation patterns. Based on regression analysis, alterations in species richness with changes in cover of forbs and shrubs were evident, and spatial heterogeneity of F. valesiaca and T. sipyleus indicated unstable vegetative patterns in heavily grazed pastures and successional changes in protected pastures. Our study results identify F. valesiaca and T. sipyleus as indicator species of vegetation suppression in condition assessments of degraded steppe rangelands.     

新疆阜康绿洲荒漠过渡带植物群落物种多样性特征

分别采用重要值、盖度、多度计算多样性指数、均匀度指数、丰富度指数、优势度指数，对新疆阜康荒漠地区植物群落的物种多样性进行了研究。结果表明：1)各类型群落的物种多样性水平表现为红柳、红砂、梭梭这三个群落较高，肉叶雾冰藜群落、叉毛蓬群落、盐爪爪群落、囊果碱蓬群落和雾冰藜群落居中，无叶假木贼群落和碱蓬群落较低；2)研究区总体群落物种多样性水平较低，随着群落演替其物种多样性水平呈现低—高—较高的变化趋势；3)对荒漠植被，以重要值或盖度为指标计测多样性指数较为可行。     

Microfungal community associated with rhizosphere soil of Zygophyllum qatarense in arid habitats of Bahrain

Population density and species diversity of microfungal communities were investigated in the rhizosphere soil of the halophytic plant Zygophyllum qatarense inhabiting saline and nonsaline habitats of the arid desert environment of Bahrain. Unlike the nonsaline habitat which is situated in the physiographic zone of multiple escarpment and backslopes, the saline site is located in the coastal lowlands and is featured by high chloride content, electrical conductivity, total soluble salts and low organic matter. Soils of both habitats are sandy, slightly alkaline, poor in nutrient sources, low in water-holding capacity and mainly dominated by a salt-tolerant flora. Quantification of data for the recovery of fungi were based on colony identification and counts by a series of ten-fold dilutions plate method, using various natural, synthetic and selective media. A total of 2780 isolates, fluctuating between 25 and 1109 per sample, were recovered during the present study among all habitats, seasons and plant sizes. Grouping of these isolates has resulted in a maximum of 28 fungal taxa varied between 5 and 15 species, of which 24 were hyphomycetes, 3 ascomycetes and one was an unknown species. All the recorded species in this study, excluding the genus Fusarium, are newly reported from the arid terrestrial habitats of Bahrain. Of the encountered fungi,Cladosporium sphaerospermum was the most dominant and frequent genus, among all plant sizes, followed by Penicillium citrinam and Aspergillus fumigatus, a finding with consistent documented data from similar arid Sahara ecosystems. Examination of data, supported by analysis of relative density values, percentage recovery rates, polar ordination and diversity indices revealed that the nonsaline habitat during the dry period yielded the highest isolate frequency, species abundance, and diversity when compared with the saline habitat. Moreover, a progressive increase in colony occurrence and species diversity was equivalently associated with increment in plant size in the nonsaline habitat. Apparently, the vast majority (moderate to low occurrence class) of the reported species are rhizosphere indigenous saprophytic cellulose-decomposers, whilst the sparse taxa (high occurrence class), e.g. C. sphaerospermum, are regarded as thermo-osmotolerant. Comparison of species richness among samples suggests that small plants inhabiting nonsaline habitats during the wet and dry season were richest in species composition. It is proposed that inter- and intra-specific variation in fungal community between the above habitats reflect not only the influence of plant age and season but also extends to critical multi-soil edaphic and biotic factors involving essentially soil moisture, salinity and root growth dynamic and exudates.     

Desertification in the Eastern Province of Saudi Arabia

The problem of land degradation in the Eastern Province of Saudi Arabia is not a new phenomenon, but during the last decade it has grown to be a serious threat to the delicate desert ecosystems, and also to the urban and industrial areas, due to an increase in sand movement. The area under investigation is situated on the Gulf coast north of Jubail and is approximately 20 km in width centered along the coastline. The total area is almost 1000 km². Eight vegetation types were classified according to the dominant and co-dominant species. A comparative study of the vegetation in a fenced, non-grazed area and in an open grazed area showed that the impact of grazing and recreational activities have largely reduced the value of the rangelands. The reduction in species diversity and density leads to an increase in wind and sand dynamics. Exposed roots, giant ripples, changes in the extension of inland sabkhas and new active dunes could be observed in many parts of the study area. It could be proved that most inland sabkhas in the area are pure deflational features with considerable dynamic due to the reduction in vegetation cover during the recent years. The monitoring of vegetation density was carried out during helicopter flights with the help of GPS and a video camera. Image processing on a personal computer allowed accurate calculations of vegetation density. Observations on active dune fields in the area showed that the size of the dune fields almost doubled in 15 months. Tests in small enclosures showed that there is a potential for natural recovery even in severely damaged areas. In just one year without grazing and with average precipitation there is a significant difference in vegetation cover. Strict grazing management and camping regulations are required to prevent further degradation and allow the recovery of damaged ecosystems. Community involvement and education is necessary for the development of an understanding of regulations and the need for preservation.     

Fire in Chihuahuan Desert grassland: Short-term effects on vegetation, small mammal populations, and faunal pedoturbation

A prescribed burn resulted in significant decreases in canopy cover of the grasses: Bouteloua eriopoda, Sporobolus flexuosus, and Aristida purpurea. One year post-burn, basal cover of B. eriopoda remained significantly lower in burned patches than in unburned areas but there were no differences in basal cover of the other perennial grasses. Only one species of the 14 summer annual species occurred in both burned and unburned plots. There were six species of spring annuals in burned patches but no spring annuals in the unburned grassland ten months post-burn. Fire killed 100% of the snakeweed shrubs (Gutierrezia sarothrae), 77% of the Ephedra torreyana shrubs, and 36% of the Yucca elata. All mesquite shrubs that were top-killed by fire, resprouted one month post-burn. Fire had no effect on abundance and species richness of rodents. There were fewer wolf spider, Geolycosa spp. burrows in burned areas than in unburned grassland. The area and volume of soil in termite galleries and sheeting were significantly larger in the unburned grassland than in the burned areas.     

Diet selection in captivity by a generalist herbivorous rodent (Octodon degus) from the Chilean coastal desert

Five captive adult female degus (Octodon degus) were offered leaves and twigs to eat from three woody (Adesmia bedwellii,Porlieria chilensisandProustia pungens) and two suffruticose (Baccharis paniculataandChenopodium petiolase) shrubs that provide cover in their natural habitat. The degus discriminated among the plant species, consuming lower amounts ofP. chilensis. Daily body mass losses of degus were significantly higher when they were fed uponP. pungensandP. chilensis. The nutritional value of plants, concomitant with degu nutritional requirements, may explain changes in shrub cover previously found to follow removal of degus from experimental plots.