陆地生态系统模型及其与气候模式耦合的回顾

陆地生态系统和气候系统通过能量通量、水汽通量、物质交换相互影响、作用。作者对陆地生态系统模型及其与气候模式耦合的研究进行综述和讨论,总结了当代5类主要全球陆地生态系统模型,即生物地理模型、生物地球化学模型、森林林窗模型、陆面生物圈模型和动态全球植被模型,以及它们与气候模式耦合的研究进展。阐述了动态全球植被模型及其与气候模式耦合研究在全球变化研究的重要作用。最后,对未来模拟研究的方向进行了分析。     

Boreal forest and tundra ecosystems as components of the climate system

The effects of terrestrial ecosystems on the climate system have received most attention in the tropics, where extensive deforestation and burning has altered atmospheric chemistry and land surface climatology. In this paper we examine the biophysical and biogeochemical effects of boreal forest and tundra ecosystems on atmospheric processes. Boreal forests and tundra have an important role in the global budgets of atmospheric CO₂ and CH₄. However, these biogeochemical interactions are climatically important only at long temporal scales, when terrestrial vegetation undergoes large geographic redistribution in response to climate change. In contrast, by masking the high albedo of snow and through the partitioning of net radiation into sensible and latent heat, boreal forests have a significant impact on the seasonal and annual climatology of much of the Northern Hemisphere. Experiments with the LSX land surface model and the GENESIS climate model show that the boreal forest decreases land surface albedo in the winter, warms surface air temperatures at all times of the year, and increases latent heat flux and atmospheric moisture at all times of the year compared to simulations in which the boreal forest is replaced with bare ground or tundra. These effects are greatest in arctic and sub-arctic regions, but extend to the tropics. This paper shows that land-atmosphere interactions are especially important in arctic and sub-arctic regions, resulting in a coupled system in which the geographic distribution of vegetation affects climate and vice versa. This coupling is most important over long time periods, when changes in the abundance and distribution of boreal forest and tundra ecosystems in response to climatic change influence climate through their carbon storage, albedo, and hydrologic feedbacks.     

陆地生态系统与全球变化相互作用的研究进展

全球变化及其对生态系统特别是陆地生态系统的影响已经严重地影响到人类生存环境与社会经济的可持续发展 ,引起了各国政府、科学家及公众的高度关注。文中从CO2 浓度倍增、温度变化、水分变化、水热与CO2 协同作用、辐射变化、臭氧变化以及人为干扰等气候环境变化对植物光合生理、生长发育、物质分配、水分利用、碳氮代谢等的影响方面阐述了全球变化影响生态系统的过程与机理 ;从地理分布范围、物候、结构与功能、生态系统的稳定性等方面分析了中国植被、森林生态系统、草原生态系统与农田生态系统对全球变化的响应 ;从植被变化引起的动力条件与热力条件的变化及植被固碳潜力的变化探讨了植被对于气候的反馈作用。在此基础上 ,基于当前全球变化研究前沿 ,提出了未来关于陆地生态系统与全球变化相互作用研究需要重视的方面 ,尤其是关于生态系统对全球变化响应的阈值研究应引起高度重视。     

Changes in Global Vegetation Distribution and Carbon Fluxes in Response to Global Warming: Simulated Results from IAP-DGVM in CAS-ESM2

Terrestrial ecosystems are an important part of Earth systems, and they are undergoing remarkable changes in response to global warming. This study investigates the response of the terrestrial vegetation distribution and carbon fluxes to global warming by using the new dynamic global vegetation model in the second version of the Chinese Academy of Sciences (CAS) Earth System Model (CAS-ESM2). We conducted two sets of simulations, a present-day simulation and a future simulation, which were forced by the present-day climate during 1981–2000 and the future climate during 2081–2100, respectively, as derived from RCP8.5 outputs in CMIP5. CO₂ concentration is kept constant in all simulations to isolate CO₂-fertilization effects. The results show an overall increase in vegetation coverage in response to global warming, which is the net result of the greening in the mid-high latitudes and the browning in the tropics. The results also show an enhancement in carbon fluxes in response to global warming, including gross primary productivity, net primary productivity, and autotrophic respiration. We found that the changes in vegetation coverage were significantly correlated with changes in surface air temperature, reflecting the dominant role of temperature, while the changes in carbon fluxes were caused by the combined effects of leaf area index, temperature, and precipitation. This study applies the CAS-ESM2 to investigate the response of terrestrial ecosystems to climate warming. Even though the interpretation of the results is limited by isolating CO₂-fertilization effects, this application is still beneficial for adding to our understanding of vegetation processes and to further improve upon model parameterizations.     

Carbon Sequestration in Arable Soils is Likely to Increase Nitrous Oxide Emissions,Offsetting Reductions in Climate Radiative Forcing

Strategies for mitigating the increasing concentration of carbon dioxide (CO₂) in the atmosphere include sequestering carbon (C) in soils and vegetation of terrestrial ecosystems. Carbon and nitrogen (N) move through terrestrial ecosystems in coupled biogeochemical cycles, and increasing C stocks in soils and vegetation will have an impact on the N cycle. We conducted simulations with a biogeochemical model to evaluate the impact of different cropland management strategies on the coupled cycles of C and N, with special emphasis on C-sequestration and emission of the greenhouse gases methane (CH₄) and nitrous oxide (N₂O). Reduced tillage, enhanced crop residue incorporation, and farmyard manure application each increased soil C-sequestration, increased N₂O emissions, and had little effect on CH₄ uptake. Over 20 years, increases in N₂O emissions, which were converted into CO₂-equivalent emissions with 100-year global warming potential multipliers, offset 75–310% of the carbon sequestered, depending on the scenario. Quantification of these types of biogeochemical interactions must be incorporated into assessment frameworks and trading mechanisms to accurately evaluate the value of agricultural systems in strategies for climate protection.     

Response of the mean global vegetation distribution to interannual climate variability

The impact of interannual variability in temperature and precipitation on global terrestrial ecosystems is investigated using a dynamic global vegetation model driven by gridded climate observations for the twentieth century. Contrasting simulations are driven either by repeated mean climatology or raw climate data with interannual variability included. Interannual climate variability reduces net global vegetation cover, particularly over semi-arid regions, and favors the expansion of grass cover at the expense of tree cover, due to differences in growth rates, fire impacts, and interception. The area burnt by global fires is substantially enhanced by interannual precipitation variability. The current position of the central United States’ ecotone, with forests to the east and grasslands to the west, is largely attributed to climate variability. Among woody vegetation, climate variability supports expanded deciduous forest growth and diminished evergreen forest growth, due to difference in bioclimatic limits, leaf longevity, interception rates, and rooting depth. These results offer insight into future ecosystem distributions since climate models generally predict an increase in climate variability and extremes. CCR Contribution # 941     

多圈层陆面过程参数化研究中遥感信息应用的进展和方向

近年来遥感技术的发展为多圈层中陆面过程和边界层研究提供了有力的工具。文章分析了目前用于陆面过程参数化研究的重要遥感信息源。并评述遥感信息在陆面过程参数化研究中的基本应用和存在问题，最后，提出发展方向和展望。     

Terrestrial biosphere carbon storage under alternative climate projections

This study investigates commonalities and differences in projected land biosphere carbon storage among climate change projections derived from one emission scenario by five different general circulation models (GCMs). Carbon storage is studied using a global biogeochemical process model of vegetation and soil that includes dynamic treatment of changes in vegetation composition, a recently enhanced version of the Lund-Potsdam-Jena Dynamic Global Vegetation Model (LPJ-DGVM). Uncertainty in future terrestrial carbon storage due to differences in the climate projections is large. Changes by the end of the century range from −106 to +201 PgC, thus, even the sign of the response whether source or sink, is uncertain. Three out of five climate projections produce a land carbon source by the year 2100, one is approximately neutral and one a sink. A regional breakdown shows some robust qualitative features. Large areas of the boreal forest are shown as a future CO₂ source, while a sink appears in the arctic. The sign of the response in tropical and sub-tropical ecosystems differs among models, due to the large variations in simulated precipitation patterns. The largest uncertainty is in the response of tropical rainforests of South America and Central Africa.     

Historical and future quantification of terrestrial carbon sequestration from a Greenhouse-Gas-Value perspective

Terrestrial ecosystems provide a range of important services to humans, including global and regional climate regulation. These services arise from natural ecosystem functioning as governed by drivers such as climate, atmospheric carbon dioxide mixing ratio, and land-use change. From the perspective of carbon sequestration, numerous studies have assessed trends and projections of the past and future terrestrial carbon cycle, but links to the ecosystem service concept have been hindered by the lack of appropriate quantitative service metrics. The recently introduced concept of the Greenhouse Gas Value (GHGV) accounts for the land-atmosphere exchanges of multiple greenhouse gases by taking into consideration the associated ecosystem pool sizes, annual exchange fluxes and probable effects of natural disturbance in a time-sensitive manner.We use here GHGV as an indicator for the carbon sequestration aspects of the climate regulation ecosystem service, and quantify it at global scale using the LPJ-GUESS dynamic global vegetation model. The response of ecosystem dynamics and ecosystem state variables to trends in climate, atmospheric carbon dioxide levels and land use simulated by LPJ-GUESS are used to calculate the contribution of carbon dioxide to GHGV. We evaluate global variations in GHGV over historical periods and for future scenarios (1850–2100) on a biome basis following a high and a low emission scenario.GHGV is found to vary substantially depending on the biogeochemical processes represented in LPJ-GUESS (e.g. carbon–nitrogen coupling, representation of land use). The consideration of disturbance events that occur as part of an ecosystem's natural dynamics is crucial for realistic GHGV assessments; their omission results in unrealistically high GHGV. By considering the biome-specific response to current climate and land use, and their projections for the future, we highlight the importance of all forest biomes for maintaining and increasing biogeochemical carbon sequestration. Under future climate and carbon dioxide levels following a high emission scenario GHGV values are projected to increase, especially so in tropical forests, but land-use change (e.g. deforestation) opposes this trend. The GHGV of ecosystems, especially when assessed over large areas, is an appropriate metric to assess the contribution of different greenhouse gases to climate and forms a basis for the monetary valuation of the climate regulation service ecosystems provide.     

The interactive climate and vegetation along the Pole-Equator belts simulated by a global coupled model

The interaction between climate and vegetation along four Pole-Equator-Pole （PEP） belts were explored using a global two-way coupled model, AVIM-GOALS, which links the ecophysiological processes at the land surface with the general circulation model （GCM）. The PEP belts are important in linking the climate change with the variation of sea and land, including terrestrial ecosystems. Previous PEP belts studies have mainly focused on the paleoclimate variation and its reconstruction. This study analyzes and discusses the interaction between modern climate and vegetation represented by leaf area index （LAI） and net primary production （NPP）. The results show that the simulated LAI variation, corresponding to the observed LAI variation, agrees with the peak-valley variation of precipitation in these belts. The annual mean NPP simulated by the coupled model is also consistent with PIK NPP data in its overall variation trend along the four belts, which is a good example to promote global ecological studies by coupling the climate and vegetation models. A large discrepancy between the simulated and estimated LAI emerges to the south of 15°N along PEP 3 and to the south of 18°S in PEP 1S, and the discrepancy for the simulated NPP and PIK data in the two regions is relatively smaller in contrast to the LAI difference. Precipitation is a key factor affecting vegetation variation, and the overall trend of LAI and NPP corresponds more obviously to precipitation variation than temperature change along most parts of these PEP belts.     

An estimate of equilibrium sensitivity of global terrestrial carbon cycle using NCAR CCSM4

Increasing concentrations of atmospheric CO₂ influence climate, terrestrial biosphere productivity and ecosystem carbon storage through its radiative, physiological and fertilization effects. In this paper, we quantify these effects for a doubling of CO₂ using a low resolution configuration of the coupled model NCAR CCSM4. In contrast to previous coupled climate-carbon modeling studies, we focus on the near-equilibrium response of the terrestrial carbon cycle. For a doubling of CO₂, the radiative effect on the physical climate system causes global mean surface air temperature to increase by 2.14 K, whereas the physiological and fertilization on the land biosphere effects cause a warming of 0.22 K, suggesting that these later effects increase global warming by about 10 % as found in many recent studies. The CO₂-fertilization leads to total ecosystem carbon gain of 371 Gt-C (28 %) while the radiative effect causes a loss of 131 Gt-C (~10 %) indicating that climate warming damps the fertilization-induced carbon uptake over land. Our model-based estimate for the maximum potential terrestrial carbon uptake resulting from a doubling of atmospheric CO₂ concentration (285–570 ppm) is only 242 Gt-C. This highlights the limited storage capacity of the terrestrial carbon reservoir. We also find that the terrestrial carbon storage sensitivity to changes in CO₂ and temperature have been estimated to be lower in previous transient simulations because of lags in the climate-carbon system. Our model simulations indicate that the time scale of terrestrial carbon cycle response is greater than 500 years for CO₂-fertilization and about 200 years for temperature perturbations. We also find that dynamic changes in vegetation amplify the terrestrial carbon storage sensitivity relative to a static vegetation case: because of changes in tree cover, changes in total ecosystem carbon for CO₂-direct and climate effects are amplified by 88 and 72 %, respectively, in simulations with dynamic vegetation when compared to static vegetation simulations.     

Modeling Dynamic Vegetation Response to Rapid Climate Change Using Bioclimatic Classification

Modeling potential global redistribution of terrestrial vegetation frequently is based on bioclimatic classifications which relate static regional vegetation zones (biomes) to a set of static climate parameters. The equilibrium character of the relationships limits our confidence in their application to scenarios of rapidly changing climate. Such assessments could be improved if vegetation migration and succession would be incorporated as response variables in model simulations. We developed the model MOVE (Migration Of VEgetation), to simulate the geographical implications of different rates of plant extirpation and in-migration. We used the model to study the potential impact on terrestrial carbon stocks of climate shifts hypothesized from a doubling of atmospheric greenhouse gas concentration. The model indicates that the terrestrial vegetation and soil could release carbon; the amount of this carbon pulse depends on the rate of migration relative to the rate of climate change. New temperate and boreal biomes, not found on the landscape today, increase rapidly in area during the first 100 years of simulated response to climate change. Their presence for several centuries and their gradual disappearance after the climate ceases to change adds uncertainty in calculating future terrestrial carbon fluxes.     

Transient climate changes in a perturbed parameter ensemble of emissions-driven earth system model simulations

We describe results from a 57-member ensemble of transient climate change simulations, featuring simultaneous perturbations to 54 parameters in the atmosphere, ocean, sulphur cycle and terrestrial ecosystem components of an earth system model (ESM). These emissions-driven simulations are compared against the CMIP3 multi-model ensemble of physical climate system models, used extensively to inform previous assessments of regional climate change, and also against emissions-driven simulations from ESMs contributed to the CMIP5 archive. Members of our earth system perturbed parameter ensemble (ESPPE) are competitive with CMIP3 and CMIP5 models in their simulations of historical climate. In particular, they perform reasonably well in comparison with HadGEM2-ES, a more sophisticated and expensive earth system model contributed to CMIP5. The ESPPE therefore provides a computationally cost-effective tool to explore interactions between earth system processes. In response to a non-intervention emissions scenario, the ESPPE simulates distributions of future regional temperature change characterised by wide ranges, and warm shifts, compared to those of CMIP3 models. These differences partly reflect the uncertain influence of global carbon cycle feedbacks in the ESPPE. In addition, the regional effects of interactions between different earth system feedbacks, particularly involving physical and ecosystem processes, shift and widen the ESPPE spread in normalised patterns of surface temperature and precipitation change in many regions. Significant differences from CMIP3 also arise from the use of parametric perturbations (rather than a multimodel ensemble) to represent model uncertainties, and this is also the case when ESPPE results are compared against parallel emissions-driven simulations from CMIP5 ESMs. When driven by an aggressive mitigation scenario, the ESPPE and HadGEM2-ES reveal significant but uncertain impacts in limiting temperature increases during the second half of the twenty-first century. Emissions-driven simulations create scope for development of errors in properties that were previously prescribed in coupled ocean–atmosphere models, such as historical CO₂ concentrations and vegetation distributions. In this context, historical intra-ensemble variations in the airborne fraction of CO₂ emissions, and in summer soil moisture in northern hemisphere continental regions, are shown to be potentially useful constraints, subject to uncertainties in the relevant observations. Our results suggest that future climate-related risks can be assessed more comprehensively by updating projection methodologies to support formal combination of emissions-driven perturbed parameter and multi-model earth system model simulations with suitable observational constraints. This would provide scenarios underpinned by a more complete representation of the chain of uncertainties from anthropogenic emissions to future climate outcomes.     

Climate change alters growing season flux dynamics in mesic grasslands

Changing climate could affect the functioning of grassland ecosystems through variation in climate forcings and by altering the interactions of forcings with ecological processes. Both the short and long-term effects of changing forcings and ecosystem interactions are a critical part of future impacts to ecosystem ecology and hydrology. To explore these interactions and identify possible characteristics of climate change impacts to mesic grasslands, we employ a low-dimensional modeling framework to assess the IPCC A1B scenario projections for the Central Plains of the United States; forcings include increased precipitation variability, increased potential evaporation, and earlier growing season onset. These forcings are also evaluated by simulations of vegetation photosynthetic capacity to explore the seasonal characteristics of the vegetation carbon assimilation response for species at the Konza Prairie in North Central Kansas, USA. The climate change simulations show decreases in mean annual soil moisture and and carbon assimilation and increased variation in water and carbon fluxes during the growing season. Simulations of the vegetation response show increased variation at the species-level instead of at a larger class scale, with important heterogeneity in how individual species respond to climate forcings. Understanding the drivers and relationships behind these ecosystem responses is important for understanding the likely scale of climate change impacts and for exploring the mechanisms shaping growing season dynamics in grassland ecosystems.     

An Analysis of Sensitivity of Terrestrial Ecosystems in China to ClimaticChange Using Spatial Simulation

A computer simulation model of regional vegetationdynamics was applied to the terrestrial ecosystems ofChina to study the responses of vegetation to elevatedCO₂ and global climatic change. The primaryproduction processes were coupled with vegetationstructure in the model. The model was parameterizedand partially validated in light of a large number of fieldobservations made throughout China on primary productivity,10 years of monthly meteorological data, 5 years of monthlynormalized differential vegetation index observed byNOAA-11 satellite, and digital vegetation and terrainmaps. Eight different climatic scenarios, set byperturbations from the present climate, 100% inatmospheric CO₂ concentration, 2 °C inmonthly mean temperature, and 20% in monthlyprecipitation, were applied to analyze the sensitivityof the Chinese terrestrial ecosystems to climaticchange. Simulation results were obtained for each ofthe climatic scenarios with the model running towardequilibrium solutions at a time step of 1 month.Preliminary validation indicated that the model wascapable of simulating the net primary productivity ofmost vegetation classes and the potential vegetationstructure in China under present climatic conditions.The simulations for the altered climatic scenariospredicted that grasslands, shrubs, and conifer forestsare more sensitive to environmental changes thanevergreen broadleaf forests in warm, wet southeastChina and desert vegetation in cold, arid northwestChina. For less than 150% of changes in vegetationstructure under altered climatic conditions, aboutthree quarters of the changes in net primaryproductivity of individual vegetation classes wereshown to be attributed to the changes in thecorresponding distribution area.     

Effects of Dynamic Vegetation on Global Climate Simulation Using the NCEP GFS and SSiB4/TRIFFID

Two global experiments were carried out to investigate the effects of dynamic vegetation processes on numerical climate simulations from 1948 to 2008.The NCEP G...     

Natural and anthropogenic climate change: incorporating historical land cover change,vegetation dynamics and the global carbon cycle

This study explores natural and anthropogenic influences on the climate system, with an emphasis on the biogeophysical and biogeochemical effects of historical land cover change. The biogeophysical effect of land cover change is first subjected to a detailed sensitivity analysis in the context of the UVic Earth System Climate Model, a global climate model of intermediate complexity. Results show a global cooling in the range of –0.06 to –0.22 °C, though this effect is not found to be detectable in observed temperature trends. We then include the effects of natural forcings (volcanic aerosols, solar insolation variability and orbital changes) and other anthropogenic forcings (greenhouse gases and sulfate aerosols). Transient model runs from the year 1700 to 2000 are presented for each forcing individually as well as for combinations of forcings. We find that the UVic Model reproduces well the global temperature data when all forcings are included. These transient experiments are repeated using a dynamic vegetation model coupled interactively to the UVic Model. We find that dynamic vegetation acts as a positive feedback in the climate system for both the all-forcings and land cover change only model runs. Finally, the biogeochemical effect of land cover change is explored using a dynamically coupled inorganic ocean and terrestrial carbon cycle model. The carbon emissions from land cover change are found to enhance global temperatures by an amount that exceeds the biogeophysical cooling. The net effect of historical land cover change over this period is to increase global temperature by 0.15 °C.     

基于PML-V2模型和站点观测数据的植被水利用率及其趋势差异分析

运用基于Penman-Monteith公式改进得到的模型PML-V2,结合12个FLUXNET站点及其对应的叶面积指数数据,进行蒸散发分离,进而计算并分析内禀水利用率(intrinsic water use efficiency, iWUE)和冠层水利用率(canopy water use efficiency, tWUE)的趋势差异。结果表明,在站点尺度上,两种植被水利用率的变化均存在不一致性。对于落叶阔叶林(deciduous broadleaf forests, DBF),i WUE的增幅比tWUE的增幅大,而在常绿针叶林(evergreen needleleaf forests, ENF)中则相反。在DBF中,冠层导度和蒸腾作用趋势的差异可在一定程度上解释两种植被水利用率的趋势差异。通过回归分析发现森林(包括DBF和ENF)的气温和大气CO₂浓度的趋势对tWUE趋势的影响更大。研究结果表明,两种植被水利用率及其趋势存在差异。基于iWUE的研究结果并不能完全反映植被的实际水利用率变化程度,因此也不能全面反映植被与大气的相互作用。本文在站点尺度明确了全球气候变...     

Estimation of critical levels of climate change influence on the natural terrestrial ecosystems on the territory of Russia

Developed are the axiomatics and criteria for estimating the critical levels of climate change influence on the natural terrestrial ecosystems based on the revelation of key climate-dependent environmental elements and model analysis of their variations. Developed is an empirical statistical vegetation model for the territory of Russia considering 15 vegetation zones including five ones in the permafrost zone. The model was used to estimate the proximity of the climate impact on the natural terrestrial ecosystems to the critical level for several climate projections.