中国海域头足类物种多样性

本文回顾了中国头足类物种多样性研究历史,分析了头足类的种类组成、分布特点,结果显示中国海域头足类记录的有效现存种达154种,隶属于34科79属：鹦鹉螺目1科1属1种,乌贼目3科12属37种,闭眼鱿目1科4属13种,开眼鱿目17科38属47种,八腕目8科20属51种,幽灵蛸目1科1属1种,未定目3科3属4种。渤海、黄海、东海和南海的头足类种类组成存在显著差异性,呈自北(渤海)向南(南海)逐渐增多的趋势：渤海9种,黄海15种,东海91种,南海140种。结合上述研究结果和国内外研究现状、发展趋势,对深入开展头足类多样性研究提出了建议。     

秋季黄海中南部鱼类群落对饵料生物的摄食量

研究鱼类与饵料生物之间食物定量关系进而为多鱼种资源评估提供依据,2000~2002年秋季(10~11月)在黄海中南部海域进行了定点底拖网调查,应用Eggers模型,计算了带鱼(Trichiurus lepturus)、小黄鱼(Pseudosciaena polyactis)、黄(Lophius litulon)、细纹狮子鱼(Liparis tanakae)等23种鱼类在秋季对饵料生物的摄食量。结果表明:黄海中南部23种鱼类在秋季对饵料生物的总摄食量约为309万t,其中,鱼(Engraulis japonicus)的摄食量最高(在250万t以上),占总摄食量的80.9%。中上层和底层鱼类对饵料生物的摄食量分别为262万t和47万t左右,占总摄食量的84.7%和15.3%,鱼和细纹狮子鱼分别是中上层和底层鱼类中最主要的捕食者。磷虾类是中上层鱼类最主要的食物来源,其次是桡足类、端足类和毛颚类;虾类和鱼类是底层鱼类最主要的食物来源,其次是磷虾类。太平洋磷虾(Euphausia pacifica)、中华哲水蚤(Calanus sinicus)、细长脚虫戎(Themisto gracilipes)、脊腹褐虾(Crangon affinis)和鱼同时是黄海中南部被摄食量最高的5种饵料生物,它们被摄食的生物量之和约为233万t,占总摄食量的75.5%。     

东海南部海域头足类群落结构特征及其与环境关系

根据2008年春(5月)、夏(8月)、秋(11月)、冬(2009年2月)四季东海区桁杆拖虾网调查期间所采集的头足类资料,分析了东海南部海域头足类种类组成及优势种,采用生态多样性指数分析了群落物种多样性,并运用典范对应分析法研究了头足类群落变化与主要环境因子的关系。结果表明:东海南部海域共捕获头足类22种,隶属于3目6科8属,优势种主要有神户乌贼(Sepia kobiensis)、双喙耳乌贼(Sepiola birostrata)、短蛸(Octopus ocellutus)和杜氏枪乌贼(Loligo duvaucelii)等;种类丰富度指数D变化范围为0—1.30,多样性指数H′变化范围为0—1.69,均匀度指数J变化范围为0.02—1.00,t检验表明,各群落多样性指数四季之间的差异均不显著(P0.05);根据典范对应分析认为,底层水温为影响调查海域头足类种类组成的主要环境因子。     

东海中南部秋季游泳动物多样性及群落结构

根据2018年11月东海中南部游泳动物底拖网调查数据,对该海域游泳动物种类组成、优势种、种类多样性及群落结构进行研究。结果表明,调查共捕获游泳动物181种,其中鱼类133种,甲壳类34种,头足类14种;优势种（IRI≥500）为七星底灯鱼（Benthosema pterotum）、带鱼（Trichiurus lepturus）、麦氏犀鳕（Bregmaceros mcclellandi）、龙头鱼（Harpadon nehereus）和日本发光鲷（Acropoma japonicum）。多样性指数结果显示,东海中南部海域丰度（d）的变化范围为0.37～5.94,种类多样性指数（H′）的变化范围为0.38～2.64,均匀度（J）的变化范围为0.14～0.71。根据聚类分析结果将东海中南部海域游泳动物群落分为以90 m等深线为界的两个群组,群组间的差异主要由七星底灯鱼、镰鲳（Pampus echinogaster）、龙头鱼、黄鲫（Setipinna tenuifilis）、麦氏犀鳕等引起。BIOENV结果显示水深与游泳动物群落结构相关性最密切（r_w=0.55）。本研究结果可为类似海域游泳动物种类多样性和群落结构的研究提供参考,也可为研究海域渔业资源开发、利用与养护等政策的制定提供科学依据。     

黄海鱼类群落分类学多样性的研究

为开展不同取样区域,不同生境间和历史数据的比较研究,根据1985年和2000年秋季在黄海进行的渔业资源底拖网调查,应用分类学多样性指数从黄海、区域和局域3个空间尺度上分析了黄海鱼类群落分类学多样性的变化。在黄海尺度上,2000年等级多样性指数△^ 和分类差异变异指数∧^ 明显降低,而1985年的分类学多样性指数值则接近于黄海鱼类主名录的理论平均值。在区域尺度上,分类多样性指数△,分类差异指数△^ 在黄海中、北部较高,南黄海较低;等级多样性指数△^ 为南黄海最高,中黄海最低,其区域尺度变化主要是由鲈形目、鲽形目、鲱形目和鲉形目种类,特别是软骨鱼类的种类数的分布变化造成的。2000年3个区域的分类学多样性指数值分别低于1985年对应值。在局域尺度上,南黄海的南部、西部和中黄海的西部以及北黄海的鱼类群落等级多样性指数△^ 远小于黄海鱼类主名录的理论平均值,仅中昔海的东部和黄海的东部的等级多样性指数值接近理论平均值。     

舟山渔场及邻近海域头足类(Cephalopod)种类组成和数量分布

根据2006年8月、2007年1月、5月和11月4个季节在舟山渔场及邻近海域(29°30'N-32°00'N,127°00'E以西)开展海洋生态系统综合调查所获得的头足类资料,用渔获率作为头足类分布的数量指标,分析了该海域的头足类种类组成、数量分布以及季节变化趋势.结果表明,该海域共鉴定头足类25种,隶属于3目8科9属...     

东、黄海2000年冬季底层鱼类群落结构及其多样性

根据2000年12月～2001年1月东、黄海冬季底层鱼类资源调查资料,分析了该海域的鱼类群落结构。本次调查共捕获鱼类4716.72kg,395796尾,138种。其中,暖水种(WW)居第1位,有81种,占60.00%;暖温种(WT)次之,有53种,占39.26%;冷温种(CT)最少,只有1种,占0.74%。全调查海域的优势种(相对重要性指数QIRI大于500)有5种:带鱼、黄鲫、发光鲷、小黄鱼和鳀;与历史资料相比,优势种个体变小、有被小型鱼类更替的趋势。以质量计算的多样性指数(种类丰富度指数D,Shannon-Wiener多样度指数H′W,种类均匀度指数J′W)呈现由北至南逐渐递增的趋势,而以尾数计算的多样性指数(H′N,J′N)呈现东海北部最高,东海中部次之,黄海南部最低的趋势。生物多样性指数中除种类丰富度指数(D)与水深、海水表温、表层盐度都呈显著性正相关外,而其它生物多样性指数与前述3种环境因子之间的相关性都不显著。单因素方差分析表明,东海北部和中部渔获组成和生物多样性指数都无显著性差别。     

广东流沙湾游泳动物种群结构和生物多样性

在流沙湾湾口设置双桩张纲张网对海湾游泳动物进行16次采样调查,结果表明:流沙湾有游泳动物161种,隶属3纲13目55科,其中:鱼类9目43科100种,虾类2目5科32种,蟹类1目4科20种,头足类2目3科9种,鱼类种类8月初最多,1月初最少;虾类种类10月底最多,1月底最少;蟹类种类8月初最多。16次采样共捕获虾类604121尾,1024kg;鱼类201339尾,1344.7kg;蟹类15115尾,49.5kg;头足类947尾,9.9kg。游泳动物生物量最高值出现在4月底、5月初和8月底,最低值出现在1月底和12月底。游泳动物优势种依次为刀额新对虾、沙栖新对虾、多鳞鱚、近缘新对虾、短棘银鲈、孔鰕虎鱼、中型新对虾、短吻鲾、刀额仿对虾、黑鳃梅童鱼、角突仿对虾、东方箬鳎、黄斑蓝子鱼和银光梭子蟹。流沙湾游泳动物丰富度指数变化较大,多样性指数、优势度值、均匀度值变化不大。流沙湾游泳动物的种群变动和水温关系密切,在水温29—32°C时的种群生物量最大。     

采用碳氮稳定同位素技术对黄海中南部鳗鱼食性的研究

根据2005年4～5月在黄海和长江口海域进行的春季底拖网调查，应用稳定同位素方法研究了黄海中南部鯷鱼（Engraulis　japonicus）及其可能摄食饵料的碳氮稳定同位素比值，结果表明：（1）黄海中南部鯷鱼的食物组成为不同粒径的浮游动物、太平洋磷虾（Euphausia pacifuca）和仔稚鱼，其中以粒径为500-900μm的浮游动物为主，贡献比例占61％～84％；仔稚鱼贡献比例占16％～21％，较传统胃含物方法分析的结果小；（2）黄海中部海域鯷鱼的碳氮稳定同位素比值平均值较南部海域高，原因可能与各海域的能量来源不同或存在微食物环有关，也可能与不同海域鯷鱼的能量转换途径不一样，即与食物链长短不一有关．     

南黄海春秋季大型底栖动物分布现状

根据 2 0 0 0年 1 0月和 2 0 0 1年 3月在 1 2 1°— 1 2 5°E、33°— 36°N海域范围内的 1 7个取样站采集的 482号采泥样品和 5 4 6号拖网样品共计 2 72种动物 ,经实验室鉴定、称重、计数、计算 ,分析研究了南黄海海域大型底栖动物在春季和秋季的分布状况。结果表明 ,在 2 72种大型底栖生物中 ,多毛类 84种 ,软体动物 76种 ,甲壳动物 83种 ,棘皮动物 2 9种。从分析结果看 ,春季和秋季大型底栖生物的栖息密度没有明显的差异 ,而生物量却变化很大 ,秋季的生物量远高于春季。这与底栖生物在春季繁殖 ,多为幼体或低龄个体 ,而秋季基本长大成熟有关。与 1 95 9— 1 960年的全国海洋综合调查对南黄海底栖生物的调查结果比较 ,表明当前该海域大型底栖生物的分布格局和生态学特点没有根本的改变 ,但春季的生物量较 1 95 9年低     

春、秋季南黄海浮游纤毛虫丰度及生物量的分布差异

Seasonal variation of marine plankton spatial distribution is important in understanding the biological processes in the ocean.In this study,we studied spatial distribution of planktonic ciliate abundance and biomass in the central deep area(station depth greater than 60 m) and the coastal shallow area(station depth less than 60 m) of the southern Yellow Sea(32°–36.5°N,121°–125°E) in spring(April) and autumn(October–November) of 2006.Our results showed that both ciliate abundance and biomass in the surface waters were higher in spring((1 490±2 336)ind./L;(4.11±7.81) μg/L) than in autumn((972±823) ind./L;(1.11±1.18) μg/L,calculated by carbon).Ciliate abundance and biomass in the surface waters of the coastal shallow area were similar in spring and autumn.However,in the central deep area,those values were much higher in spring((1 878±2 893) ind./L;(5.99±10.10)μg/L) than in autumn((738±373) ind./L;(0.74±0.76) μg/L).High values of ciliate abundance and biomass occurred in the central deep area in spring and in the coastal shallow area in autumn.Mixotrophic ciliate Laboea strobila was abundant in the central deep area in spring,when a phytoplankton bloom occurred.However,in autumn,L.strobila was abundant in the coastal shallow area.Boreal tintinnid Ptychocyli obtusa was found in spring.Both L.strobila and P.obtusa were concentrated in the surface waters when their abundance was more than 1 000 ind./L.Peaks of these species were in the subsurface waters when their abundance was less than 400 ind./L.This study showed that both high abundance and biomass of ciliates occurred in different areas in southern Yellow Sea seasonally.     

Studies on the species composition and biodiversity of free-living marine nematodes in the southern Huanghai Sea

1 IntroductionThe conservation, persistentuse and basic re-search ofbiodiversity are increasingly becom ing oneofthe centraltopics concerned by internationalaca-dem iccirclesandothergroupsdueto itshugevaluetom ankind living and suffered seriousthreats. Fr…     

南黄海和中国东海中挥发性卤代烃的分布与海气通量

Distributions and sea-to-air fluxes of five kinds of volatile halocarbons(VHCs) were studied in the southern Yellow Sea(SYS) and the East China Sea(ECS) in November 2007. The results showed that the concentrations of 1,1,1-trichloroethane(C2H3Cl3), 1,1-dichloroethene(C2H2Cl2), 1,1,2-trichloroethene(C2HCl3), trichloromethane(CHCl3) and tetrachloromethane(CCl4) in the surface water were 0.31–4.81, 2.75–21.3, 1.21–17.1, 5.02–233 and 0.045–4.47 pmol/L, respectively, with the average values of 1.89, 12.20, 6.93, 60.90 and 0.33 pmol/L. On the whole, the horizontal distributions of C2H3Cl3, C2H2Cl2 and CCl4 were affected mainly by anthropogenic activities, while C2HCl3 and CHCl3 were influenced by biological factors as well as anthropogenic activities. In the study area, the concentrations of VHCs(except C2HCl3) exhibited a decreasing trend from inshore to offshore sites, with the higher values occurring in the coastal waters. The sea-to-air fluxes of C2H3Cl3, C2HCl3, CHCl3 and CCl4 were calculated to be-56.00–(-5.68),-7.31–123.42, 148.00–1 309.31 and-83.32–(-1.53) nmol/(m2·d), respectively, with the average values of-6.77, 17.14, 183.38 and-21.27 nmol/(m2·d). Our data showed that the SYS and ECS in autumn was a sink for C2H3Cl3 and CCl4, while it was a source for C2HCl3 and CHCl3 in the atmosphere.     

Distribution of phytoplankton in the East China Sea and the southern Yellow Sea in spring in relation to environmental variables and dimethylsulfide compounds

The coastal ecosystems are highly sensitive to climate change and are usually influenced by variations in phytoplankton communities and water physiochemical factors. In the present study, the phytoplankton community, chlorophyll a（Chl a） and their relationships with environmental variables and dimethylsulfide（DMS） and dimethylsulfoniopropionate（DMSP） were investigated in spring 2017（March 24 to April 16） in the East China Sea（26.0°–33.0°N, 120.0°–128.0°E） and southern Yellow Sea（31.0°–36.0°N, 12...     

黄渤海区捕捞结构的研究

根据2010年对黄渤海沿海32个主要渔港、渔村和5家网具生产厂家的实地调查和现场测量资料,结合2009年度三省一市(河北、辽宁、山东、天津)的渔具渔法调查报告以及1983~2009年中国渔业年鉴中记载的捕捞产量分类统计资料,对黄渤海区捕捞结构进行了研究.研究结果表明,黄渤海区主要捕捞品种有30种之多,其中年渔获量超过万吨的有14种鱼类、5种甲壳类、4种头足类、2种海蜇和6种贝类.渔业资源品种的多样性决定了黄渤海区捕捞结构的多样化和多层次的特点,现有9大类19型23式130种网型或作业方式,同时受渔业资源变动的影响,捕捞网具也在变化,不断有旧的渔具消失和新的网具出现.其中,拖网是主要作业方式,贡献率为38.47%~51.79%,且近几年还有上升的趋势;其次为刺网,贡献率从1983年最低时的7.66%逐步上升到目前的30.14%;第三为张网,贡献率从1985年最高时的34.31%下降到目前的10.49%;围网和钓具的贡献率一直较低,均不足4%,其他渔具的贡献率为5.98%~13.30%.目前黄渤海区渔业结构的调整方向应当是减少拖网作业,控制刺网总量,限制张网、陷阱类渔具和地笼,鼓励发展钓渔业     

夏、秋季南黄海小型底栖动物空间分布格局及其环境影响因素

为研究南黄海小型底栖动物的空间分布格局及其环境影响因素,于2020年8月（夏季）和11月（秋季）对南黄海进行了两个航次的野外观测和采样,对小型底栖动物的类群组成、丰度、生物量、垂直分布、群落结构及其与环境因子的关系进行了研究。结果显示,共鉴定出小型底栖动物类群15个,其中自由生活海洋线虫为最优势类群,在两个航次中分别占小型底栖动物总丰度的75.6%和84.6%。其他较重要的类群还包括底栖桡足类、轮虫类和枝角类等。夏季和秋季小型底栖动物的平均丰度分别为（514.9±32.1）ind./(10 cm²) 和（350.8±30.7）ind./(10 cm²),平均生物量（干质量）分别为（651.7±98.0）μg/(10 cm²)和（589.2±37.1）μg/(10 cm²)。小型底栖动物在时空分布上存在差异。在季节分布上,小型底栖动物丰度和类群组成存在极显著差异。结合环境因子分析结果可知,沉积物中值粒径是引起差异的主要环境因子。在空间分布上,夏季小型底栖动物丰度和类群组成在不同水深间存在极显著差异,秋季小型底栖动物丰度和类群组成在不同水深间差异不显著。推测黄海冷水团是影响夏季小型底栖动物空间分布差异的主要因素。本研究中小型底栖动物的数量和类群多样性相较于国内其他对南黄海小型底栖动物的研究较低,其中沉积物叶绿素a含量及有机质含量是引起南黄海小型底栖动物丰度变化的重要因素。海洋线虫与桡足类的丰度比值（N/C比值）评估显示秋季该区域存在有机污染,这一结果与应用大型底栖动物对同一区域进行环境评价的结果不一致,对于应用N/C比值评价环境质量还需要进一步的研究。     

黄海中部太平洋磷虾的大小组成和丰度分布的季节变化及其与环境因子的关系

The seasonal size structure and spatial abundance distributions of Euphausia pacifica populations were investigated in the central part of southern Yellow Sea from August 2009 to May 2010.The abundance and biomass of E.pacifica were higher in spring and summer,and lower in autumn and winter.The mean abundance and biomass(calculated by carbon)were 74.94 ind./m~3 and 8.23 mg/m~3,respectively.Females with total length(TL)ranging between 10 and 19 mm in summer had a substantial contribution to the population biomass,whereas larvae of TL of 3–7 mm in spring were the main contributor to the population abundance.The sex ratio(female:male)showed a female bias in four seasons.Its value peaked in summer,and then decreased in autumn,spring,and winter successively.Cohort analysis revealed that the length-frequency distribution of E.pacifica could be characterized as one group with large animals(mean TL12 mm)accompanied by one or two subgroups of small individuals(mean TL7 mm).Regarding the spatial distribution,juveniles and adults of E.pacifica tend to concentrate in relatively deep water with low temperature(～11℃)and high salinity(32),whereas its larvae showed more abundance in inshore water with rich chlorophyll a,low salinity(32),and warm temperature(11℃),especially in summer and autumn.Associations changed seasonally between stage-specific abundance and environmental factors.     

南黄海春季水温分布特征的分析

利用美国海军的空间分辩率为10′×10′月平均的GDEM三维水温资料,研究了南黄海春季水温的分布特征及其演变过程。分析结果较清晰地显示了春季南黄海的水温分布如何从冬季的垂直均匀型过渡到夏季的层化结构。分析还表明:春季南黄海水温的水平和垂直结构皆比冬季更为复杂,并出现若干个较特殊的水文现象,例如,在34°40′~36°20′N的南黄海西侧出现了“青岛冷水团”,而在35°30′~37°20′N的南黄海东侧,初次发现存在着一个类似性质的冷水团,称其为“仁川外海冷水团”。此外,在冷水团的邻近海域还存在着中层冷水。     

Community structure of picoplankton abundance and biomass in the southern Huanghai Sea during the spring and autumn of 2006

During spring and autumn of 2006,the investigations on abundance,carbon biomass and distribution of picoplankton were carried out in the southern Huanghai Sea(Yellow Sea,sHS) . Three groups of picoplankton-Synechococcus(Syn) ,Picoeukaryotes(PEuk) and heterotrophic bacteria(BAC) were identified,but Prochlorococcus(Pro) was undetected. The average abundance of Syn and PEuk was lower in spring(5.0 and 1.3 × 10 3 cells/cm 3,respectively) than in autumn(92.4 and 2.7 × 10 3 cells/cm 3,respectively) ,but it was opposite for BAC(1.3 and 0.7 × 10 6 cells/cm 3 in spring and autumn,respectively) . And the total carbon biomass of picoplankton was higher in spring(37.23 ± 11.67) mg/m 3 than in autumn(21.29 ± 13.75) mg/m 3 . The ratios of the three cell abundance were 5:1:1 341 and 30:1:124 in spring and autumn,respectively. And the ratios of carbon biomass of them were 5:7:362 and 9:4:4 in spring and autumn,respectively. Seasonal distribution characteristics of Syn,PEuk,BAC were quite different from each other. In spring,Syn abundance decreased in turn in the central waters(where phytoplankton bloom in spring occurred) ,the southern waters and inshore waters of the Shandong Peninsula(where even Syn was undetected) ;the high values of PEuk abundance appeared in the central and southern waters and the inshore of the Shandong Peninsula;the abundance of BAC was nearly three order of magnitude higher than that of photosynthetic picoplankton,and high values appeared in the central waters. In autumn,Syn abundance in central waters was higher than that in surrounding waters,while for PEuk abundance,it decreased in turn in the inshore waters of the Shandong Peninsula,the southern waters and the central waters;BAC presented a complicated blocky type distribution. Sub-surface maximum of each group of picopalnkton appeared in both spring and autumn. Compared with the available literatures concerning the studied area,the range of Syn abundance was larger,and the abundance of BAC was higher. In addition,the conversion factors for calculating picoplanktonic carbon biomass were discussed,with the conversion factors which are different from previous studies in the same surveyed waters. The result of regression analysis showed that there was distinct positive correlation between BAC and photosynthetic picoplankton in spring(r=0.61,P 0.001) ,but no correlation was found in autumn.