The Pleistocene reindeer of the North Sea—initial palaeontological data and archaeological remarks

The interdisciplinary North Sea Project aims at investigating the biotic history of the Pleistocene in the Southern Bight of the North Sea. Humans were part of these biotopes too as Palaeolithic flint artefacts prove. Based on a large fossil record and radiocarbon dating, it becomes clear that reindeer was part of the Late Pleistocene Mammoth-fauna.Upper Palaeolithic hunters often are described as reindeer-hunters. Despite a large fossil reindeer record and collection of Palaeolithic flint artefacts, no evidence was found for co-existence of humans and reindeer or for reindeer-hunting.     

Last Interglacial peopling of Siberia: the Middle Palaeolithic site Ust''-Izhul'', the upper Yenisei area

Multidisciplinary Quaternary investigations in the Minusinsk Basin in the upper Yenisei River region and other southern Siberian continental depressions have produced evidence of prehistoric peopling pre‐dating the last glacial stage (>100 ka BP). Abundant ‘pebble tools’ and bone artefacts exposed from eroded alluvia of the Yenisei River terraces indicate repeated occupation of this territory since the Middle Pleistocene. A new stage of expansion of the early human occupation habitat occurred around the last interglacial (OIS 5e) by a Middle Palaeolithic (Neanderthaloid?) population characterized by a core and flake stone industry and open‐air occupation sites. The key camp/kill‐processing site at Ust‘‐Izhul’, dated to c. 125 ka BP and documenting complex behavioural activities, is so far the most complete in situ pre‐Late Palaeolithic site found in Siberia. This unique record provides new insights into the timing and the palaeoenvironmental conditions of the Pleistocene colonization of north‐central Asia.     

Coastal curios? An analysis of ex situ beach finds for mapping new Palaeolithic sites at Happisburgh,UK

Recent archaeological discoveries from exposures of the Cromer Forest-bed Formation at Happisburgh, UK, have radically changed interpretations of the nature and timing of early hominin occupation of northern latitudes, but this in situ archaeology is only one part of the picture. Surface finds of Pleistocene mammalian remains have been found along this coastline for centuries, with stone tools adding to this record over the past 7 years. The ex situ nature of these finds, however, means they are often seen as limited in the information they can provide. This work contributes to a growing body of research from a range of landscape and environmental contexts that seeks to demonstrate the value and importance of these ex situ assemblages. Here the focus is on Palaeolithic flint artefacts and Pleistocene mammalian remains recovered by a group of local collectors through systematic, GPS-recorded beach collection from 2013 to 2017, and their use in developing a methodology for working with ex situ Palaeolithic finds in coastal locations. The results demonstrate significant patterning that identifies unexplored exposures both onshore and offshore, considerably expanding the known extent of deposits and facilitating new insights into the wider archaeological landscape associated with the earliest occupation of northern Europe.     

A re-evaluation of the timing of the earliest reported human occupation of Britain: the age of the sediments at Happisburgh, eastern England

Lower Palaeolithic artefacts have been reported at Happisburgh, north Norfolk, in sediments that have been assigned to the late Early Pleistocene, in either marine isotope stage (MIS) 25 or 21, using magnetostratigraphy, biostratigraphy and clast lithology. However, the proposal that these sediments were deposited by the ancestral River Thames is inconsistent both with the established late Early Pleistocene palaeogeography of the region and with the dispositions of the contemporaneous Thames terraces. The Happisburgh deposits were evidently emplaced by a local river, which reworked older sediments that from their lithology had been derived largely from the Bytham River rather than the Thames catchment. Nonetheless, the potential significance of this sedimentary succession for early human dispersal and behaviour requires a conservative assessment of its youngest possible age. Although its basal part is clearly Early Pleistocene, there is nothing to preclude an early Middle Pleistocene age for the overlying sediments that have yielded the artefacts and the mammalian biostratigraphic evidence. It is indeed arguable that these sediments date from the cooling transition at the end of MIS 15c, and are thus younger than the artefact-bearing succession at Pakefield. Pending the availability of additional dating evidence, future discussion of the Happisburgh site should be qualified with respect to any claim for an Early Pleistocene age for the human occupation indicated.     

The Homo habitat niche: using the avian fossil record to depict ecological characteristics of Palaeolithic Eurasian hominins

Although hardly applied to human palaeoecology, bird fossils offer a unique opportunity for quantitative studies of the hominin habitat. Here we reconstruct the Homo habitat niche across a large area of the Palaearctic, based on a database of avian fauna for Pleistocene sites. Our results reveal a striking association between Homo and habitat mosaics. A mix of open savannah-type woodland, wetlands and rocky habitats emerges as the predominant combination occupied by Homo across a wide geographical area, from the earliest populations of the Lower Palaeolithic to the latest hunter-gatherer communities of the Upper Palaeolithic. This observation is in keeping with the view that such landscapes have had long standing selective value for hominins.     

Reassessment of the ‘Abbevillien’ in the perspective of new discoveries from the Lower Palaeolithic and Quaternary sites of Abbeville (Somme,northern France)

North-West Europe yields few traces of early human occupation, in particular for the Acheulean. In this context, the Somme Valley in northern France offers a route to Britain during various Pleistocene low sea levels, and has provided numerous evidence of Lower Palaeolithic human occupation through fieldwork initiated during the 19th century. These localities are associated with the original definition in the 1930s by the French prehistorian Abbé Henri Breuil of the ‘Abbevillien’ (Abbevillian facies), based on lithic pieces including crudely made bifaces recovered in particular in some famous key localities of Abbeville, Carpentier, Léon and Moulin Quignon quarries. The history of the term and its definition subsequently gave rise to debates concerning the chronocultural framework of Palaeolithic assemblages among the scientific community of prehistorians over time, from Jacques Boucher de Perthes, Gabriel de Mortillet, Geoffroy d'Ault du Mesnil, Victor Commont, Henri Breuil and François Bordes. New investigations at these three localities, all associated with the High Terrace of the Somme system, pushed back the age of the expansion of the Acheulean both in northern France and in Western Europe to c. 670–650 000 years. They imply that early hominins were able to settle in North-West Europe during both climatic temperate and cold phases. Our work, including new excavations and associated field observations of the three Abbeville localities involved at the onset of the controversy, allows a re-examination of the Abbevillian and contributes to the discussion of the history of Prehistoric Science and the Earliest ‘Acheuleans’ in North-West Europe.     

Palaeolithic and Quaternary research in the Trent Valley (UK): contributions by early collectors

A brief history is presented here of the activities of the three most notable collectors of Lower and Middle Palaeolithic artefacts from Quaternary river terrace deposits in the Trent Valley, derived from archival material that has been largely ignored by previous research. Two, Mr. Fred W.G. Davey and Mr. George F. Turton, were local amateur collectors who did not publish the results of their work themselves and were reliant on collaboration with established archaeologists. The third, Mr. A. Leslie Armstrong, was an archaeologist best known for his work at Creswell Crags and Grimes Graves. Armstrong also made many Palaeolithic discoveries in the Trent Valley but published few details of his own material. Although such details of early Palaeolithic research in the Midlands and North of Britain are predominantly of historical interest only, they nonetheless provide a number of insights into the apparent lack of interest shown in areas north of the ‘Severn-Wash Line’ by collectors of Palaeolithic artefacts. Importantly, the dataset assembled by the Trent Valley Palaeolithic Project (TVPP) and summarized here is the only complete record of the known artefacts and archival material from the Trent, a proportion of which is now unavailable to the research community, having entered unknown private ownership since being studied. The second part of the paper relates this early research to current knowledge of the British Palaeolithic at its most northern fringes and to recent developments in reconstructing the Quaternary evolution of the River Trent.     

Fluvial deposits as an archive of early human activity

River terraces are well established as an important source of Lower and Middle Palaeolithic artefacts in Europe, large collections having been assembled there during the years of manual gravel extraction. Now that many terrace sequences can be reliably dated and correlated with the oceanic record, potentially useful patterns can be recognized in the distribution of artefacts. The earliest appearance of artefacts in terrace staircases, marking the arrival of the first tool-making hominins in the region in question, is the first of several archaeological markers within fluvial sequences. The Lower to Middle Palaeolithic transition, with the appearance of Levallois, is another. Others may be more regional in significance: the occurrences of Clactonian (Mode 1) industry, twisted ovate handaxes and bout coupé handaxes, for example. IGCP Project no. 449 instigated the compilation of fluvial records from all over the ‘old world’. Comparison between British and Central European sequences confirms the established view that there is a demarcation between handaxe making in the west and flake/core industries in the east. Other centres of activity reported here have been in the Middle East (Syria), South Africa and India. Data from such areas will be key in deciphering the story of the earlier ‘out-of-Africa’ migration, that by pre-Homo sapiens people. There is clear evidence for diachroneity between the first appearances of different industries, in keeping with the well-established idea of northward migration.     

Pleistocene Rhine–Thames landscapes: geological background for hominin occupation of the southern North Sea region

This paper links research questions in Quaternary geology with those in Palaeolithic archaeology. A detailed geological reconstruction of The Netherlands' south‐west offshore area provides a stratigraphical context for archaeological and palaeontological finds. Progressive environmental developments have left a strong imprint on the area's Palaeolithic record. We highlight aspects of landscape evolution and related taphonomical changes, visualized in maps for critical periods of the Pleistocene in the wider southern North Sea region. The Middle Pleistocene record is divided into two palaeogeographical stages: the pre‐Anglian/Elsterian stage, during which a wide land bridge existed between England and Belgium even during marine highstands; and the Anglian/Elsterian to Saalian interglacial, with a narrower land bridge, lowered by proglacial erosion but not yet fully eroded. The Late Pleistocene landscape was very different, with the land bridge fully dissected by an axial Rhine–Thames valley, eroded deep enough to fully connect the English Channel and the North Sea during periods of highstand. This tripartite staging implies great differences in (i) possible migration routes of herds of herbivores as well as hominins preying upon them, (ii) the erosion base of axial and tributary rivers causing an increase in the availability of flint raw materials and (iii) conditions for loess accumulation in northern France and Belgium and the resulting preservation of Middle Palaeolithic sites. Copyright © 2011 John Wiley & Sons, Ltd.     

Vegetation context and climatic limits of the Early Pleistocene hominin dispersal in Europe

The vegetation and the climatic context in which the first hominins entered and dispersed in Europe during the Early Pleistocene are reconstructed, using literature review and a new climatic simulation. Both in situ fauna and in situ pollen at the twelve early hominin sites under consideration indicate the occurrence of open landscapes: grasslands or forested steppes. The presence of ancient hominins (Homo of the erectus group) in Europe is only possible at the transition from glacial to interglacial periods, the full glacial being too cold for them and the transition interglacial to glacial too forested. Glacial–interglacial cycles forced by obliquity showed paralleled vegetation successions, which repeated c. 42 times during the course of the Early Pleistocene (2.58–0.78 Ma), providing 42 narrow windows of opportunity for hominins to disperse into Europe.The climatic conditions of this Early Pleistocene vegetation at glacial-interglacial transitions are compared with a climatic simulation for 9 ka ago without ice sheet, as this time period is so far the best analogue available. The climate at the beginning of the present interglacial displayed a stronger seasonality than now. Forest cover would not have been hampered though, clearly indicating that other factors linked to refugial location and soils leave this period relatively free of forests. Similar situations with an offset between climate and vegetation at the beginning of interglacials repeated themselves throughout the Quaternary and benefitted the early hominins when colonising Europe.The duration of this open phase of vegetation at the glacial–interglacial transition was long enough to allow colonisation from the Levant to the Atlantic.The twelve sites fall within rather narrow ranges of summer precipitation and temperature of the coldest month, suggesting the hominins had only a very low tolerance to climate variability.     

The artefacts from the present land surface at the Palaeolithic site of Warren Hill,Suffolk, England

Over 2000 handaxes located in museum collections point to the archaeological importance of Warren Hill, a Lower Palaeolithic site in the path of the pre-Anglian Bytham River in East Anglia that was worked for gravel until about 1950. Yet apart from the initial report by Solomon (1933), detailed archaeological study was not undertaken until 2002. This report describes the surface fieldwalking carried out from 1964 to the time of the 2002 excavations, during which over 400 Palaeolithic artefacts have been recovered. Over 92% of these are flakes, some retouched, thus redressing the bias towards handaxes in the museum collections, and suggesting that flake tools played a major role in Lower Palaeolithic daily life. Also represented are notched pieces, scrapers, cores and a few Neolithic flakes. The surface comprises a lag deposit in which artefacts are concentrated in two main zones, contrasting with the subsurface material which is sparse and well-spread through the lithostratigraphy. The Palaeolithic artefacts are mostly rolled and edge-damaged, and patination is variable, suggesting a complex history. The two types of handaxe noted by Solomon – crude and fine – are confirmed. The preservation of typological integrity amongst surface finds that appear to lie in the path of a major fluvial feature is either a relic of humanly assembled, pre-Anglian bankside clusters eroded into the stream, or it may signal the arrival of post-Anglian humans who discovered a lag deposit rich in artefacts and flint clasts and proceeded to use, resharpen and re-arrange them. The dense clustering noted at Warren Hill is placed in a world context. Amongst the surface gravels an assemblage of flakes together with early 20th century rubbish was discovered; it is interpreted as a collector's dump of unsaleable artefacts.     

San Quirce (Palencia,Spain): new chronologies for the Lower to Middle Palaeolithic transition of south-west Europe

San Quirce is an open-air archaeological site situated on a fluvial terrace in the Duero basin (Palencia, northern Iberia). This paper presents new and consistent chronologies obtained for the sedimentary sequence using post-infrared infrared stimulated luminescence (pIR-IR) dating of K-feldspars and single-grain thermally transferred optically stimulated luminescence (TT-OSL) dating of quartz. The new dating results indicate that the sequence is older than ~200 000 years and place San Quirce Level III within marine isotope stages (MIS) 8 and 7, between 274 ± 13 ka and 238 ± 13 ka. The main lithic assemblage at San Quirce comes from Level III. The predominant tool types found in this level are hammerstones, manuports and flakes, with a small proportion of cores and a significant presence of denticulates. Adaptation to local environmental conditions resulted in distinctive cultural habits, which were embedded in the cultural tradition of hominins occupying the site during the final third of the Middle Pleistocene. San Quirce preserves a simple cultural tradition that was employed by local hominins to engage in a diverse array of activities, and highlights the cultural diversity that appears to have been a characteristic feature of the Lower to Middle Palaeolithic transition 300–200 ka.     

Age of Middle Pleistocene fauna and Lower Palaeolithic industries from Kent's Cavern,Devon

Kent's Cavern has long been known as potentially among the oldest Palaeolithic sites in the country, with the basal Breccia deposit containing a sparse Lower Palaeolithic industry. The sediment consists of a chaotic clayey conglomerate emplaced as a series of debris flows, which entered the cave via blocked entrances at its southwest end. The Breccia contains a fauna dominated by the bear Ursus deningeri, with lion Felis leo and the voles Arvicola cantiana and Microtus oeconomus, establishing a late Cromerian age for the deposit. The artefacts comprise an industry of crudely manufactured handaxes and flakes, and show damage suggesting that they were brought into the cave by the debris flows, and may thus predate the sediment and fauna. We demonstrate an age of >340 ka for the Breccia using two independant dating methods, consistent with existing models of the age of the British Middle Pleistocene sequence.     

The British Pleistocene fluvial archive: East Midlands drainage evolution and human occupation in the context of the British and NW European record

This paper reviews the Pleistocene evolution and human occupation of the River Trent, the major fluvial artery draining Midland Britain, and places it within a modern Quaternary context. In contrast to the sedimentary records of the River Thames and the erstwhile Bytham system, which extend back to the early Pleistocene, present knowledge of the terrace sequence of the Trent, its tributary systems and associated ancestral courses extends back only to the Anglian glaciation (Marine Isotope Stage (MIS) 12), although the regional pre-Anglian drainage configuration is demonstrably complex. The post-Anglian sequence is well developed, with major terrace sand and gravel aggradations associated with each subsequent cold stage. Temperate-climate sediments correlating with MIS 7 and 5e have been recorded, although deposits relating to earlier interglacials during MIS 11 and 9 have yet to be identified. Evidence for human occupation in the form of Lower and Middle Palaeolithic artefacts has been recorded from terrace sediments correlated with MIS 8 and MIS 4, but the majority of this material is heavily rolled and abraded, suggesting significant reworking from older deposits. This review demonstrates that there is a rich palaeo-environmental record from the Trent but the lack of a high-resolution chronostratgraphic framework raises issues about correlation with other systems.     

Early Human Evolution in the Western Palaearctic: Ecological Scenarios

This review presents the themes of a special issue dealing with environmental scenarios of human evolution during the Early Pleistocene (2.6–0.78 Ma; MIS 103-MIS 19) and early Middle Pleistocene (0.78–0.47 Ma; MIS 19-base of MIS 12) within the western Palaearctic. This period is one of dramatic changes in the climates and the distribution of Palaearctic biota. These changes have played their role in generating adaptive and phyletic patterns within the human ancestry, involving several species such as Homo habilis, “Homo georgicus”, Homo erectus, Homo antecessor and Homo heidelbergensis. In the archaeological record, these species include the Oldowan (Mode 1) and Acheulian (Mode 2) lithic technologies. Taphonomic considerations of palaeoecological research in hominin-bearing sites are provided and evaluated. Syntheses are provided for north Africa, western Asia, the Mediterranean Basin, Britain, and continental Europe. Palaeoenvironmental reconstructions based on multidisciplinary data are given for Ain Boucherit, Ain Hanech and El-Kherba in Algeria, Dmanisi in Georgia, Atapuerca, Cueva Negra, and the Orce Basin in Spain, Monte Poggiolo and Pirro Nord in Italy, Pont-de-Lavaud in France, and Mauer in Germany. The state of the art with the Out of Africa 1 dispersal model is reviewed. A source-sink dynamics model for Palaeolithic Europe is described to explain the morphological disparity of H. heidelbergensis (we will sometimes use the informal name “Heidelbergs”) and early Neanderthals. Other aspects debated here are the selective value of habitat mosaics including reconstructions based on mammal and avian databases, and the role of geological instability combined with topographic complexity. This review is completed by addressing the question of whether the appearance of evolutionary trends within hominins is concentrated in regions of highest worldwide biological diversity (biodiversity hotspots). It is concluded that the keys for the activation of evolutionary change in hominins may have been geological instabilities, and a shifting physiographical heterogeneity combined with high biodiversity and ecological interaction.     

Palaeoenvironments of early hominins in temperate and Mediterranean Eurasia: new palaeobotanical data from Palaeolithic key-sites and synchronous natural sequences

The present study focuses on new palaeobotanical data (pollen, phytoliths and fruits) from three of the oldest Early Palaeolithic sites in Eurasia (Dmanisi in Georgia, Ca’ Belvedere di Monte Poggiolo in Italy and Pont-de-Lavaud in France). The main aim is to examine the ecological factors associated with the first human dispersals out of Africa into Eurasia. The palaeoecological data are discussed with regards to chronology and geographical location of the settlements. The vegetation pattern of each site consists of temperate Eurasian and Mediterranean taxa, subtropical trees being more or less recorded depending on climatic and geographic features. The new palaeobotanical data show that different vegetation structures occurred between the first human dispersal in Caucasus and the later dispersals in western Eurasia. However, in all cases hominins seemed to be adapted to temperate ecosystems. In Caucasus, human occupation took place at 1.7 Ma in a forest-steppe environment, while at ca 1 Ma human populations occupied various settings such as open landscapes or dense forests. These data suggest that during the Early Pleistocene, human populations evolved and dispersed in western Eurasia, gradually increasing their degree of adaptation to diversified environments.     

Hominin variability,climatic instability and population demography in Middle Pleistocene Europe

We propose a population model for Middle Pleistocene Europe that is based on demographic “sources” and “sinks”. The former were a small number of “core” or populations in glacial refugia in southern Europe from which hominins expanded northwards in interstadial and interglacial periods; occupation outside glacial refugia would have been restricted to warm or temperate periods, and populations at the northern limit of the Middle Pleistocene range would have been “sink” populations in that they depended upon recruitment from source populations further south. Southwest Asia would also have been a likely source of immigrant, source populations. We argue as an alternative to an “ebb and flow” model in which groups retreated to refugia when conditions worsened that local extinction outside refugia would have been frequent. In extreme situations, Europe may have been a population “sink” (i.e. unpopulated) that was replenished from source populations in Southwest Asia. We suggest that this pattern of repeated colonisation and extinction may help explain the morphological variability of European Middle Pleistocene hominins, particularly Homo heidelbergensis and its apparent non-lineal evolution towards Homo neanderthalensis.     

A demographic model for Palaeolithic technological evolution: The case of East Asia and the Movius Line

The Pleistocene record of East Asia continues to pose controversial questions for palaeoanthropology, especially with regard to Palaeolithic technological patterns. In recent years, an increased understanding of the effect of demography on cultural transmission has improved our understanding of the incidence, proliferation, and elaboration of technological traditions. Here, we present a generalised null model of Lower–Middle Palaeolithic technological evolution, which expressly links cultural transmission theory and demographic factors (i.e. population size, density, and social interconnectedness). Consistent with our model, Africa exhibits evidence of major technological innovations during the Early to Middle Pleistocene, due to a constant source of population and growth due to accumulation through time. In comparison, Pleistocene East Asian assemblages are dominated by Mode 1-type technologies, and only a few localized occurrences of bifacial technology are currently known. We detail evidence suggesting that during much of the Pleistocene a combination of biogeographical, topographical, and dispersal factors are likely to have resulted in relatively lower effective population sizes in East Asian hominins compared with western portions of the Old World, particularly Africa. Thus, the Movius Line – as is the case with its namesake ‘Wallace's line’ – must be examined in terms of its biogeographical context, if the divergent evolutionary trajectories of entities either side of it are to be understood. Most parsimoniously, the Movius Line sensu lato is thus a ‘line’ which represents the crossing of a demographic threshold. Under the parameters of our (testable) null model, geographically and temporally sporadic occurrences of bifacial technology in East Asia are the product of short-lived instances of technological convergence. As a consequence, the in situ evolution of Levallois (Mode 3) was inhibited in East Asia due to the constraints of relatively smaller effective population sizes.     

The last 750 ka in loess–palaeosol sequences from northern France: environmental background and dating of the western European Palaeolithic

This study presents an overview of Middle Pleistocene loess–palaeosol sequences (LPS) in northern France and discusses the palaeoclimatic significance of the pedosedimentary record in the context of western European LPS and of global climatic cycles for the last 750 ka. In this area, the oldest loess deposits (early Middle Pleistocene) are preserved in sedimentary traps (leeward scarps of fluvial terraces and dissolution sinkholes). They result from local deflation processes reworking Pleistocene sandy fluvial deposits or relicts of Tertiary sands. A large extension of typical calcareous loess over the landscape, the Loess Revolution, is then observed during MIS 6, with heavy mineral assemblages testifying to long-distance transport from the polar desert area of the dried eastern Channel. A correlation scheme is proposed between the global records of northern France in continental environments and both global palaeoclimatic records and other main western European LPS. After 30 years of research, northern France LPS stand as a fundamental archive of the impact of interglacial–glacial climatic cycles as well as millennial events. Finally, these works provide a robust chronoclimatic framework for the study of the western European Late Acheulean and Middle Palaeolithic and for the relative dating of the various fluvial terraces that they fossilise.     

Last glacial ecology and geoarchaeology of the Central Trans‐Ural area: the Sosva River Upper Palaeolithic Complex,western Siberia

Chlachula, J. & Serikov, Y. B. 2010: Last glacial ecology and geoarchaeology of the Central Trans‐Ural area: the Sosva River Upper Palaeolithic Complex, western Siberia. Boreas, 10.1111/j.1502‐3885.2010.00166.x. ISSN 0300‐9483. Quaternary and geoarchaeology studies from the eastern limits of the Ural Mountains provide multiple lines of evidence of the Palaeolithic peopling of this geographically marginal and still poorly explored territory of western Siberia following the mid‐last glacial (MIS 3) warming. A complex of investigated open‐air localities in the Sosva River basin (the north‐central Trans‐Ural area) at the periphery of the western Siberian Plain, distinguished by very high concentrations of Pleistocene megafaunal remains previously regarded as ‘mammoth cemeteries’, indicate, in conjunction with the associated diagnostic ivory/bone and stone industry, open occupation sites during the Last Glacial (MIS 2). Fossil faunal remains, dominated by mammoth (98%) together with bird and fish species, indicate various methods of exploitation of the Late Pleistocene natural resources and successful behavioural adaptation to the last glacial sub‐polar tundra‐steppe environment. The taphonomy and composition of the well‐preserved skeletal remains from the main occupation sites suggest both active hunting and anthropogenic ‘scavenging’ practices. The contextual geology and the cultural and biotic multi‐proxy records from the Trans‐Ural Upper Palaeolithic Complex provide new insights into the timing and palaeoecological conditions of the Pleistocene human occupation of north‐central Asia.