The influence of wind forcing on across-shelf transport in the Florida Keys

Acoustic Doppler current profiles and current meter data are combined with wind observations to describe the transport of water leaving Florida Bay and moving onto the inner shelf on the Atlantic side of the Florida Keys. A 275-day study in the Long Key Channel reveals strong tidal exchanges, but the average ebb tide volume leaving Florida Bay is 19% greater than the average flood tide volume entering the bay. The long-term net outflow averages 472 m³ s⁻¹. Two studies in shelf waters describe the response to wind forcing during spring and summer months in 2004 and during fall and winter months in 2004–2005. During the spring–summer study, southeasterly winds have a distinct shoreward component, and a two-layer pattern appears. Surface layers move shoreward while near-bottom layers move seaward. During the winter study, the resultant wind direction is parallel to the Keys and to the local isobaths. The entire water column moves in a nearly downwind direction, and across-shelf transport is relatively small. During the summer wet season, Florida Bay water should be warmer, fresher, and thus less dense than Atlantic shelf waters. Ebbing bay water should move onto the shelf as a buoyant plume and be held close to the Keys by southeasterly winds. During the winter dry season, colder and saltier Florida Bay water should leave the tidal channels with relatively high density and be concentrated in the near-bottom layers. But little across-shelf flow occurs with northeasterly winds. The study suggests that seasonally changing wind forcing and hydrographic conditions serve to insulate the reef tract from the impact of low-quality bay water.     

Eurasian Arctic shelf hydrography: Exchange and residence time of southern Laptev Sea waters

The hydrography of the Laptev Sea is significantly influenced by river water and sea-ice processes, which are highly variable over the annual cycle. Despite of an estuarine structure the inner and outer shelf regions are decoupled at times as documented by the stability of a warm intermediate layer formed during summer below the Lena River plume. We demonstrate that a remnant of this warm layer is preserved below the fast ice until the end of winter, while only slightly farther to the north, offshore of the landfast ice in the polynya region, the pycnocline is eroded and no signature of this layer is found. The warm intermediate layer (WIL) formed during summer can be used as tracer for Laptev Sea shelf waters throughout the winter. Thereby, residence times of southern Laptev Sea waters can be estimated to be at least from summer to the end of winter/spring of the following year.     

Patterns of sea surface temperature variability on seasonal to sub-annual scales at and offshore the Río de la Plata estuary

SST variability on seasonal to sub-annual scales in the coastal region of South America between 30° and 39°S, largely influenced by the Rio de la Plata estuary’s plume, and its relation to wind variability are explored. Data are six years of daily ensembles of gridded satellite SST and sea surface winds with spatial resolutions of about 11 and 25 km, respectively. Observations from oceanographic cruises are used to validate the results. It is found that the seasonal cycle can be explained in terms of two modes. The first one, characterizing fall-early winter/spring-early summer, is related to the radiative cycle. The second one, corresponding to late summer and winter, displays warm/cold anomalies along the Uruguayan coast forced by the prevailing winds during those seasons. In the upper estuary and the northern part of the area of influence of the freshwater plume, variability in sub-annual scales is significant. A large portion of this variance is related to zonal wind anomalies that force warm/cold SSTs along that coast. Cold anomalies of up to −5 °C occur under anomalously intense easterly winds, indicating upwelling. These events are very frequent and show large persistence, occurring up to one and a half months. They also display a marked seasonal cycle – being more frequent in late spring and summer – large inter-annual variability and seem to be modulated by the continental runoff. When discharge is low, the freshwater plume retracts to the west, reducing the inner-shelf stratification and increasing the likelihood of a full upwelling to the surface. In winter, short time-scale SST variability is mostly due to variability in the atmospheric cold fronts crossing the region. Weaker or less frequent (stronger or more frequent) fronts produce a generalized warming (cooling) over the region. As the estuary heats (colds) faster than the shelf, a warm (cold) anomaly develops in the upper Río de la Plata. On inter-annual time scales, probably because ENSO activity was weak during the studied period, SST variability was not important.     

Wave climate simulation for southern region of the South China Sea

This study investigates long-term variability and wave characteristic trends in the southern region of the South China Sea (SCS). We implemented the state-of-the art WAVEWATCH III spectral wave model to simulate a 31-year wave hindcast. The simulation results were used to assess the inter-annual variability and long-term changes in the SCS wave climate for the period 1979 to 2009. The model was forced with Climate Forecast System Reanalysis winds and validated against altimeter data and limited available measurements from an Acoustic Wave and Current recorder located offshore of Terengganu, Malaysia. The mean annual significant wave height and peak wave period indicate the occurrence of higher wave heights and wave periods in the central SCS and lower in the Sunda shelf region. Consistent with wind patterns, the wave direction also shows southeasterly (northwesterly) waves during the summer (winter) monsoon. This detailed hindcast demonstrates strong inter-annual variability of wave heights, especially during the winter months in the SCS. Significant wave height correlated negatively with Niño3.4 index during winter, spring and autumn seasons but became positive in the summer monsoon. Such correlations correspond well with surface wind anomalies over the SCS during El Nino events. During El Niño Modoki, the summer time positive correlation extends northeastwards to cover the entire domain. Although significant positive trends were found at 95 % confidence levels during May, July and September, there is significant negative trend in December covering the Sunda shelf region. However, the trend appears to be largely influenced by large El Niño signals.     

Characteristics of seasonal and spatial variations of primary production over the southeastern Bering Sea shelf

In situ primary production data collected during 1978–1981 period and 1997–2000 period were combined to improve understanding of seasonal and spatial distribution of primary production in the southeastern Bering Sea. Mean daily primary production rates showed an apparent seasonal cycle with high rates in May and low rates in summer over the entire shelf of the southeastern Bering Sea except for oceanic region due to lack of data. There was also an increasing trend of primary production rates in the fall over the inner shelf and the middle shelf. There was a decreasing trend of primary production rates between late April and mid-May over the inner shelf while there was an abrupt increase between late April and mid-May over the middle shelf and the outer shelf. In the shelf break region, there was an increasing pattern in late May. These suggest that there was a gradual progression of the development of the spring phytoplankton bloom from the inner shelf toward the shelf break region. There was also a latitudinal variability of primary production rate over the middle shelf, probably due to either spatial variations of the seasonal advance and retreat of sea ice or horizontal advection of saline water in the bottom layer. Annual rates of primary production across the southeastern Bering Sea shelf were 121, 150, 145, 110, and 84 g C m⁻² yr⁻¹ in the inner shelf, the middle shelf, the outer shelf, the shelf break, and oceanic region, respectively. High annual rates of primary production over the inner shelf can be attributed to continuous summer production based on regenerated nitrogen and/or a continuous supply of nitrogen at the inner front region, and to fall production. There were some possibilities of underestimation of annual primary production over the entire shelf due to lack of measurement in early spring and fall, which may be more apparent over the shelf break and oceanic region than the inner shelf, the middle, and the outer shelf. This study suggests that the response of primary production by climate change in the southeastern Bering Sea shelf can be misunderstood without proper temporal and seasonal measurement.     

Modeling the Mozhaisk Reservoir Genetic Structure

A box model of water mass composition is used to determine the genetic structure of the main water mass (MWM) of the Mozhaisk Reservoir in the high-water 1986–1987 and for four constructed scenarios. The greatest difference between the genetic compositions of the spring, summer, and autumn modifications of MWM manifested itself in that the winter reservoir water of the previous water management year predominates in a low-water year. Elevated proportions of snowmelt flood water were recorded in the spring and summer of a medium-water year (with high snowmelt flood and low rain floods) and in the winter of a low-water year. Possible variations (up to 27%) in the salinity and color index of MWM in years with different genetic composition are demonstrated and shown to be attributable to different water abundance in these years. The genetic structure of the reservoir water is supposed to affect possible variations in the phosphorus content of the reservoir MWM.     

Temporal changes in suspended sediment transport in a gullied loess basin: the lower Chabagou Creek on the Loess Plateau in China

Suspended sediment dynamics are still imperfectly understood, especially in the loess hilly region on the Loess Plateau, with strong temporal variability, where few studies heretofore have been conducted. Using a dataset up to eight years long in the Lower Chabagou Creek, the variability in suspended sediment load at different temporal scales (within‐flood variability, monthly–seasonal and annual) is analyzed in this paper. The results show that, on the within‐flood scale, most of the sediment peaks lag behind peak discharges, implying that slope zones are the main sediment source area; independent of the occurring sequences of the peaks of sediment and discharge, all the events could present an anti‐clockwise hysteresis loop resulting from the abundant material and the influence of hyperconcentrated flows on suspended sediment concentration. At monthly and seasonal scales, there is a ‘store–release’ process, i.e. sediment is prepared in winter, spring and late autumn, and exported in summer and early autumn. At the annual scale, the high variability in concentration and sediment yield are highly correlated with water yield, resulting from the number and magnitude of floods recorded yearly, and almost all the suspended load is transported during these events. Copyright © 2008 John Wiley & Sons, Ltd.     

松潘台拱背氡值资料变化分析

松潘拱背水氡值自1977年11月投入观测以来,其年变化规律呈现的是冬低夏高的形态。通过分析发现,汶川8．0级地震以前,水氡观测值既存在1～3年的中长期趋势变化又表现出3个月左右的短期变化,但临震突跳变化并不突出;而气氡观测值在地震孕育、发生和震后存在以短临突跳的方式相对应,变化幅度在50％以上。但由于观测时间短,这个认识有待进一步探讨研究。2007年底该台增上了数字化气氡观测仪器。通过总结反思,发现松潘台拱背观测水点氡观测值的变化与龙门山断裂带破坏性地震的孕育、发生有一定的对应关系,这些变化为今后研究地下水氡含量变化与附近地震的发生提供了一个很好的对应例子。松潘地震台的水氡和气氡观测值变化能较好地反映龙门山构造带应力积累和地震孕育信息。     

The effects of river discharge and seasonal winds on the shelf off southeastern South America

The Río de la Plata waters form a low salinity tongue that affects the circulation, stratification and the distributions of nutrients and biological species over a wide extent of the adjacent continental shelf. The plume of coastal waters presents a seasonal meridional displacement reaching lower latitudes (28°S) during austral winter and 32°S during summer. Historical data suggests that the wind causes the alongshore shift, with southwesterly (SW) winds forcing the plume to lower latitudes in winter while summer dominant northeasterly (NE) winds force its southward retreat. To establish the connection between wind and outflow variations on the distribution of the coastal waters, we conducted two quasi-synoptic surveys in the region of Plata influence on the continental shelf and slope of southeastern South America, between Mar del Plata, Argentina and the northern coast of Santa Catarina, Brazil. We observed that: (A) SW winds dominating in winter force the northward spreading of the plume to low latitudes even during low river discharge periods; (B) NE winds displace the plume southward and spread the low salinity waters offshore over the entire width of the continental shelf east of the Plata estuary. The southward retreat of the plume in summer leads to a volume decrease of low salinity waters over the shelf. This volume is compensated by an increase of Tropical waters, which dominate the northern shelf. The subsurface transition between Subantarctic and Subtropical Shelf Waters, the Subtropical Shelf Front, and the subsurface water mass distribution, however, present minor seasonal variations. Along shore winds also influence the dynamics and water mass variations along the continental shelf area. In areas under the influence of river discharge, Subtropical Shelf Waters are kept away from the coastal region. When low salinity waters retreat southward, NE winds induce a coastal upwelling system near Santa Marta Cape. In summer, solar radiation promotes the establishment of a strong thermocline that increases buoyancy and further enhances the offshore displacement of low salinity waters under the action of NE winds.     

CHANGES IN GLACIER AND ALPINE RUNOFF IN THE NORTH CASCADE RANGE,WASHINGTON, USA 1985–1993

From 1985 to 1993, the mean summer temperature was 1.1°C above the long-term mean and the mean winter precipitation was 11% below the long-term mean at the eight Washington State Cascade Mountain weather stations. The effect of this climate fluctuation on glacier and alpine runoff has been examined in five North Cascade basins. From 1985 to 1993 the two basins with less than 1% glacier-covered area experienced mean 1 July to 30 September (late summer) runoff 36% below the long-term mean. The three moderately glaciated basins (3, 6 and 14% glaciated, respectively) experienced a 13% decline in late summer runoff for the same period. A significant change in late summer runoff has occurred in the North Cascades and this change is less pronounced in glacier basins. The cause of the change is decreased winter precipitation and earlier onset of spring melting of the alpine snowpack, followed by above average summer temperatures and an earlier summer melt of alpine snowpack. The smaller decrease in runoff in glacial basins is due to increased ablation and consequent glacier runoff due to high summer temperatures. However, glacier retreat is also reducing glacier runoff.     

Thermal and haline variability over the central Bering Sea shelf: Seasonal and interannual perspectives

We examine multi-year conductivity-temperature-depth (CTD) data to better understand temperature and salinity variability over the central Bering Sea shelf. Particular consideration is given to observations made annually from 2002 to 2007 between August and October, although other seasons and years are also considered. Vertical and horizontal correlation maps show that near-surface and near-bottom salinity anomalies tend to fluctuate in phase across the central shelf, but that temperature anomalies are vertically coherent only in the weakly or unstratified inner-shelf waters. We formulate heat content (HC) and freshwater content (FWC) budgets based on the CTD observations, direct estimates of external fluxes (surface heat fluxes, ice melt, precipitation (P), evaporation (E) and river discharge), and indirect estimates of advective contributions. Ice melt, P−E, river discharge, and along-isobath advection are sufficient to account for the mean spring-to-fall increase in FWC, while summer surface heat fluxes are primarily responsible for the mean seasonal increase in HC, although interannual variability in the HC at the end of summer appears related to variability in the along-isobath advection during the summer months. On the other hand, FWC anomalies at the end of summer are significantly correlated with the mean wind direction and cross-isobath Ekman transport averaged over the previous winter. Consistent with the latter finding, salinities exhibit a weak but significant inverse correlation between the coastal and mid-shelf waters. The cross-shelf transport likely has significant effect on nutrient fluxes and other processes important to the functioning of the shelf ecosystem. Both the summer and winter advection fields appear to result from the seasonal mean position and strength of the Aleutian Low. We find that interannual thermal and haline variability over the central Bering Sea shelf are largely uncoupled.     

Late Quaternary clay minerals off Middle Vietnam in the western South China Sea: Implications for source analysis and East Asian monsoon evolution

High-resolution clay mineral records combined with oxygen isotopic stratigraphy over the past 450 ka during late Quaternary from Core MD05-2901 off Middle Vietnam in the western South China Sea are re-ported to reconstruct a history of East Asian monsoon evolution. Variations in Illite, chlorite, and kaolinite contents indicate a strong glacial-interglacial cyclicity, while changes in smectite content present a higher frequency cyclicity. The provenance analysis indicates a mixture of individual clay minerals from various sources surrounding the South China Sea. Smectite derived mainly from the Sunda shelf and its major source area of the Indonesian islands. Illite and chlorite originated mainly from the Mekong and Red rivers. Kaolinite was provided mainly by the Pearl River. Spectral analysis of the kaolin-ite/(illite chlorite) ratio displays a strong eccentricity period of 100 ka, implying the ice sheet-forced win-ter monsoon evolution; whereas higher frequency changes in the smectite content show an ice sheet-forced obliquity period of 41 ka, and precession periods of 23 and 19 ka and a semi-precession period of 13 ka as well, implying the tropical-forced summer monsoon evolution. The winter monsoon evolution is generally in coherence with the glacial-interglacial cyclicity, with intensified winter monsoon winds during glacials and weakened winter monsoon winds during interglacials; whereas the summer monsoon evolution provides an almost linear response to the summer insolation of low latitude in the Northern Hemisphere, with strengthened summer monsoon during higher insolation and weakened summer monsoon during lower insolation. The result suggests that the high-latitude ice sheet and low-latitude tropical factor could drive the late Quaternary evolution of East Asian winter and summer monsoons, respectively, implying their diplex and self-contained forcing mechanism.     

Numerical study of seasonal circulation and variability over the inner shelf of the northern South China Sea

This study examines seasonal circulation, hydrography, and associated spatial variability over the inner shelf of the northern South China Sea (NSCS) using a nested-grid coastal ocean circulation model. The model external forcing consists of tides, atmospheric forcing, and open boundary conditions based on the global ocean circulation and hydrography reanalysis produced by the Hybrid Coordinate Ocean model. Five numerical experiments are conducted with different combinations of external forcing functions to examine main physical processes affecting the seasonal circulation in the study region. Model results demonstrate that the monthly mean circulation in the study region features the Guangdong Coastal Current (GCC) over coastal waters and the South China Sea Warm Current (SCSWC) in the offshore deep waters. The GCC produced by the model flows nearly southwestward in winter months and northwestward in summer months, which agrees with previous studies. The SCSWC flows roughly northeastward and is well defined in summer months. In winter months, by comparison, the SCSWC is superseded by the southwestward strong wind-driven currents. Analysis of model results in five different experiments demonstrates that the monthly mean circulation over coastal and inner shelf waters of the NSCS can be approximated by barotropic currents forced by the southwestward monsoon winds in winter months. In summer months, by comparison, the monthly mean circulation in the study region is affected significantly by baroclinic dynamics associated with freshwater runoff from the Pearl River and advection of warm and saline waters carried by the SCSWC over the NSCS.     

Spatial and temporal variability in nutrient concentrations in Liverpool Bay,a temperate latitude region of freshwater influence

This paper presents data for the temporal and spatial distribution of nutrients in Liverpool Bay between 2003 and 2009 and an analysis of inputs of nutrients from the major rivers. The spatial distribution of winter nutrient concentrations are controlled by the region of freshwater influence (ROFI) in Liverpool Bay through the mixing of riverine freshwater and Irish Sea water, with strong linear relationships between nutrient concentration and salinity between December and February. The location of highest spring and summer phytoplankton biomass reflects the nutrient distributions as controlled by the ROFI. Analysis of 7 years of data showed that the seasonal cycle of winter maximum nutrient concentrations in February and drawdown in April/May is a recurrent feature of this location, with the timing of the drawdown varying by several weeks between years. A comparison of observed nutrient concentrations in Liverpool Bay with those predicted from inputs from rivers has been presented. Nutrient concentrations in the rivers flowing into Liverpool Bay were highly variable and there was reasonable agreement between predicted freshwater nutrient concentrations using data from this study and riverine nutrient concentrations weighted on the basis of river flow, although the exact nature of mixing between the rivers could not be determined. Predicted Irish Sea nutrient concentrations in the winter were lower than those reported for the input waters of the North Atlantic, supporting findings from previous work that nitrogen is lost through denitrification in the Irish Sea.     

Transport over a narrow shelf: Exuma Cays, Bahamas

Current meter and temperature data were collected over a 402-day period from an outer shelf and a tidal channel study site in the Exuma Cays, Bahamas. The shelf width is less than 2 km, and floods and ebbs through a nearby tidal channel extend across the entire shelf and reduce coherence of wind forcing and along-shelf flow. The data are used in perturbation analyses to investigate the across-shelf turbulent transport of heat and momentum over seasonal time scales. Data show a net landward transport of both heat and momentum over the course of the study, but the perturbation products contain distinct seasonal cycles. In fall and winter months, across-shelf heat and momentum fluxes are landward, while during spring and summer months fluxes are seaward. Comparison of shelf-water temperature with the temperature of bank water leaving on the ebb suggests that seasonal cycles of across-shelf heat and momentum in shelf waters are influenced by the seasonal export of relatively warm and cool water from Great Bahama Bank.Responsible Editor: Iris Grabemann     

A modeling study of the physical processes affecting the development of seasonal hypoxia over the inner Louisiana-Texas shelf: Circulation and stratification

The physical processes affecting the development of seasonal hypoxia over the Louisiana-Texas shelf were examined using a high-resolution, three-dimensional, unstructured-grid, Finite Volume Coastal Ocean Model (FVCOM). The model was forced with the observed freshwater fluxes from the Mississippi and Atchafalaya Rivers, surface winds, heat fluxes, tides and offshore conditions. The simulations were carried out over a six-month period, from April to September 2002, and the model performance was evaluated against several independent series of observations that included tidal gauge data, Acoustic Doppler Current Profiler (ADCP) data, shipboard measurements of temperature and salinity, vertical salinity and sigma-t profiles, and satellite imagery. The model accurately described the offshore circulation mode generated over the Louisiana-Texas shelf by the westerly winds during summer months, as well as the prevalent westward flow along the coast caused by the easterly winds during the rest of the study period. The seasonal cycle of stratification also was well represented by the model. During 2002, the stratification was initiated in early spring and subsequently enhanced by the intensity and phasing of riverine freshwater discharges. Strong stratification persisted throughout the summer and was finally broken down in September by tropical storms. The model simulations also revealed a quasi-permanent anticyclonic gyre in the Louisiana Bight region formed by the rotational transformation of the Mississippi River plume, whose existence during 2002 was supported by the satellite imagery and ADCP current measurements. Model simulations support the conclusion that local wind forcing and buoyancy flux resulting from riverine freshwater discharges were the dominant mechanisms affecting the circulation and stratification over the inner Louisiana-Texas shelf.     

A 200-year precipitation index for the central English Lake District / Un indice de précipitation de 200 ans pour la Région des Lacs en Angleterre

Abstract

Abstract The geographical context and hydroclimatology of the English Lake District means that the region is an important monitor of changes to nationally significant environmental assets. Using monthly rainfall series for sites in and around the central Lake District, a continuous ~200-year precipitation index was constructed for a representative station close to Grasmere. The bridged series shows a significant decline in summer rainfall since the 1960s, offset by increases in winter and spring that are strongly linked to North Atlantic forcing. Over longer time periods, the index exhibits several notable dry (1850s, 1880s, 1890s, 1930s, 1970s) and wet (1820s, 1870s, 1920s, 1940s, 1990s) decades. These patterns are strongly reflected by reservoir inflow series and by indicators of the biological status of the region’s freshwater lakes. It is argued that long-term climate indices will become increasingly important as managers seek to evaluate recent and project environmental changes within the context of long-term natural variability.     

EFFECTS OF CLIMATE CHANGE ON INLAND WATERS OF THE PACIFIC COASTAL MOUNTAINS AND WESTERN GREAT BASIN OF NORTH AMERICA

The region designated as the Pacific Coastal Mountains and Western Great Basin extends from southern Alaska (64°N) to southern California (34°N) and ranges in altitude from sea level to 6200 m. Orographic effects combine with moisture-laden frontal systems originating in the Pacific Ocean to produce areas of very high precipitation on western slopes and dry basins of internal drainage on eastern flanks of the mountains. In the southern half of the region most of the runoff occurs during winter or spring, while in the northern part most occurs in summer, especially in glaciated basins. Analyses of long-term climatic and hydrological records, combined with palaeoclimatic reconstructions and simulations of future climates, are used as the basis for likely scenarios of climatic variations. The predicted hydrological response in northern California to a climate with doubled CO₂ and higher temperatures is a decrease in the amount of precipitation falling as snow, and substantially increased runoff during winter and less in late spring and summer. One consequence of the predicted earlier runoff is higher salinity in summer and autumn in San Francisco Bay. In saline lakes, the incidence of meromixis and the associated reduction in nutrient supply and algal abundance is expected to vary significantly as runoff fluctuates. In subalpine lakes, global warming will probably will lead to increased productivity. Lacustrine productivity can also be altered by changes in wind regimes, drought-enhanced forest fires and maximal or minimal snowpacks associated with atmospheric anomalies such as El Niño–Southern Oscillation (ENSO) events. Reduced stream temperature from increased contributions of glacial meltwater and decreased channel stability from changed runoff patterns and altered sediment loads has the potential to reduce the diversity of zoobenthic communities in predominately glacier-fed rivers. Climatic warming is likely to result in reduced growth and survival of sockeye salmon in freshwater, which would, in turn, increase marine mortality. Further research activities should include expanded studies at high elevations and of glacier mass balances and glacial runoff, applications of remote sensing to monitor changes, further refinement of regional climatic models to improve forecasts of future conditions and continued analyses of long-term physical, chemical and biological data to help understand responses to future climates. © 1997 John Wiley & Sons, Ltd.     

The reproductive phenology of Isoetes lacustris L.: Results of field studies in Scandinavian lakes

The phenologic change of the leaf rosette structure of Isoetes lacustris L. was studied in 26 lakes of temperate, boreal, or subarctic Scandinavia between 59° and 70° n.l. The investigations were carried out during six defined seasons: late winter, spring, early summer, late summer, autumn, and early winter. From 640 plants, gained with the aid of SCUBA, six leaf types were distinguished: immature megasporophylls, mature megasporophylls, immature microsporophylls, mature microsporophylls, and sporophylls that had released their spores and leaves with undeveloped sporangia. Mean numbers per rosette of each leaf type were established in each study lake and study season, resulting in a common pattern embracing all lakes studied.

Megasporophylls are developed throughout the year, whenever the water temperature is about 10 °C. Their share was always more than 30%, excepting winter. Microsporophylls are produced preferentially in spring/early summer when the days are longest; they amount to more than 50% of the rosette leaves during this period, but only to some 10% in the remaining seasons. The spores mature and are released between late summer and early winter. It is concluded that not all spores mature in the year of their birth, and those that do not mature are released in the early summer of the following year, as well as the old empty leaves become detached.     

Remote sensing and modelling of bio-physical distribution patterns at Porcupine and Rockall Bank,Northeast Atlantic

Monthly composites of multi-year sea surface temperature (SST) and chlorophyll-a (Chl-a) have been used in combination with ocean model simulations to study bio-physical distribution patterns at Porcupine and Rockall Bank, two large submarine banks in the Northeast Atlantic in close proximity to the European shelf edge. Seven years (January 1998–December 2004) of remotely sensed data have been collated to create monthly climatological fields and to analyse principal spatio-temporal characteristics. At both banks, a region of cooler SST is found over the summit region compared to warmer waters of the surrounding ocean, less apparent in summer when capped by the seasonal thermocline. Enhanced Chl-a levels are found over both banks with a lifetime partly exceeding the bloom period. At Rockall Bank, both SST and Chl-a signals are more pronounced and persistent showing a 30% increase in annual Chl-a levels over the summit area with an even higher ratio in spring and autumn. A combination of physical processes appears to promote the enhanced productivity over both banks through the generation of a quasi-steady dome of cold, dense water during winter convection and upwelling events. This cold dome is associated with the presence of a retentive circulation based on Taylor cap dynamics and tidal rectification processes. The larger and more persistent enhancement of Chl-a levels over Rockall Bank would appear due to its isolated nature as well as its size. In contrast, Porcupine Bank is partly attached to the Irish shelf edge and exposed to the poleward flowing shelf edge current which may strip passive particles from the central bank region. Satellite derived Chl-a spring/summer distributions over the banks have been used to initialise model simulations of passive tracer dispersion. Timescales for the observed lifetime of the remotely sensed Chl-a patches are consistent with model derived retention timescales and simple scaling for the dispersion of passive biological material over the banks. Surface particle residence times over Rockall Bank are estimated to exceed Porcupine Bank values by a factor of two. Finally, the tidal contribution to individual particle motion is found to be large in some Rockall Bank areas, but less important at Porcupine Bank.