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1.
Using an interdisciplinary three-dimensional physical and biogeochemical model developed for the Black Sea, the long-term evolution of marine dynamics and ecosystem is investigated. The hydrophysical fields were calculated from a model of Black Sea circulation with assimilation of hydrographic survey and satellite measurement data from 1971 to 2001. The circulation model reproduces well processes of various scales in both space and time (particularly the seasonal course and interannual variability of main hydrophysical fields). The resulting flow fields are then used to calculate the long-term evolution of the components of the lower level of the food chain in the Black Sea ecosystem. The biogeochemical model used in the calculations is based on the nitrogen cycle and includes a parameterization of the main biological and chemical interactions and processes in the upper layer of the Black Sea. The numerical experiments indicated that the biogeochemical component of the model rather successfully reproduces the main features and evolution trends in the Black Sea ecosystem for the period under consideration: the growth in the phytoplankton biomass during eutrophication and changes in seasonal cycles of the main ecosystem components. Also, the hydrophysical processes were shown to be important for a reliable reproduction of long-term changes in the ecosystem.  相似文献   

2.
Partial pressure of CO2 (pCO2) in surface seawater has been measured in the northeastern Pacific Ocean at Station P and along Line P since 1973. These data have been divided into ‘oceanic’ and ‘coastal/transition’ zones, and the seasonal and interannual variability and the long-term trends for each zone have been examined. The oceanic zone shows little seasonality in surface seawater pCO2, with undersaturation throughout the year. A strong, biologically-driven seasonal cycle is offset by variation in temperature-dependent solubility of CO2. The coastal/transition zone shows a decline in pCO2 from winter–spring through summer and fall that is likely the result of seasonal stratification and convection rather than coastal upwelling. Interannual variability all along Line P is correlated with the multivariate ENSO index (MEI), with lower seawater pCO2 associated with El Niño conditions. Correlations with the Pacific Decadal Oscillation Index are similar but weaker, in part because there are few data prior to the 1976 regime shift. The long-term trend in seawater pCO2 in the oceanic zone is +1.36±0.16 μatm year?1, indistinguishable from the atmospheric growth rate, and varies little among the seasons. In the coastal/transition zone a slow increase in the pCO2 of surface seawater relative to that of the atmosphere has led to increasing undersaturation, particularly in spring. Aliasing of the seasonal and interannual variability due to sampling frequency may explain part of the observed trend in the coastal/transition zone, but real changes in physical or biological processes are also possible and require more detailed study.  相似文献   

3.
Thirteen years (1998–2010) of satellite-measured chlorophyll a are used to establish spatial patterns in climatological phytoplankton biomass seasonality across the California Current System (CCS) and its interannual variability. Multivariate clustering based on the shape of the local climatological seasonal cycle divides the study area into four groups: two with spring-summer maxima representing the northern and southern coastal upwelling zones, one with a summer minimum offshore in mid-latitudes and a fourth with very weak seasonality in between. Multivariate clustering on the seasonal cycles from all 13 years produces the same four seasonal cycle types and provides a view of the interannual variability in seasonal biogeography. Over the study period these seasonal cycles generally appear in similar locations as the climatological clusters. However, considerable interannual variability in the geography of the seasonal cycles is evident across the CCS, the most spatially extensive of which are associated with the 1997–1999 El Niño-Southern Oscillation (ENSO) signal and the 2005 delayed spring transition off the Oregon and northern and central California coasts. We quantify linear trends over the study period in the seasonal timing of the two seasonal cycles that represent the biologically productive coastal upwelling zones using four different metrics of phenology. In the northern upwelling region, the date of the spring maximum is delaying (1.34 days yr−1) and the central tendency of the summer elevated chlorophyll period is advancing (0.63 days yr−1). In the southern coastal upwelling region, both the initiation and cessation of the spring maximum are delaying (1.78 days yr−1 and 2.44 days yr−1, respectively) and the peak is increasing in duration over the study period. Connections between observed interannual shifts in phytoplankton seasonality and physical forcing, expressed as either basin-scale climate signals or local forcing, show phytoplankton seasonality in the CCS to be influenced by changes in the seasonality of the wind mixing power offshore, coastal upwelling in the near-shore regions and basin-scale signals such as ENSO across the study area.  相似文献   

4.
The seasonal variability of surface chlorophyll in the northern Humboldt Current System is studied using satellite data, in situ observations and model simulations. The data show that surface chlorophyll concentration is highest in austral summer and decreases during austral winter, in phase opposition with coastal upwelling intensity. A regional model coupling ocean dynamics and biogeochemical cycles is used to investigate the processes which control this apparently paradoxical seasonal cycle. Model results suggest that the seasonal variability of the mixed layer depth is the main controlling factor of the seasonality. In winter, the mixed layer deepening reduces the surface chlorophyll accumulation because of a dilution effect and light limitation. In summer, biomass concentrates near the surface in the shallow mixed layer and nitrate limitation occurs, resulting in a biomass decrease in the middle of summer. Intense blooms occur during the spring restratification period, when winter light limitation relaxes, and during the fall destratification period, when the surface layer is supplied with new nutrients. Model sensitivity experiments show that the seasonal variations in insolation and surface temperature have little impact on the surface chlorophyll variability.  相似文献   

5.
To understand the confounding fishing effect and physical influence on fish production, catch time series in the East China Seas were analyzed. Principal component analysis partitioned 18 catch time series into interannual trends and variations. While the trends were attributed to growth in fishing effort, variations in catches were related to precipitation and monsoon wind speed. Correlations of catch variations with the physical variables suggest that land-based runoff and monsoon circulation of the diluted coastal water masses are the physical forces dominating catch variability and the influences are largely through the associated nutrient supply on primary production. Runoff inputs nutrients to the coastal ecosystem, while monsoons drive their distribution. Offshore diffusion of the coastal water masses by the summer monsoon increases distribution and efficiency of nutrients and has a positive effect on fish production. Southerly transport of coastal currents alongshore by the winter monsoon confines nutrient distribution and induces nutrient loss from the northern waters. This process reduces overall and northern production, but increases production to the south. A long-term variation in catches was identified, which corresponds to a trend in the local winter monsoon as well as large-scale atmospheric changes. Prediction of the catch variation by the local wind speed suggests that large-scale atmospheric circulation determines the trend in the local winter monsoon, and the local winter monsoon that drives nutrient distribution should be directly responsible for the long-term variation of fish production in the East China Seas.  相似文献   

6.
The seasonal and interannual dynamics of the biomass and spatial distribution of a macrophyte meadow were explored in a Mediterranean coastal lagoon (Albufera des Grau, Balearic Islands) from 2002 to 2007. The dynamics in the main physicochemical variables were also analysed to assess the factors involved in the spatiotemporal variability of the submerged macrophytes.  相似文献   

7.
The response of the Gulf of Alaska (GOA) circulation to large-scale North Pacific climate variability is explored using three high resolution (15 km) regional ocean model ensembles over the period 1950-2004. On interannual and decadal timescales the mean circulation is strongly modulated by changes in the large scale climate forcing associated with PDO and ENSO. Intensification of the model gyre scale circulation occurs after the 1976-1977 climate shift, as well as during 1965-1970 and 1993-1995. From the model dynamical budgets we find that when the GOA experiences stronger southeasterly winds, typical during the positive phase of the PDO and ENSO, there is net large-scale Ekman convergence in the central and eastern coastal boundary. The geostrophic adjustment to higher sea surface height (SSH) and lower isopycnals lead to stronger cyclonic gyre scale circulation. The opposite situation occurs during stronger northwesterly winds (negative phase of the PDO).Along the eastern side of the GOA basin, interannual changes in the surface winds also modulate the seasonal development of high amplitude anticyclonic eddies (e.g. Haïda and Sitka eddies). Large interannual eddy events during winter-spring, are phase-locked with the seasonal cycle. The initial eddy dynamics are consistent with a quasi-linear Rossby wave response to positive SSH anomalies forced by stronger downwelling favorable winds (e.g. southwesterly during El Niño). However, because of the fast growth rate of baroclinic instability and the geographical focusing associated with the coastal geometry, most of the perturbation energy in the Rossby wave is locally trapped until converted into large scale nonlinear coherent eddies. Coastally trapped waves of tropical origin may also contribute to positive SSH anomalies that lead to higher amplitude eddies. However, their presence does not appear essential. The model ensembles, which do not include the effects of equatorial coastally trapped waves, capture the large Haïda and Sitka eddy events observed during 1982 and 1997 and explain between 40% and 70% of the tidal gauges variance along the GOA coast.In the western side of the GOA basin, interannual eddy variability located south of the Alaskan Stream is not correlated with large scale forcing and appears to be intrinsic. A comparison of the three model ensembles forced by NCEP winds and a multi-century-long integration forced only with the seasonal cycle, shows that the internal variability alone explains most of the eddy variance. The asymmetry between the eddy forced regime in the eastern basin, and the intrinsic regime in the western basin, has important implications for predicting the GOA response to climate change. If future climate change results in stronger wintertime winds and increased downwelling in the eastern basin, then increased mesoscale activity (perhaps more or larger eddies) might occur in this region. Conversely, the changes in the western basin are not predictable based on environmental forcing. Eastern eddies transport important biogeochemical quantities such as iron, oxygen and chlorophyll-a into the gyre interior, therefore having potential upscale effects on the GOA high-nutrient-low-chlorophyll region.  相似文献   

8.
文章比较了卫星专用传感器微波成像仪/探测仪(SSM/I&SSMIS)、遥感系统数据集(RSS V7R01)、欧洲中期天气预报中心第5代再分析数据(ERA5)和现代回顾分析的研究与应用第2版数据(MERRA-2) 4种观测和再分析资料在刻画全球大洋水汽气候态中的异同点, 初步探究了不同尺度的大气柱水汽总量(TCWV)的变化特征和长期趋势。研究结果表明, 4种数据TCWV的空间分布、季节和年际变化较为一致。从1988至2018年, TCWV总体呈增加趋势, 其中热带海洋的年际变化显著, 增加趋势较强, 且和厄尔尼诺-南方涛动(ENSO)高度相关。利用再分析数据做短时间(如1991—1997年)的TCWV趋势分析时要慎重使用ERA5和MERRA-2的数据。在研究热带区域长期水汽变化趋势时, 需谨慎使用MERRA-2的数据。  相似文献   

9.
Dissolved organic nitrogen (DON) dynamics in the North Sea was explored by means of long-term time series of nitrogen parameters from the Dutch national monitoring program. Generally, the data quality was good with little missing data points. Different imputation methods were used to verify the robustness of the patterns against these missing data. No long-term trends in DON concentrations were found over the sampling period (1995–2005). Inter-annual variability in the different time series showed both common and station-specific behavior. The stations could be divided into two regions, based on absolute concentrations and the dominant times scales of variability. Average DON concentrations were 11 μmol l−1 in the coastal region and 5 μmol l−1 in the open sea. Organic fractions of total dissolved nitrogen (TDN) averaged 38 and 71% in the coastal zone and open sea, respectively, but increased over time due to decreasing dissolved inorganic nitrogen (DIN) concentrations. In both regions intra-annual variability dominated over inter-annual variability, but DON variation in the open sea was markedly shifted towards shorter time scales relative to coastal stations. In the coastal zone a consistent seasonal DON cycle existed with high values in spring–summer and low values in autumn–winter. In the open sea seasonality was weak. A marked shift in the seasonality was found at the Dogger Bank, with DON accumulation towards summer and low values in winter prior to 1999, and accumulation in spring and decline throughout summer after 1999. This study clearly shows that DON is a dynamic actor in the North Sea and should be monitored systematically to enable us to understand fully the functioning of this ecosystem.  相似文献   

10.
The South African coastline has been invaded by numerous alien species. Rare pre-invasion (1980) and post-invasion datasets (2001 and 2012) exist for Marcus Island, a small land-tied island in Saldanha Bay, South Africa. These snapshot datasets of the island’s intertidal invertebrate community were complemented with monitoring across seasons, from 2014 to 2016. Invertebrate communities were compared among the summers of 1980, 2001, 2012, 2014, 2015 and 2016 to assess interannual differences, while invertebrates and algae were monitored quarter-annually to assess seasonal changes. In addition, the population dynamics of the alien mussel Mytilus galloprovincialis were monitored. Differences in invertebrate communities between consecutive summers were significant but much smaller than changes induced by the arrival of alien species. Invertebrate and seaweed communities differed among years and shore zones but not among seasons, whereas species diversity differed among years, seasons and shore zones, with zones having the strongest influence. The invasion by M. galloprovincialis, and ensuing spatial and temporal variability in its recruitment, emerged as the most important factor influencing community composition, overshadowing interannual and seasonal changes. This work highlights that the impacts of alien species can be distinguished from natural variability by combining long-term monitoring with surveys at finer temporal scales. This is an important step in extending our understanding of the impacts of marine alien species.  相似文献   

11.
A long-term spin-up and a subsequent interannual simulation are conducted for the ocean–ice component of the climate model intercomparison project (CMIP)-class earth system model of the Japan Meteorological Agency/Meteorological Research Institute. This experiment has three purposes: first is to assess the ability of our model with the Coordinated Ocean–ice Reference Experiments (COREs) forcing in reproducing the present ocean-climate; second is to understand the ocean-climate variability for the past 60 years; third is to present an example of evaluating an ocean–ice interannual variability simulation. The Pacific Ocean is focused on for the last two purposes. After integrating for about 1500 years with repeated use of a detrended CORE interannual forcing, the model reaches a quasi-steady state where the present climate is reproduced satisfactorily. Then, the interannual variability simulation is conducted with the retrieved forcing trend and the result is analyzed. The simulation is successful at reproducing the long-term variability in the Pacific and surrounding oceans. Brief analyses of the tropical and mid-latitude upper layer, deep circulation, and the Arctic sea ice are presented. A caveat in treating other parts of the globe is due to the recent intense convection in the Southern Ocean caused by a remarkably increasing trend of the Southern Hemisphere westerly. Overall, the current simulation with our CMIP-class ocean–ice model is shown to be useful for studying the present ocean-climate variability, specifically in the Pacific sector. It could also be used as a benchmark control experiment that facilitates further research, model development, and intercomparison.  相似文献   

12.
A lower trophic level NPZD ecosystem model with explicit iron limitation on nutrient uptake is coupled to a three-dimensional coastal ocean circulation model to investigate the regional ecosystem dynamics of the northwestern coastal Gulf of Alaska (CGOA). Iron limitation is included in the NPZD model by adding governing equations for two micro-nutrient compartments: dissolved iron and phytoplankton-associated iron. The model has separate budgets for nitrate (the limiting macro-nutrient in the standard NPZD model) and for iron, with iron limitation on nitrate uptake being imposed as a function of the local phytoplankton realized Fe:C ratio. While the ecosystem model represents a simple approximation of the complex lower trophic level ecosystem of the northwestern CGOA, simulated chlorophyll concentrations reproduce the main characteristics of the spring bloom, high shelf primary production, and “high-nutrient, low-chlorophyll” (HNLC) environment offshore. Over the 1998–2004 period, model-data correlations based on spatially averaged, monthly mean chlorophyll concentrations are on average 0.7, with values as high as 0.9 and as low as 0.5 for individual years. The model also provides insight on the importance of micro- and macro-nutrient limitation on the shelf and offshore, with the shelfbreak region acting as a transition zone where both nitrate and iron availability significantly impact phytoplankton growth. Overall, the relative simplicity of the ecosystem model provides a useful platform to perform long-term simulations to investigate the seasonal and interannual CGOA ecosystem variability, as well as to conduct sensitivity studies to evaluate the robustness of simulated fields to ecosystem model parameterization and forcing. The ability of the model to differentiate between nitrate-limited, and iron-limited growth conditions, and to identify their spatial and temporal occurrences, is also a first step towards understanding the role of environmental gradients in shaping the complex CGOA phytoplankton community structure.  相似文献   

13.
A three-dimensional ocean biogeochemical model of the tropical Atlantic Ocean was run for more than half a century (1949–2000) in order to characterize the ocean biogeochemical response to variable forcing over this period. The seasonal cycle in the equatorial upwelling zone agrees reasonably well with observations and other published simulations but underestimates phytoplankton biomass under strong upwelling conditions. Away from the equator, modelled nutrient flux and biological production are maximal in each hemisphere's winter season, and appear to be proximately forced by evaporative cooling and wind stirring rather than by Ekman upwelling. The fraction of the total variance associated with the seasonal cycle is considerably smaller for modelled biogeochemical fields than for sea-surface temperature over this long simulation, and much of the biogeochemical variance is associated with interdecadal changes. The model results suggest that the tropical Atlantic became more productive following the Pacific climate shift of 1976 and remained so until about 1989. Summer surface nitrate concentrations during the 1990s were lower than those in the 1980s. The relationship between the equatorial and off-equatorial regimes may have changed following the 1976 event, with equatorial variability dominating the basin-wide variance patterns after 1976.  相似文献   

14.
The variability of the sea surface temperature(SST) in the China seas has been studied in seasonal,interannual and interdecadal scales based on the monthly data of HadISST spanning from 1870 to 2007. The main results obtained are SST in the China offshore changes most actively at the seasonal scale with the intensity diminishing from north to south,as the temperature differences between summer and winter reaching 17 and 4 C in the northern and southern areas,respectively. Moreover,seasonal variation near the coastal regions seems relatively stronger than that far from the coastline;significant interannual variations are detected,with the largest positive anomaly occurring in 1998 in the overall area. But as far as different domains are concerned,there exists great diversity,and the difference is also found between winter and summer. Differed from the seasonal variations,where the strongest interannual variability takes place,resides to the south of that of the seasonal ones in the northern section,nevertheless in the South China Sea,the most significant interannual variability is found in the deep basin;interdecadal changes of summer,winter and annual mean SST in different domains likewise present various features. In addition,a common dominant warming in recent 20 a are found in the overall China offshore with the strongest center located in the vicinity of the Changjiang Estuary in the East China Sea,which intensifies as high as 1.3 C during the past 130 a.  相似文献   

15.
The ecosystem model ERSEM II has been used to hindcast the development of the ecosystem of the North Sea during the years 1955 to 1993. The simulation was driven by the box-aggregated output from a general circulation model of the North Sea of corresponding duration; radiation, river inputs, atmospheric input and boundary conditions at the borders to the Atlantic Ocean and to the Baltic Sea were applied as realistically as possible. The general features of the eutrophication process are reproduced in the hindcast: the coastal areas show strong changes in nutrient concentrations in the hindcast as well as in the observations. Eutrophication not only shows up in the nutrient concentrations, but also in primary production. The simulated spatial distributions of phosphate, nitrate and primary production compare well with the observed ones. In addition, the hindcast simulates considerable trend-like changes of the nutrients in the southern part of the North Sea, where the nutrients are transported from the continental coastal strip to the southern central North Sea. The line from the river Humber to southern Norway separates the region of noticeable anthropogenic influence of riverine and atmospheric input from the northern area, which is mainly influenced by the Atlantic nutrient inflow. The observed annual cycles in the central and northern North Sea are quite well reproduced by the hindcast. The comparison of the hindcast with the long-term observations at two sites in the continental coastal zone of the North Sea shows that the long-term behaviour of phosphate, nitrate and silicate is simulated well. Primary production is increased in summers during the main period of eutrophication, 1975 to 1989, in the hindcast and in the observations. The flagellates at Helgoland, however, experience much more pronounced annual cycles with much less interannual variability in the hindcast than in the observations.  相似文献   

16.
Principal components analysis was used to examine monthly data on sea level obtained from tide-gauge records from the west coast of southern Africa over the period 1957—1975. After a simple correction had removed the effect of local atmospheric pressure, this statistical analysis revealed the structure of nearshore ocean variability over time periods of months to years. The first principal component, accounting for over 55 per cent of the variability in monthly mean sea level, was a near uniform rise and fall everywhere along the west coast of southern Africa. The spectrum of this large-scale structure showed a strong interannual trend, which correlated well with long-term trends of temperature anomaly over the entire South-East Atlantic, and a high-frequency term, which could not be resolved at a time period of one month. The second principal component showed the contrasting response of the northern and southern sites on a seasonal scale, and it could be correlated with seasonal wind data. These results compared well with similar analysis carried out in the North Pacific Ocean.  相似文献   

17.
The seasonality of primary productivity plays an important role in nutrient and carbon cycling. We quantify the seasonality of satellite-derived, oceanic net primary production (NPP) and its interannual variability during the first decade of the SeaWiFS mission (1998 to 2007) using a normalized seasonality index (NSI). The NSI, which is based upon production half-time, t(1/2), generally becomes progressively more episodic with increasing latitude in open ocean waters, spanning from a relatively constant rate of primary productivity throughout the year (mean t(1/2) ~5 months) in subtropical waters to more pulsed events (mean t(1/2) ~3 months) in subpolar waters. This relatively gradual, poleward pattern in NSI differs from recent estimates of phytoplankton bloom duration, another measure of seasonality, at lower latitudes (~40°S–40°N). These differences likely reflect the temporal component of production assessed by each metric, with NSI able to more fully capture the irregular nature of production characteristic of waters in this zonal band. The interannual variability in NSI was generally low, with higher variability observed primarily in frontal and seasonal upwelling zones. The influence of the El Niño–Southern Oscillation on this variability was clearly evident, particularly in the equatorial Pacific, where primary productivity was anomalously episodic from the date line east to the coast of South America in 1998. Yearly seasonality and the magnitude of annual production were generally positively correlated at mid-latitudes and negatively correlated at tropical latitudes, particularly in a region bordering the Pacific equatorial divergence. This implies that increases of annual production in the former region are attained over the course of a year by shorter duration but higher magnitude NPP events, while in the latter areas it results from an increased frequency or duration of similar magnitude events. Statistically significant trends in the seasonality, both positive and negative, were detected in various patches. We suggest that NSI be used together with other phenomenological characteristics of phytoplankton biomass and productivity, such as the timing of bloom initiation and duration, as a means to remotely quantify phytoplankton seasonality and monitor the response of the oceanic ecosystem to environmental variability and climate change.  相似文献   

18.
We consider the decadal evolution of the Black-Sea ecosystem on the basis of a three-dimensional coupled model consisting of the Black-Sea circulation model and a biogeochemical block. The circulation model is based on the widely used POM (Princeton Ocean Model) model. The calculated hydrophysical fields are interpolated then to levels of the biogeochemical model, which covers the upper 150-m layer. We demonstrate the close relationship between the interannual variation of hydrophysical fields and the evolution of the main elements of the ecosystem. The period under consideration (1992–2001) is characterized by the warming of the Black-Sea upper layer, which can be traced by the trend of a growing surface temperature. It follows from the results of modeling that the process of warming is also revealed in the subsurface hydrophysical characteristics and the dynamics of the main elements of the Black-Sea ecosystem.  相似文献   

19.
齐庆华  蔡榕硕 《海洋学报》2017,39(11):37-48
气候变暖背景下,全球平均海洋变暖和海平面上升显著,为人类社会的可持续发展带来巨大挑战。上层海洋热力状况是海平面变化的主导因子之一。本文围绕"21世纪海上丝绸之路"途经海区(文中简称为丝路海区)上层海洋热含量异常的区域性时空特征,分析探讨了丝路海区热比容海平面异常的时空变化、演变特征及可能影响,以期为"21世纪海上丝绸之路"海洋环境安全保障提供服务支撑。结果表明,自20世纪70年代中后期,丝路海区上层(0~700 m)海洋已明显变暖,尤其20世纪90年代中后期增暖幅度显著加大。近60年来,在丝路海区热带海洋中,西太平洋的北赤道流区及以北海域、东海黑潮流域以及南海北部和南部海区、阿拉伯海西北部海域、马来西亚西北部海域及南印度洋部分海域具有长期增暖趋势。热带西太平洋暖池区整体增暖不明显,主要与印度洋中部海域呈反位相变化,且明显受到季节和年际变化的调制。长江口附近沿岸、南海北部沿岸、中南半岛南部沿岸以及阿拉伯海西北部沿岸的近岸海域长期增暖明显,自20世纪90年代中后期,中南半岛东部和西部沿海、澳大利亚西部沿海以及我国东南沿海热比容海平面上升明显。近岸热比容海平面的季节演变对沿海地区社会和经济发展会造成一定影响。此外,东亚夏季风与东海、黄海和渤海热比容海平面的上升显著相关,同时,ENSO、太平洋年代际振荡和印度洋偶极子的发生也均与我国东南沿海和印度洋西部沿海热比容海平面上升明显关联。特别是,气候变暖情形下,各种区域性致灾因子和气候变率的协同影响会对丝路海区海岸带和沿海地区的防灾减灾与社会经济发展带来较大挑战,开展海岸带和沿海地区全球变化综合风险研究成为当前首要任务。  相似文献   

20.
The temporal and spatial distributions of zooplankton biomass and larval fish recorded during 27 months (December 1995-December 1998) off the Pacific coast of central México are analyzed. A total of 316 samples were obtained by surface (from 40-68 to 0 m) oblique hauls at 12 sampling sites using a Bongo net. Two well-defined periods were observed: a pre-ENSO period (December 1995-march 1997) and an ENSO event (July 1997-September 1998) characterized by impoverishment of the pelagic habitat. The highest biomass concentrations occurred at coastal stations during the pre-ENSO period. During the El Niño period no spatial patterns were found in coastal waters. The months with highest biomass were those in which the lowest sea surface temperature (SST) occurred (January-May), and this pattern was also observed during the ENSO period. A typical, although attenuated, seasonal environmental pattern with enhanced phytoplankton (diatoms and dinoflagellates) was prevalent during the El Niño event in nearshore waters. During the El Niño period the phytoplankton was mainly small diatoms (microphytoplankton), while dinoflagellates were practically absent. The most parsimonious generalized linear models explaining spatial and temporal distribution of larval fish species included the ENSO index (MEI), upwelling index (UI) and distance to the coast. The environmental variability defined on an interannual time-scale by the ENSO event and the seasonal hydroclimatic pattern defined by the UI (intra-annual-scale) controlled the ecosystem productivity patterns. The small-scale distribution patterns (defined by a cross-shore gradient) of plankton were related to the hydroclimatic seasonality and modulated by interannual anomalies.  相似文献   

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