AMS 14C chronology of the world's southernmost woolly mammoth (Mammuthus primigenius Blum.)

The world's definite southernmost woolly mammoth record is a molar from Ji’nan (around 36°N), Shandong Province, China. AMS ¹⁴C dating of the specimen, gave a conventional ¹⁴C age of 33,150±250 BP. The period of 40–30 ka BP corresponds to the later phase of the Marine Oxygen Isotope Stage 3 (MIS 3a), recognized as the global interstadial of the last glacial period. However, it is known that the winter monsoon strengthened in Asia during the period 35–33 ka BP, and the age of the woolly mammoth specimen from Ji’nan corresponds to that age. The specimen suggests that this area became cold and dry in 33 ka BP, and grassland or open forest, suitable habitat for woolly mammoths, developed during this short time span. This age is similar to the age of the southernmost woolly mammoth record in Europe, therefore supporting a hypothesis by Porter and An [1995. Correlation between climate events in the North Atlantic and China during the last glaciation. Nature 375, 305–308] that an important component of Chinese palaeoclimates may be linked to changes in North Atlantic oceanic conditions.     

Population-level genotyping of coat colour polymorphism in woolly mammoth (Mammuthus primigenius)

Patterns in the spatial or temporal distribution of genotypes may be indicative of natural selection. Previous work on the woolly mammoth melanocortin-1 receptor (Mc1r) gene identified three polymorphic positions that suggest Pleistocene populations may have harboured both light- and dark-haired mammoths (Rompler et al., 2006, 313: 62). Here, we extend this work and present the first population-level analysis of a functional gene in an extinct species. We genotyped the Mc1r gene in 47 woolly mammoth samples excavated from sites across the central portion of the woolly mammoths’ former range to examine the extent of variation of this polymorphism through time and across space. Only one individual was found to be heterozygous, indicating that the frequency of the ‘light’ mutant allele was very low. We conclude that light-coloured woolly mammoths would have been very rare, and may even have been non-existent if the ‘light’ mutant allele was strongly selected against in its homozygotic form. With the increasing availability of large-scale sequencing technologies, population-level datasets capable of identifying local adaptation will become increasingly attainable.     

The Sevsk woolly mammoth (Mammuthus primigenius) site in Russia: Taphonomic,biological and behavioral interpretations

Excavations at Sevsk, Bryansk Region, Russia, by the Paleontological Institute of the Russian Academy of Sciences in 1988–1991 recovered 3800 bones of woolly mammoth (Mammuthus primigenius Blum.) representing a minimum of 33 individuals. The locality is one of the largest naturally occurring deposits of mammoth remains in Europe and is inferred to be a catastrophic death assemblage. The material includes five skeletons of juvenile mammoths, from 1 month to 6 or 7 years of age, as well as partial skeletons and isolated bones of adult individuals. A femur and humerus of an approximately 10–12-month-old fetus are also among the finds. Morphological features suggest that the Sevsk mammoths belonged to one family group; the age structure and sexual composition of the assemblage do not differ significantly from that of a family group of Modern African elephants. In contrast to other localities in Siberia and central Russia, relatively more (about 45% of individuals) prepubertal animals are preserved at Sevsk. Radiocarbon dates indicate that the mammoths died about 14,000 years ago. Data from diatoms, pollen and rodents, as well as archeological evidence, corroborate this age, and provide the basis for a paleoenvironmental reconstruction at the end of the Valdaian Glaciation in western Russia.     

New data on the morphology of a foetal mammoth (Mammuthus primigenius) from the Late Pleistocene of southwestern Siberia

Twenty-six bones and fragments of bones belonging to a Mammuthus primigenius (Blumenbach, 1799) foetus have been discovered in osteological material coming from the late Pleistocene site of Shestakovo, Chebulinsk area, Kemerovo Province. The age of the site is estimated as 24,000–20,000 years BP (Karginian–Sartanian). The stage of intra-uterine development of the foetus can be estimated as 19–20 months. This foetal skeleton is the most complete of all known foetus materials belonging to this species.     

Extinction of the woolly mammoth (Mammuthus primigenius) and woolly rhinoceros (Coelodonta antiquitatis) in Eurasia: Review of chronological and environmental issues

Kuzmin, Y. V. 2009: Extinction of the woolly mammoth (Mammuthus primigenius) and woolly rhinoceros (Coelodonta antiquitatis) in Eurasia: Review of chronological and environmental issues. Boreas, 10.1111/j.1502‐3885.2009.00122.x. ISSN 0300‐9483. The current evidence for date and environmental preferences of the extinction of two middle–late Pleistocene megafaunal species, the woolly mammoth (Mammuthus primigenius Blum.) and woolly rhinoceros (Coelodonta antiquitatis Blum.), is presented in this review. It is suggested that extinction of these large herbivores in Eurasia was closely related to landscape changes near the Pleistocene–Holocene boundary (c. 12 000–9000 uncalibrated radiocarbon years ago, yr BP), mainly involving the widespread forest formations in the temperate and arctic regions of northern Eurasia and the loss of grasslands crucial to the existence of woolly mammoth and rhinoceros. However, some woolly mammoth populations survived well into the Holocene (up to c. 3700 yr BP), showing that the process of final extinction was fairly complex, with delays in some regions of up to several millennia. The possible role of Palaeolithic humans in the extinction of Late Pleistocene megafauna is also considered.     

Initiation of the last glaciation in northern europe

The bio- and chronostratigraphy of the Eemian interglacial (marine isotope substage 5e) and an Early Weichselian glaciation (5d-a) established from representative and detailed sequences can be correlated with the deep-sea oxygen isotope stratigraphy, ice-core data, sea-level fluctuations and coupled ice sheet-climate models. Biostratigraphic sequences from Fennoscandian key sections are correlated with reference sequences from Estonia and from sections located near or beyond the margins of the last glaciation. Organic sediments previously attributed to Early and Middle Weichselian interstadial periods in Finland are argued to be redeposited and mixed older (last interglacial) material. Pollen and diatom spectra of the undisturbed materials suggest that the Eemian climatic optimum was followed by a continuously cooling climate and a regressive marine level. If only undisturbed sequences are considered, the major climatic fluctuations of the Early Weichselian, apparent in Central and Western Europe, are not apparent in the sequences from the central part of the glaciated terrain. Instead, some sequences are truncated by sediments indicating approaching ice sheets soon after the interglacial. This may imply that the ice sheet grew over Finland during the first Early Weichselian stadial. The preservation of the interglacial beds and the lack of younger non-glacial sediments support the interpretation that the area remained ice-covered until the final deglaciation. During the Early Weichselian, the Norwegian coast was probably occasionally ice free, similar to the coastal zone of Greenland today. The authors' interpretation of the Fennoscandian organic deposits of the last glaciation may also explain similar observations from the central parts of the Laurentide ice sheet.     

Chronology and new research on the Schaefer mammoth (?Mammuthus primigenius) site,Kenosha County,Wisconsin, USA

A summary of previous and new research on the Schaefer mammoth site is presented. The Schaefer site in extreme southeastern Wisconsin, USA was excavated in 1992 and 1993. Seventy-five percent of a mammoth ?Mammuthus primigenius, the woolly mammoth (but comparable as well with M. jeffersonii, the Jefferson mammoth) was recovered. Drifted wood specimens and plant macro fossils were also recovered from the site.Twenty-five specimens of wood have been identified as Picea sp., Picea/Larix sp., or unidentified hardwood. Macro fossils in the form of cones have been identified as black spruce (Picea mariana). These preliminary data do not appear to support the traditional environment inferred for the woolly mammoth and points to a need for study of the animal's attribution to species. The animal was a male, 36 years of age at death.Thirteen AMS 14C assays on bone cluster between 12,290 and 12,570 BP. Sixteen dates on wood specimens intimately associated with bone yield a range of dates from 11,980 to 12,940 BP.The remains exhibit multiple cut and wedge marks interpreted as cultural. Non-diagnostic cultural lithics were recovered. A disarticulated bone pile deposited in a low energy environment is consistent with human interaction.     

First finding of a mammoth female (<Emphasis Type="Italic">Mammuthus primigenius</Emphasis> Blum.) on the Taimyr Peninsula

The first finding of a mammoth female skeleton from the Kastykhtakh River valley, Taimyr Peninsula, is described. The skeleton consists of 104 elements including the skull and lower jaw; small distal limb bones, tail vertebrae, and one tusk are absent. There are teeth of the two last changes in the jaws. The skeleton elements have individual features: sigmoid contact of the low second and third molars, crack in the neural arch of the atlas, and false joint and calluses on places of the rib fractures. The calibrated radiocarbon dating of the tubular bone is 32 070-30 565 yr (Kargin interglacial time).     

Local disappearance of bivalves in the Azores during the last glaciation

The Pleistocene (Eemian) outcrops of Lagoinhas and Prainha, located at Santa Maria Island (Azores), were investigated and their fossil mollusc content reported. These studies revealed that the last glaciation affected two groups of molluscs: the ‘warm‐guest’ gastropods with West African or Caribbean affinities (e.g. Conus spp., Cantharus variegatus, Bulla amygdala, Trachypollia nodulosa) and shallow bivalve species mainly associated with sandy habitats (Ensis minor, Lucinella divaricata, and probably Laevicardium crassum). In this paper we focus on this group of bivalves, which has since locally disappeared from the Azores. We relate the local disappearance of these bivalves in the Azores with the lack of sand in the shelf. The specific characteristics of the Santa Maria shelf combined with the sea‐level drop during the Weichselian prevented deposition of the lowstand deposits and permitted erosion of the previous ones, leaving the shelf without a sediment cover. Copyright © 2008 John Wiley & Sons, Ltd.     

How cold was it for Neanderthals moving to Central Europe during warm phases of the last glaciation?

Precise estimates of mean annual temperature (MAT) for when Neanderthals occupied Central Europe are critical for understanding the role that climatic and associated environmental factors played in Neanderthal migrations and in their ultimate extinction. Neanderthals were continuously present in the relatively warm regions of southern and Western Europe in the Pleistocene but only temporarily settled Central Europe (CE), presumably because of its colder and less hospitable climate. Here, we present a new approach for more spatially and temporally accurate estimation of palaeotemperatures based on the stable oxygen isotope composition of phosphates extracted from animal teeth found at sites linked directly to concurrent Neanderthal occupation. We provide evidence that Neanderthals migrated along the Odra Valley of CE during warmer periods throughout the Upper Pleistocene. The MATs during these migrations were about 6.8 °C for the warm phase of Oxygen Isotope Stage OIS 5a–d (prior to the OIS4 cold event) at ～115–74,000 yr BP and about 6.3 °C during the early OIS 3 warm phase ～59–41,000 yr BP. Our results show that temperatures during these phases peaked 2–4 °C above longer term estimates from ice cores and pollen records. We argue that our approach can provide valuable insights into evaluating the role of climate in human migration patterns in the Pleistocene.     

Fluctuations of the South Patagonian Ice-field during the last glaciation and the Holocene

Mapping along a transect from the southeastern margin of the South Patagonian Ice-field in Torres del Paine National Park (Chile) to the limits of fresh moraines of the last glacial cycle indentified eight glacier advances. The four younger ones have been dated by dendrochronology, tephrochronology and radiocarbon dating. Although the bases of 10 m deep bogs were sampled, close limiting radiocarbon dates were not obtained because bog formation in this rain-shadow area appears not to have commenced until ca.12000 yr ago. The outermost Little Ice Age moraine formed during the seventeenth century and three inner ones were deposited around ad 1805, 1845 and after 1890. Densely vegetated older moraines contiguous with Little Ice Age deposits are possibly of late Holocene age. Tephra from the eruption of Reclus volcano at ca. 11 880 yr BP was incorporated by a readvance that deposited large multiple moraines 10–16 km from the modern ice-front; the oldest basal peat found inside the moraine has been dated to ca. 9200 yr BP. These bracketing dates indicate that some eastern outlet glaciers of the ice-field advanced at a time when some western tidewater outlet glaciers terminated inside their modern limits. This questions the view of J. H. Mercer and other that Patagonian glaciers did not readvance during the late-glacial interval. A stadial event also occurred when the glaciers were some 18–20 km from their modern positions and is closely dated to ca. 11880 yr BP because Reclus pumice flushed down-glacier forms thick upper beds in outwash deltas deposited in proglacial lakes. The four older moraines pre-date the late-glacial eruption of Reclus but are not dated closely. Comparison of their spatial extent with well-dated moraines in the Chilean Lakes Region suggests that they may mark advances culminating at ca. 14000 yr BP, ca. 20000 yr BP and earlier.     

Stratigraphic and geochronologic contexts of mammoth (Mammuthus) and other Pleistocene fauna,Upper Missouri Basin (northern Great Plains and Rocky Mountains), U.S.A.

Mammoth fossils and other Pleistocene vertebrates from the Upper Missouri Basin, in the northern Plains and Rocky Mountains of the western interior of North America, have been dated to the late Pleistocene and are associated with Wisconsinan deposits. Mammoth remains have also been found in older stratigraphic contexts. For example, Mammuthus columbi and other fossils from the Doeden Locality are in pre-Wisconsinan terrace gravels along the Yellowstone River; the deposits are likely Illinoian or Sangamonian. Faunas that appear to be associated with the Wisconsinan interstadial, before the Last Glacial Maximum, are found in intermountain valleys and mountain settings (the Merrell Locality, Blacktail Cave, Natural Trap Cave) and on the Plains in both glaciated and unglaciated regions (Box Creek, Wibaux gravel pit). Localities containing faunas dated to the time interval from about the Last Glacial Maximum to the end of the Younger Dryas chronozone (late Wisconsinan) include the youngest fossil-bearing strata at Merrell, Blacktail Cave, Natural Trap Cave; deposits at Sheep Rock Spring, Indian Creek, MacHaffie, False Cougar Cave, Shield Trap Cave; and mammoths found at Sun River, Glendive, Colby, and the Deer Creek drainage.     

Vegetation and climates in southern Tasmania since the last glaciation

Enclosed basins (glacial and nonglacia) of Tasmania contain the most comprehensive record in Australia of trends in a regional vegetation and climate since the late Pleistocene. Seven pollen sequences, each continuous and extending back at least 10,000 years, are used to reconstruct the history of postglacial vegetation and climate in Southern Tasmania (42°S–43°30′S). Interpretations are supported by a study of the modern pollen rain. Postglacial climates in Tasmania were characterized by a strong west-to-east decrease in precipitation. During the late Pleistocene, climates were markedly colder and drier than at present, and the vegetation was largely devoid of trees. A major rise in temperature between ca. 11,500 and 9500 yr B.P., accompanied by rising effective precipitation, resulted in the expansion of Eucalyptus, then other trees, across Tasmania. This warming trend may have been temporarily reversed during the early postglacial. Dry climates delayed the development of forest in inland eastern Tasmania until after ca. 9500 yr B.P. There is no evidence for a major change in climate since this temperature rise. Two broad phases of development have occurred within the postglacial forests. The first was an early Holocene phase during which Nothofagus cunninghamii cool temperate rain forest developed in western Tasmania and on the slopes of mountains in central and southeastern Tasmania. Eucalyptus sclerophyll forests developed in eastern Tasmania and have remained dominant there since. By ca. 7800 yr B.P. rain-forest communities were established beyond present-day limits. The second phase was a mid to late Holocene phase during which forests and alpine vegetation became more open in structure, leading to the re-expansion of Eucalyptus and shade-intolerant species. During the early to mid Holocene, climates in Southern Tasmania were wetter and (? then) warmer than at present. Maximum and minimum dates for this “optimum” are 8000 and 5000 yr B.P. Since then, climates have become increasingly rigorous, possibly through an increased incidence of inequable “weather types” leading to an increase in the frequency of drought and frost. Structural changes in the postglacial vegetation of Southern Tasmania closely parallel those at equivalent latitudes in New Zealand and Chilean South America, hence are likely to reflect the same primary cause.     

Radiocarbon dates of late quaternary mammals in the Archangelsk Region and their contribution to reconstructions of the last glaciation in Eastern Europe

Twelve new AMS (Accelerator Mass Spectrometry) dates of large Quaternary mammal remains were reported: mammoth (Mammuthus primigenius, bison (Bison priscus), and musk ox (Ovibos pallantis) found in the Archangelsk Region. The absolute age of the identified samples varies from 46 000 to 22 000 calibrated years ago. These data suggest that a substantial part of the Archangelsk Region was not covered by ice during the indicated time interval.     

Mass balance and sliding velocity of the Puget lobe of the cordilleran ice sheet during the last glaciation

An estimate of the sliding velocity and basal meltwater discharge of the Puget lobe of the Cordilleran ice sheet can be calculated from its reconstructed extent, altitude, and mass balance. Lobe dimensions and surface altitudes are inferred from ice limits and flow-direction indicators. Net annual mass balance and total ablation are calculated from relations empirically derived from modern maritime glaciers. An equilibrium-line altitude between 1200 and 1250 m is calculated for the maximum glacial advance (ca. 15,000 yr B.P.) during the Vashon Stade of the Fraser Glaciation. This estimate is in accord with geologic data and is insensitive to plausible variability in the parameters used in the reconstruction. Resultant sliding velocities are as much as 650 m/a at the equilibrium line, decreasing both up- and downglacier. Such velocities for an ice sheet of this size are consistent with nonsurging behavior. Average meltwater discharge increases monotonically downglacier to 3000 m³/sec at the terminus and is of a comparable magnitude to ice discharge over much of the glacier's ablation area. Palcoclimatic inferences derived from this reconstruction are consistent with previous, independently derived studies of late Pleistocene temperature and precipitation in the Pacific Northwest.     

Depositional evidence for marginal oscillations of the Irish Sea ice stream in southeast Ireland during the last glaciation

Late Devensian/Midlandian glacial deposits on the southeast Irish coast contain a record of sedimentation at the margins of the Irish Sea ice stream (ISIS). Exposures through the Screen Hills reveal a stratigraphy that documents the initial onshore flow of the ISIS ('Irish Sea Till') followed by ice stream recession and readvances that constructed glacitectonic ridges. Ice-contact fans (Screen Member) were deposited in association with subglacial deformation tills and supraglacial/subaqueous mass flow diamicts. In SE Ireland, the ISIS moved onshore over proglacial lake sediments which were intensely folded, thrust and cannibalized producing a glacitectonite over which laminated and massive diamictons were deposited as glacitectonic slices. Ice marginal recession and oscillations are documented by: (a) ice-proximal, subaqueous diamict-rich facies; (b) isolated ice-contact glacilacustrine deltas; (c) syn-depositional glacitectonic disturbance of glacilacustrine sediments and overthrusting of ice-contact outwash; (d) offshore moraine ridges; and (e) changing ice flow directions and facies transitions. Diagnostic criteria for the identification of dynamic, possibly surging, ice-stream margins onshore include thrust-block moraines, tectonized pitted outwash and stacked sequences of glacitectonites, deformation tills and intervening stratified deposits. In addition, the widespread occurrence of hydrofracture fills in sediments overridden and locally reworked by the ISIS indicate that groundwater pressures were considerably elevated during glacier advance. The glacigenic sediments and landforms located around the terrestrial margins of the ISIS are explained as the products of onshore glacier flow that cannibalized and tectonically stacked pre-existing marine and glacilacustrine sediments. Localized tectonic thickening of subglacially deformed materials at the former margins of glaciers results in zones of net erosion immediately up-ice of submarginal zones of net accretion of subglacial till. The more stable the ice-stream margin the thicker and more complex the submarginal sedimentary stack.     

Interactions between the Greenland Ice Sheet and the Liverpool Land coastal ice cap during the last two glaciation cycles

The sedimentary record from the Ugleelv Valley on central Jameson Land, East Greenland, adds new information about terrestrial palaeoenvironments and glaciations to the glacial history of the Scoresby Sund fjord area. A western extension of a coastal ice cap on Liverpool Land reached eastern Jameson Land during the early Scoresby Sund glaciation (≈the Saalian). During the following glacial maximum the Greenland Ice Sheet inundated the Jameson Land plateau from the west. The Weichselian also starts with an early phase of glacial advance from the Liverpool Land ice cap, while polar desert and ice‐free conditions characterised the subsequent part of the Weichselian on the Jameson Land plateau. The two glaciation cycles show a repeated pattern of interaction between the Greenland Ice Sheet in the west and an ice cap on Liverpool Land in the east. Each cycle starts with extensive glacier growth in the coastal mountains followed by a decline of the coastal glaciation, a change to cold and arid climate and a late stage of maximum extent of the Greenland Ice Sheet. Copyright © 2005 John Wiley & Sons, Ltd.     

History of sedimentation in the northern Sea of Okhotsk during the last 1.1 Ma

Results of the lithogeochemical and mineralogical investigations of the superlong (46 m) core MD01-2415 recovered from the northern Sea of Okhotsk during the cruise of the R/V Marion Dufresne (France) in 2001 are described. Sediments from the core bottom are 1.1 Ma old (Nürnberg and Tiedemann, 2004). Lithological characteristics, physical properties, grain size composition, and chemical composition (based on the neutron activation analysis) of clay minerals are studied. Together with published data, our original data provide insights into the cyclicity, trend, and local features of sedimentation in the study region.