太白山高山林线植被的数量分析

通过对太白山南坡高山林线（ａｌｐｉｎｅｔｉｍｂｅｒｌｉｎｅ）及其附近的草本植物群落的聚类,排序,物种多样性以及生态种组等研究分析表明：１．太白山高山植物群落的物种多样性随着海拔高度的升高而增加;２群落交错带（ｅｃｏｔｏｎｅ）的物种多样性要比相邻群落内部高;３．在太白山高山带,随着海拔升高,种一面积相关值呈波动增加,但在群落交错带比相邻群落的内部小。     

太白山高山带环境特征

对比国内外关于高山带景观划分的多种观点，采用气候树线以上部分为高山带的划分方法，重点讨论太白山高山带自然环境，包括第四纪冰川遗迹和现代冰缘作用影响下的高山、亚高山地貌，研究区内主要气候要素的分布状、特别气温和降水的时空分配分布格局，土壤有机质的空间分异，以及在这种环境条件下高山带的植物区系和植被分布格局。太白山高山带的各种环境要素之间密切相关，是表现特殊的自然综合体。在全球气候变化背景下，其自然环境特点及变化值得密切关主。     

中国东部暖温带高山林线乔木的光合作用及其与环境因子的关系

高山林线对于气候变化的影响非常敏感。气候变化首先从个体的水平影响林线的内部结构，然后才影响到作为整体的林线的推移，因此研究乔木个体生理活动与气候条件之间的关系，对于研究高山林线对气候变化的响应具有重要意义。本文通过对们于中国东部暖温带的五台山、关帝山、太白山森林上限附近白杆（Picea meyeri）、华北落叶松（Larix principis-ruprechtii)和太白红杉（L.chinensis)生理活动与环境因子的野外测定，分析了光合作用与环境因子之间的关系，得出以下初步结论：（1）研究区内林线乔木的光合作用不存在单一的限制因子；（2）有关研究表明，光照与温度共同作用于高山林线乔木的光合作用，在光照较低的情况下，温度成为光合作用的限制因子，本研究进一步证实了这一现象；（3）树线、林缘、郁闭林内三种不同生境条件下乔木光合作用的对比表明，林缘的生境条件最适合植物光合作用。虽然林缘的湿度条件中等，但良好的光照和温度条件对光合作用有促进作用。     

Dynamics of the Alpine Treeline Ecotone under Global Warming: A Review

The alpine treeline ecotone is defined as a forest-grassland or forest-tundra transition boundary either between subalpine forest and treeless grassland, or between subalpine forest and treeless tundra. The alpine treeline ecotone serves irreplaceable ecological functions and provides various ecosystem services. There are three lines associated with the alpine treeline ecotone, the tree species line (i.e., the highest elevational limit of individual tree establishment and growth), the treeline (i.e., the transition line between tree islands and isolated individual trees) and the timber line (i.e., the upper boundary of the closed subalpine forest). The alpine treeline ecotone is the belt region between the tree species line and the timber line of the closed forest. The treeline is very sensitive to climate change and is often used as an indicator for the response of vegetation to global warming. However, there is currently no comprehensive review in the field of alpine treeline advance under global warming. Therefore, this review summarizes the literature and discusses the theoretical bases and challenges in the study of alpine treeline dynamics from the following four aspects: (1) Ecological functions and issues of treeline dynamics; (2) Methodology for monitoring treeline dynamics; (3) Treeline shifts in different climate zones; (4) Driving factors for treeline upward shifting.     

大兴安岭北部大白山高山林线动态与气候变化的关系

高山林线植被对气候变化十分敏感,已成为全球变化研究的热点.研究了大兴安岭北部大白山高山林线的树木生长和群落更新动态及其与气候变化、火干扰等因素的关系.结果表明,林线树木的生长对气候变化十分敏感,但其敏感性随着海拔的降低而减弱;在高海拔,林线树木的径向生长与上年生长季后期(8月)降水负相关,而与上年初秋(9月)温度正相关,这限制作用随着海拔的降低而逐渐消失;与此相反,低海拔树木生长与当年冬末春初(3月)的温度负相关,但随着海拔上升这种限制作用消失.分析结果还表明,本地区的群落更新主要受火干扰驱动,而与气候变化没有显著关系.不同树种在火灾后更新的时间存在差异,这种差异又因海拔的不同而异,反映出不同树种的更新策略及对环境变化适应能力的差异.大兴安岭北部的高山林线在树木生长对气候变化的敏感性、与气候因子的关系及群落更新动态等方面均与干旱区林线有明显的差异,这些差异与气候条件、树种、更新驱动力等方面的不同有密切的关系.     

天山北坡高山林线分布的生态地理特征

综合利用多源遥感影像和实地勘察资料识别天山北坡高山林线分布格局,结合区域气象数据和土壤理化性质,分析天山北坡林线分布的生态地理特征。结果表明:①天山北坡林线分布高度大约在2 600～2 850 m,从西向东林线分布高度呈上升趋势,奇台至巴里坤段林线高度上升最为显著;伊犁河谷段与玛纳斯段林线垂直宽度较宽。②影响天山北坡林线分布高度的关键气候因子为生长季温度（如年生物学温度3.35 ℃,最热月均温10.49 ℃,生长季均温8.26 ℃）,特别是年生物学温度,能较好的指示天山北坡高山林线分布位置,且各气候指标均在全国均值范围之内,而影响巴音布鲁克地区森林发育的主要原因为冬季低温干旱。③伊犁林线过渡带和玛纳斯林线过渡带有机质、全氮及全磷的含量最高;酸碱性大致以阜康林线为界,向西呈酸性,向东呈碱性;土壤营养物质主要分布于表层（0～10 cm）,深层(30～80 cm)含量低且变化不显著,具有明显的“表聚现象”;下层土壤pH值从西向东逐渐由弱酸性向弱碱性过渡;电导率空间变异性较强,各层变化特征不显著。     

中国高山林线的分布高度与气候的关系

通过研究我国高山林线的分布高度沿纬度、经度的变化格局,和对高山林线处的温度和基带降水等气候指标的分析,对我国高山林线分布高度与气候因子的关系进行探讨。结果表明：(1) 我国高山林线高度表现出明显的纬向和经向变化,总体趋势是：在北纬30^o以北,高山林线高度随纬度升高而下降,下降速率为112 m/度左右;在30^oN以南,则表现出较大的东西部差异：在东部,高山林线高度变化不明显,西部则随纬度增加呈上升趋势。在相似的纬度上,高山林线高度呈现出从东向西升高的趋势。高山林线在藏东南的洛隆、丁青、工布江达一带 (约29^o~32^oN,94^o~96^oE) 达到4 600 m,为世界最高林线高度,并以此为中心向四周降低。(2) 影响高山林线高度的主导气候因子为生长季温度条件。我国高山林线高度的温度指标为年生物温度3.5 ^oC,温暖指数14.2 ^oC·月,生长季平均温度8.2 ^oC。该指标相应海拔高度的地理差异,导致了我国高山林线高度的纬向、经向变化,和从沿海到内陆林线高度的差异。(3) 降水对高山林线高度有显著影响。在中高纬度地区,相同纬度上干旱区域的高山林线高于较湿润区域,降水量是通过温度间接作用于林线高度的。     

The Role of Thresholds and Fine-Scale Processes in Driving Upper Treeline Dynamics in the Bighorn Mountains,Wyoming

This research investigates whether abrupt changes indicative of a threshold response occurred in the spatial and temporal patterns of tree establishment within upper treeline ecotones in the Bighorn Mountains of north-central Wyoming, with a particular focus on assessing whether treeline advance occurred. Dendroecological techniques were used to reconstruct the spatiotemporal patterns of tree establishment at multiple spatial scales (local and landscape). Increases in the elevational extent of treeline and tree density above timberline were reconstructed for each decade of the 20^th century. Regime-shift analysis was used to detect threshold changes in tree establishment. Marked increases in tree establishment and density occurred at local and landscape scales during the 20th century, particularly during the post-1970 period and in areas above timberline. Varying degrees of treeline advance occurred during this time, yet establishment upslope from timberline was often contingent on the availability of sheltered microsites in the lee of boulders. This highlights the importance of both broad-scale climate inputs and fine-scale site conditions in governing the rate and pattern of tree establishment in high-elevation treeline environments. Overall, the abrupt changes in tree establishment over time and space since 1970 suggest that bioclimatic thresholds can strongly influence ecotonal dynamics at upper treeline.     

高山树线交错带的景观格局与生态过程

由于对气候变化的敏感性, 近年来高山树线交错带成为全球变化研究中的热点问题。高 山树线交错带所指示的是一个复杂生态系统的特征, 反映出了由低处植被向高山植被的转化, 其 景观格局是综合了种子定居, 树木生长以及树木繁殖等多个树木生态过程和特定高山环境下的 地理特征格局的适应结果。一方面, 树木通过自身生理生态学上的调整对高山极端环境进行适 应。一方面, 高山特定地理环境特征对树线的景观格局进行影响, 因而高山树线交错带成为研究 景观格局- 生态过程相互关系的一个重要载体。通过运用3S 技术, 可以将高山树线交错带研究中 的树线景观结构和生理生态过程结合在一起, 并验证在其景观格局形成过程中地理特征和人为 因子的相对重要性。     

Geomorphic Controls of Spatial Pattern and Process at Alpine Treeline*

This study examines the role of surface geomorphic features in tree establishment at the alpine treeline in Glacier National Park, Montana, through the presentation of a multiscale, conceptual model of biogeomorphic relationships at alpine treeline. Empirical observations gathered through a multiscale field methodology over three summers serve as a base for the model. The model highlights the importance of surface geomorphic features, specifically boulders and terrace risers, in creating favorable local site conditions, largely by protecting seedlings from wind. The sheltering effect of surface features enables initial seedling establishment, and in some cases survival, above current treeline locations, thereby initiating a positive feedback effect that encourages subsequent tree establishment. Geomorphic features are therefore important in linking scales of pattern and process at the alpine treeline ecotone.     

‘WHY FAMILIES MOVE’: A RE-EXAMINATION

Abstract

This study examines the role of surface geomorphic features in tree establishment at the alpine treeline in Glacier National Park, Montana, through the presentation of a multiscale, conceptual model of biogeomorphic relationships at alpine treeline. Empirical observations gathered through a multiscale field methodology over three summers serve as a base for the model. The model highlights the importance of surface geomorphic features, specifically boulders and terrace risers, in creating favorable local site conditions, largely by protecting seedlings from wind. The sheltering effect of surface features enables initial seedling establishment, and in some cases survival, above current treeline locations, thereby initiating a positive feedback effect that encourages subsequent tree establishment. Geomorphic features are therefore important in linking scales of pattern and process at the alpine treeline ecotone.     

山体效应对北半球林线分布的影响分析

通过搜集整理了北半球516 个林线数据, 结合WorldClim 气象数据计算了林线数据点上的大陆度, 并依据SRTM高程数据提取了林线处的山体基面高度(作为山体效应的代用因子), 然后以纬度、大陆度和山体基面高度为解释变量, 建立三元回归模型。结果表明:线性回归模型的判定系数R²为0.904, 二次回归模型的R²高达0.912。相比先前不考虑基面高度的林线分布模型(R² = 0.79), 纳入了山体基面高度的林线分布模型能够更加有效的拟合半球尺度的林线分布; 结果还表明, 山体基面高度对北半球林线高度分布的贡献率达到了48.94% (p =0.000), 而纬度和大陆度分别为45.02% (p = 0.000) 和6.04% (p = 0.000)。这揭示了山体效应对半球尺度林线分布具有重要的影响。基面高度在北美洲地区对林线高度的贡献率最大(50.49%, p=0.000), 在欧亚大陆东部地区为48.73% (p = 0.000), 在欧亚大陆西部地区为43.6% (p=0.000)。这一结果说明山体效应对林线分布高度的影响虽有区域差异, 但都有较高的贡献率。     

Sigmoid wave transitions at alpine treeline

Ecotones that are controlled by a single environmental gradient or a set of correlated environmental factors are hypothesized to show a sigmoid wave form in percentage cover as the ecotone is crossed. This study investigates the sigmoid wave hypothesis at the alpine treeline in Glacier National Park, Montana, USA. We find that approximately half of the sampled locations exhibit a sigmoid wave form in percentage cover across the ecotone. These results indicate that in some cases treeline is controlled by a single environmental gradient or a set of correlated environmental factors. However, in other cases, additional factors that are not correlated with the primary controlling gradient act to change the form of the transition. We investigated surficial geology as a possible controlling factor that would not be correlated with the primary climatic gradients found along transects across the alpine treeline. In at least some of the cases, surficial geology type was shown to be significantly associated with differences between sigmoidal and non‐sigmoidal transitions. These results support previous evidence for sigmoid wave transitions at the boreal forest—tundra ecotone, but also show that the alpine treeline case is more complex. We argue that in cases where ecotones are used as sites for monitoring the effects of climate change on vegetation, care should be taken to utilize only sites that exhibit a sigmoidal transition so as to minimize the effects of non‐climate‐related controlling factors on the interpretations.     

Comparing the seasonal variation of parameter estimation of ecosystem carbon exchange between alpine meadow and cropland in Heihe River Basin, northwestern China

Grasslands and agro-ecosystems occupy one-third of the global terrestrial area. However, great uncertainty still exists about their contributions to the global carbon cycle. This study used various com...     

长白山北坡林线长白落叶松与岳桦对环境适应的对比分析

以长白山北坡林线长白落叶松（Larix olgensis）和岳桦（Betula ermanii）为研究对象,通过野外样方调查、树轮技术分析,采用趋势线模型拟和、单因素方差等统计方法从各种参数分析长白落叶松和岳桦径向树高生长的对比关系并揭示2个种群动态对环境适应性的对比差异。结果显示：垫状生长形的长白落叶松海拔最高（均值：2 017 m）,其次为旗形树（2 005 m）,正常形态的海拔最低（1 992 m）,所在生境差异显著（p < 0.05）。正常形态的长白落叶松生长速度大于岳桦（树高方程斜率：长白落叶松0.750 9>岳桦0.253 6）,随着旗形树的出现,速度相似（斜率：长白落叶松0.375 4>岳桦0.253 6）,变成垫状后,岳桦占优。长白落叶松受到压制种群扩张速度不如岳桦,长白山林线上侵表现为以岳桦种群为主力,长白落叶松仍是岳桦主要的伴生树种。     

岷江上游林树下线地理分布格局及其空间移动特征

林树下线是干旱河谷背景下一种特殊的地理生态现象,针对林树下线的定位和比对研究是揭示山地环境自然变化过程与人类活动环境效应的有效途径之一。选取岷江上游林树下线为研究对象,采用遥感和GIS聚类分析方法,基于1999-2009年两个时期、聚落区和非聚落区两类区域的比对研究,揭示林树下线的空间分异特征及其移动规律。结果表明：① 林树下线分布具有集聚性特征,主要分布于海拔1400~3800 m V型河谷谷肩上部,这一特征与山区降水随海拔升高的梯度变化相适应;② 6°~15°坡度是林树下线分布的峰值,林树下线随着坡度递增分布明显减少;③ 林树下线坡向分异显著,平均高程表现为南坡高于北坡、东坡高于西坡,最小值为西北方向,源于阴坡水分/湿度更适于植被生长;④ 聚落区林树下线10年间向低海拔空间移动达108.6 m,而非聚落区仅为38 m,这一结果可为定量评估1999年起实施的退耕还林等生态建设工程成效提供科学判据。     

藏东南色季拉山急尖长苞冷杉林线地带地上生物量随海拔的变化特征

从亚高山森林到高山林线,逐渐严酷的生长环境影响了生物量的积累和分配.本研究采用相对生长法和样方收获法估算了藏东南色季拉山急尖长苞冷杉林在禾同海拔(4 190 m,4 270 m和4 326 m)的地上生物量,分析了群落中不同层次地上生物量、各器官生物量及其分配比例随海拔的变化格局.结果表明:1.急尖长苞冷杉群落地上生物量在181～284 t/hm2之间,其中,乔木层在11～-248 t/hm2,灌木层为35～62 t/hm2,草本层为0.9～1.5 t/hm2;2.群落地上生物量随海拔升高急剧降低,降低的平均幅度为73.1 t/(hm2·100 m);3.随海拔升高,群落地上生物量分配到非光合器官(树干和枝条)的比例显著降低,分配到叶的比例呈增加趋势.随海拔的增加,急尖长苞冷杉群落通过增加叶生物量所占比例(高叶寿命)以延长养分在植物体内的存留时间,提高生态系统的养分利用效率,从而适应高海拔地区的低温胁迫环境.     

Precipitation and Temperature Estimation Error at Alpine Treeline Ecotones Using the Mountain Climate Simulator Model (MT-CLIM)

In order to improve modeling of alpine treeline responses to climate change, estimations of snowfall at treeline sites are needed. The MT-CLIM climate model was evaluated for this purpose by extrapolating precipitation and temperature from standard weather stations at lower elevations to 30 alpine SNOTEL study sites across the western United States. Quantification of the topography between the base stations and the SNOTEL sites was used in inverse distance weighting and compared to straight-line weighting. The predicted temperature and precipitation under different weighting methods were compared to observed data over three months during the winter of 2006-2007. The errors were mapped and their spatial pattern analyzed. Error patterns indicate strong gradients, particularly in the Pacific Northwest, that are suggestive of areas where additional characteristics of atmosphere-land interactions and boundary layer climatology need to be considered in modeling applications.     

祁连山高山灌丛生物量及其分配特征

分布于青海云杉林上线的祁连山高山灌丛是祁连山水源涵养林效益最佳林型,研究高山灌丛生物量是生态系统生产力的重要体现,也是群落结构和功能的主要测度之一。以祁连山排露沟流域3 300～3 700 m高山灌丛为研究对象,采用标准地、样方收获法以及壕沟挖掘方法,对不同海拔高度10个固定样地灌丛生物量,器官生物量进行调查,分析了不同海拔梯度灌丛的叶、枝、须根、细根、粗根烘干重数据及其生物量分配特征。结果表明：祁连山高山灌丛总生物量为12 869.39±3 306.16 kg/hm²(平均值±标准差,n=10),其中器官分配以枝生物量所占比例最高,达32.21%,叶、须根、细根、粗根的比例分别为15.70%、14.06%、11.13%和26.90%。不同海拔梯度灌丛生物量器官分配比例差异较大,地上生物量平均为6 097.17kg/hm²,地下生物量平均为6 772.22kg/hm²,不同海拔根茎比在0.56～1.93之间变化,海拔3 500 m处根茎比达最大。祁连山高山灌丛生物量与海拔呈现显著的负相关（R² = 0.898 7,p＜0.01）,随着海拔的升高,灌丛总生物量,地上以及地下生物量均呈现下降的趋势。研究结果可为内陆河流域生态环境的保护及其高山灌丛对全球气候变化的响应研究提供重要理论依据和应用资料。     

Spatial estimation of long-term seasonal and annual groundwater resources: application of WetSpass model in the Werii watershed of the Tekeze River Basin,Ethiopia

Treelines are widely studied worldwide in relation to climate changes because they are hypothesized to be sensitive climate proxies. However, forest treeline expansion toward higher altitudes may be influenced both by a warming climate and by other factors, such as surface morphology and, in the European Alps, the decline in alpine farming. Our results from five valleys in the inner and peripheral regions of the Alps show that present-day treeline altitudes mostly depend on anthropogenic and orographicgeomorphologic factors. Climatic treelines are limited to steep and inaccessible slopes, and occur at higher altitudes and farther from mountain peaks in the inner regions than in the peripheral regions of the mountain range. Looking for sites in which to study treeline responses to climate change, we recommend investigating the inner regions of the Alps where treelines are farther from human disturbances and from geomorphologic constraints, potentially resulting in freer upward shifts under warmer temperature conditions. We also found that, in the valleys selected, human disturbance is mainly concentrated about 165 m below non-climatic treelines, suggesting a homogeneous influence on treelines, regardless of geographic position.