Additional coralline algae from the Lower Miocene to Late Holocene sediments of the Porbandar Group,Gujarat

The Neogene-Quaternary sediments well exposed in the Porbandar Basin are characterized by coralline algae. The limestone, calc-arenites and marls are the dominant lithology of this sequence. The present paper documents ten species of coralline algae. Among these, the nongeniculate coralline algae are represented by four species and geniculate coralline algae include six species. The sediments yielding this algal flora range in age from the Lower Miocene to Late Holocene.     

Coralline algae from the Aramda Reef Member of the Chaya Formation,Mithapur, Gujarat

The Quaternary sediments of the Aramda Reef Member of the Chaya Formation exposed in the Mojap coast near Mithapur, Gujarat are characterized by well-developed coralline algal build-ups. These algal build-ups are exceptionally rich in coralline algae and corals. In the present paper, thirteen species belonging to eight genera of coralline algae are described. Out of these, seven species (Titanoderma nataliae, Lithophyllum nitorum, Lithophyllum quadratum, Spongites sp. Brandano et al., 2005, Sporolithon lvovicum, Mesophyllum fructiferum and Lithothamnion praefruticulosum) are the new records for India. Four species (Titanoderma pustulatum, Sporolithon intermedium, Mesophyllum commune and Phymatolithon sp.) are first time recorded from the study area. Among the major framework builders of coralline algae of the Aramda Reef Member are Lithophyllum, Titanoderma, Sporolithon, Mesophyllum and Lithothamnion.     

Generic distinguishing characteristics and stratigraphic ranges of fossil corallines: An update

Corallines or coralline algae are carbonate secreting and strongly calcified red algae of the order Corallinales of division Rhodophyta. Architecturally, the corallines have two groups, the nongeniculate and geniculate coralline forms. Corallines are used as a potential tool for paleoecology, paleoenvironments and paleobathymetry. Coralline algae are builder of porous and permeable carbonate reservoir rocks for hydrocarbon and reefs rich in hydrocarbon. The old approach, ca. prior to 1977, of taxonomy of fossil coralline genera has been replaced by the modern approach that established during the last decade using certain distinguishing features such as arrangement of basal filaments, cell fusions, conceptacle perforations and orientation of filaments around conceptacles of living corallines. The earliest confirmed fossil record of coralline algae is from the Hauterivian (Early Cretaceous) and from the Hauterivian to the Pleistocene 9 nongeniculate coralline genera, namely Distichoplax, Lithophyllum, Lithoporella, Lithothamnion, Mesophyllum, Neogoniolithon, Phymatolithon, Spongites and Sporolithon and 7 geniculate genera, viz. Amphiroa, Arthrocardia, Calliarthron, Corallina, Jania, Metagoniolithon and Subterraniphyllum having different stratigraphic ranges are unequivocally known as fossils. After 1977, we do not have a comprehensive publication giving the generic distinguishing characteristics and stratigraphic ranges of both nongeniculate and geniculate corallines. The present paper gives an update of distinguishing characteristics of fossil coralline algal genera and their stratigraphic ranges.     

Neogene-quaternary calcareous algae from Saurashtra basin,Western India: Implications on paleoenvironments and hydrocarbon exploration

The Cenozoic and Quaternary sediments of Saurashtra basin encompass signatures of sea level fluctuations, environments of deposition and presence of hydrocarbons. These sediments are divided into seven formations that in order of superposition are: Deccan Trap (late Cretaceous to Eocene), Gaj (early Miocene), Dwarka (middle Miocene to early Pliocene), Miliolite (middle Pleistocene), Chaya (late Pleistocene to late Holocene), Katpur (middle Holocene) and Mahuva Formation (late Holocene). The fossil calcareous algae are significant constituents of these sediments and in all 88 species so far have been recorded by different workers from three prominent areas: Dwarka-Okha area (38 coralline algal species:28 nongeniculate and 10 geniculate), Porbandar area (37 coralline algal species:25 nongeniculate and 12 geniculate species) and Diu area (13 calcareous algal species:6 nongeniculate coralline algal species, 5 geniculate coralline algal species; 1 dasycladalean algal species and 1 halimedacean algal species) of the Saurashtra basin. The present paper provides a checklist of these 88 calcareous algal species documented from the different formations of this basin and their implications for paleoenvironments and petroleum exploration.     

柴北缘塔塔楞环斑花岗岩的岩相学和地球化学特征

塔塔楞环斑花岗岩是柴达木盆地北缘一个古生代复式岩体。该环斑花岗岩在主量元素上, 具有富SiO2、K2O和FeO*, 高K2O/Na2O和FeO*/MgO的特点, 其平均值分别为72.86％、5.17％和3.35％,2.22和10.73;∑REE在279.1×10－6～300.3×10－之间,（La/Lu）N为11.32～13.14,δEu在0.28～0.38之间;Ba、Rb、Pb、Th等元素的含量高,而Sr、Cr、Ni、V等元素的含量低。与经典环斑花岗岩相比,二者在岩相学上相同,在地球化学上也有相似之处,即该岩体也表现为高钾、富铁和LREE,Eu亏损的特征,但部分微量元素与典型环斑花岗岩有一定差异。岩体的形成时代和区域构造背景的综合分析显示,该岩体可能是早古生代后碰撞或后造山伸展构造环境下的产物。     

西沙海域西琛1井生物礁性质及储层岩石学特征

西沙海域西琛1井生物礁主要是由红藻门壳状珊瑚藻、有节珊瑚藻和绿藻门仙掌藻等钙藻组成的植物礁,其次为珊瑚礁。礁相类型主要有礁核相和礁后泻湖相。岩石矿物成分单一,以碳酸盐矿物为主,包括低镁方解石和铁白云石;结构组分有生物骨架、粒屑、泥晶和亮晶;结构类型有生物格架结构、生物障积结构、生物节片结构、生物捆扎结构和生物粘结结构。岩石类型包括骨架石灰岩/白云岩、粘结石灰岩/白云岩、粒屑石灰岩/白云岩。储集空间类型有粒间孔、生物体腔孔和藻架孔等原生孔隙和铸模孔、裂缝、颗粒内溶蚀孔、藻类溶孔和扩大的粒间溶孔等次生孔隙。孔隙组合类型以粒间孔＋溶孔＋晶间孔最为发育,储集性能较好。     

Mg-lattice associations in red coralline algae

Recent investigations have shown red coralline algae to record ambient temperature in their calcite skeletons. Temperature recorded by variation in Mg concentrations within algal growth bands has sub-annual resolution and high accuracy. The conversion of Mg concentration to temperature is based on the assumption of Ca replacement by Mg within the algal calcite skeleton at higher temperatures. While Mg-temperature relationships in coralline algae have been calibrated for some species, the location of Mg within the calcite lattice remains unknown. Critically, if Mg is not a lattice component but associated with organic components this could lead to erroneous temperature records. Before coralline algae are used in large scale climate reconstructions it is therefore important to determine the location of Mg. Synchrotron Mg-X-ray absorbance near edge structure (XANES) indicates that Mg is associated with the calcite lattice in Lithothamnion glaciale (contemporary free-living, contemporary encrusting and sub-fossil free-living) and Phymatolithon calcareum (contemporary free-living) coralline algae. Mg is deposited within the calcite lattice in all seasons (L. glaciale & P. calcareum) and thallus areas (P. calcareum). These results suggest L. glaciale and P. calcareum are robust Mg-palaeotemperature proxies. We suggest that similar confirmation be obtained for Mg associations in other species of red coralline algae aiding our understanding of their role in climate reconstruction at large spatial scales.     

Coralline algae from the prang formation (middle-late Eocene) of the Lumshnong Area,Jaintia Hills,Meghalaya

The present paper records nine species of coralline algae from the Prang Formation of middle-late Eocene age from the Jaintia Hills, Meghalaya. The algae are associated with the larger foraminifera including Nummulites, Alveolina and Discocyclina throughout the succession. The temporal distribution of algal species indicates that seven species are known from the Eocene. Of the remaining two, one ranges from the Palaeocene to Eocene and the other is Palaeocene in age. The coralline algae and larger foraminifers are differentiated into two associations, the lower one indicating inner-ramp environment of high energy and the upper one showing deposition in relatively calm waters of deeper ramp environment.     

Holocene carbonate sedimentation on the west Eucla Shelf, Great Australian Bight: a shaved shelf

The southern continental margin of Australia is a cool-water carbonate sedimentary province located in a high-energy, swell-dominated oceanographic setting. A vibrocore transect of ¹⁴C-dated sediments across the centre of the Eucla Shelf is the first record of Holocene shelf deposition in the Great Australian Bight. Much of the seafloor shallower than 70 m water depth, the base of wave abrasion, is bare Cenozoic limestone, in some places encrusted by (?) Late Pleistocene, coral-rich, limestone that is cemented by high-magnesium calcite (12 mole% MgCO₃). The areally extensive, 100 km-wide, hard, bored substrate supports an epibiota of coralline algae, minor bryozoans and soft algae or is covered by patches of Holocene sediment up to 1.5 m thick; generally a basal bivalve lag (< 3 ka) overlain by quartzose-bioclastic palimpsest sand. This pattern of active carbonate production but little accretion on the wave-swept mid- to inner-shelf is similar to that on other parts of the southern Australian continental margin. The term shaved shelf is proposed for this style of carbonate platform, formed by alternating periods of sediment accretion, cementation and erosion.

The palimpsest sand is typically rich in bivalves, coralline algae and locally, detrital dolomite. Outer shelf Holocene sediment, below the base of wave abrasion but inboard of the shelf edge, is a metre-thick unit of fine, microbioclastic muddy sand with minor delicate bryozoans overlying a 9–13 ka rhodolith gravel. Some of this outer shelf sediment appears to have been resedimented. The shelf edge is a sandy and rocky seafloor with active bryozoan growth and sediment production.

The Holocene sediments are enriched in coralline algal particles and conspicuous large foraminifers (cf. Marginopora) and depleted in bryozoans, as compared to coeval deposits on the Lacepede and Otway shelves off southeastern Australia. These differences are interpreted to reflect warmer waters of the Leeuwin Current and prevalent downwelling in this area as opposed to the general upwelling and colder waters in the east.     

New record of coralline algae from the Holocene sediments of Agatti Island,Lakshadweep, India

The present paper examines the coralline algal assemblage recovered from the Holocene sediments of Agatti Island, Lakshadweep, India. The assemblage comprises nine species of coralline algae which include Amphiroa fragilisma, Lithophyllum nitorum, Lithophyllum incrustans, Lithoporella melobesioids, Spongites sp., Porolithon craspedium, Aethesolithon problematicum, Clathromorphum parcum, Melobesoideae gen. et spec. indet. Clathromorphum parcumis an endophytic coralline alga reported for the first time from the Indian subcontinent. Out of these, seven species belong to the family Corallinaceae and the two represent the family Hapalidiaceae of the class Rhodophyceae. Coralline algal association of this area is dominated by Lithophylloideae, Melobesioideae and Mastophoroideae which include encrusting to fragmented growth forms. The associated branching corals are well developed and adapted to shallow, warm water, low turbid, protected lagoonal environment.     

Reefal coralline algal build-ups within the Arctic Circle: morphology and sedimentary dynamics under extreme environmental seasonality

Carbonate frameworks secreted by phototrophic organisms within the Arctic Circle are not well documented. Underwater surveys of the inner-shelf off Troms, northern Norway (70°N), reveal extended fixed algal build-ups which are fringed by rhodolith belts affected by storms. Reefal growth by coralline algae under temperature and light regimes of extreme seasonality is made possible because of a decoupling of carbon fixation during summer and utilization of stored carbon during the period of winter darkness. Although the annual growth of the framework constructing algae is comparatively low, the annual carbonate production rate is similar to subtropical-tropical counterparts because of a remarkably high standing stock. Early diagenetic alteration is restricted to intraparticle cementation processes which start in vivo. Bioerosional destruction is the dominant control on the preservation of high latitude build-ups. Preservation of Holocene autochthonous coralline algal biostromes is enhanced by rapid burial during storm events. Redeposition during storms is the most important process in forming a distinct sedimentary facies zonation.     

Mg fractionation in crustose coralline algae: Geochemical, biological, and sedimentological implications of secular variation in the Mg/Ca ratio of seawater

The Mg/Ca ratio of seawater has varied significantly throughout the Phanerozoic Eon, primarily as a function of the rate of ocean crust production. Specimens of the crustose coralline alga Neogoniolithon sp. were grown in artificial seawaters encompassing the range of Mg/Ca ratios shown to have existed throughout the Phanerozoic. Significantly, the coralline algae’s skeletal Mg/Ca ratio varied in lockstep with the Mg/Ca ratio of the artificial seawater. Specimens grown in seawater treatments formulated with identical Mg/Ca ratios but differing absolute concentrations of Mg and Ca exhibited no significant differences in skeletal Mg/Ca ratios, thereby emphasizing the importance of the ambient Mg/Ca ratio, and not the absolute concentration of Mg, in determining the Mg/Ca ratio of coralline algal calcite. Specimens grown in seawater of the lowest molar Mg/Ca ratio (mMg/Ca = 1.0) actually changed their skeletal mineralogy from high-Mg (skeletal mMg/Ca > 0.04) to low-Mg calcite (skeletal mMg/Ca < 0.04), suggesting that ancient calcitic red algae, which exhibit morphologies and modes of calcification comparable to Neogoniolithon sp., would have produced low-Mg calcite from the middle Cambrian to middle Mississippian and during the middle to Late Cretaceous, when oceanic mMg/Ca approached unity. By influencing the original Mg content of carbonate facies in which these algae have been ubiquitous, this condition has significant implications for the geochemistry and diagenesis of algal limestones throughout most of the Phanerozoic. The crustose coralline algae’s precipitation of high-Mg calcite from seawater that favors the abiotic precipitation of aragonite indicates that these algae dictate the precipitation of the calcitic polymorph of CaCO₃. However, the algae’s nearly abiotic pattern of Mg fractionation in their skeletal calcite suggests that their biomineralogical control is limited to polymorph specification and is generally ineffectual in the regulation of skeletal Mg incorporation. Therefore, the Mg/Ca ratio of well-preserved fossils of crustose coralline algae, when corrected for the effect of seawater temperature, may be an archive of oceanic Mg/Ca throughout the Phanerozoic. Magnesium fractionation algorithms that model algal skeletal Mg/Ca as a function of seawater Mg/Ca and temperature are presented herein. The results of this study support the empirical fossil evidence that secular variation of oceanic Mg/Ca has caused the mineralogy and skeletal chemistry of many calcifying marine organisms to change significantly over geologic time.     

Microborer ichnocoenoses in Quaternary corals from New Caledonia: reconstructions of paleo-water depths and reef growth strategies in relation to environmental changes

Coral reef growth and development depend on several environmental factors, including tectonic and climatic parameters and local ecological drivers. Reef growth is especially sensitive to sea-level variations. Paleo-water depth reconstructions are essential tools used to determine reef growth patterns during different periods of reef growth. Assemblages of corals and/or coralline algae have been commonly used in such paleodepth reconstructions. This study shows that using microendolith ichnocoenoses can sometimes provide better accuracy than traditional coralgal analyses, particularly in the depth-range 0–10 m where coralgal assemblages usually show broad distribution ranges. Holocene and Pleistocene cores from two barrier reef sites on the west coast of Grande Terre in New Caledonia are examined here. Holocene reef development at these sites feature examples of microendolith ichnocoenoses that document rapid environmental changes and small sea-level variations of about 2–5 m in amplitude, and record these changes with more accuracy than coral and coralline algae assemblages which are highly dependant on the hydrodynamic energy of the setting. During the Pleistocene, which was less chronologically constrained, the microendolith ichnocoenoses also reflect paleo-water depths and reef-growth patterns at different periods of reef history.     

Facies analysis of Pleistocene limestones from Neil West Coast Formation,Neil Island,Ritchie’s archipelago of South Andaman,India

Petrographic thin section analysis of the samples collected from the type section of Neil West Coast Formation, situated in the west coast of Neil Island yielded moderately preserved coralline red algae, benthic and planktic foraminifers, coral fragments, echinoid spines and gastropod shells. The coralline red algae are represented by both non-geniculate and geniculate forms. The non-geniculate forms belong to melobesids, lithophylloids and mastophoroides. The geniculate forms are represented by species of Amphiroa, Corallina, and Jania. However, the diversity and abundance of coralline algal forms are less in comparison to the benthic foraminifers those are represented by Amphistegina, Neorotalia, Ammonia, Elphidium, Operculina, Assilina, Amphisorus and texularids. Planktic foraminifers like Globigerinoides and other biogenic components viz., gastropod shells, echinoid spines and coral fragments are also common. A foraminiferal-algal grainstone facies has been recognized as observed in the field as well as in thin section analysis. The overall assemblage of the biogenic components and facies analysis indicate intertidal to near shore environment of deposition with high energy condition and increased hydrodynamic activity.     

Sedimentary facies,production rates and facies models for recent coralline algal gravels,Co. Galway,Ireland

Five sedimentary facies are described from SCUBA diving examination and sampling of Mannin Bay, Ireland. A Bank facies is built up by the unattached coralline algae Lithothamnium corallioides and Phymatolithon calcareum. This autochthonous facies occurs in shallow sheltered environments. In exposed areas a rippled Clean Algal Gravel facies is found composed of coralline and molluscan debris. In intermediate energy areas a Muddy Algal Gravel facies is found with small amounts of live corallines. Sheltered creeks have a Mud facies which is partly carbonate and partly terrigenous. The shallow water coralline algal sediments are overlapped by a Fine Sand facies of mixed biogenic composition. Each facies is characterized by particular phenotypic growth forms of the unattached corallines. Rates of organic calcium carbonate production are obtained which are found to be similar to rates from shallow tropical non-reef environments. The carbonate sediments of Mannin Bay are compared with similar sediments from Kilkerrin Bay. Ireland, from Brittany and from Falmouth Harbour. From these comparisons, facies models are proposed for these carbonate sediments. The major factor controlling facies distribution is coastal morphology. The present day shelf is considered to be too exposed to preserve complete sequences of the shallow water sediments.     

Carbonate sediments of Elizabeth and Middleton Reefs close to the southern limits of reef growth in the southwest Pacific

Elizabeth and Middleton Reefs are atoll-like structures that have developed on top of volcanic edifices and are close to the southern environmental limit of reef development in the southwest Pacific. Reef morphology and vertical accretion rates during the Holocene appear similar to those on other more tropical reefs. Sediment samples were collected from the lagoon of both reefs and around the flanks of Middleton Reef. A distinctly chlorozoan assemblage was observed with coral, molluscs, Halimeda, coralline algae and foraminifers being the dominant sediment constituents. Lagoon sediment samples show little variation within or between reefs, lacking the concentric zonation characteristic of larger atolls. Samples collected from the flanks of Middleton Reef, and subsurface material from vibrocores, differ compositionally from the surficial lagoon sand and were typically more tropical in character. A comparison of the sediment constituents from these reefs with those of samples from within a fringing reef and from the shelf around Lord Howe Island, further south, indicated regional patterns in sediment composition. Halimeda rapidly decreased in abundance with increased latitude, and appeared confined to deeper water, whereas coralline red algae increased significantly. The rapid change in these major sediment contributors is coincident with the general decrease in coral growth rates with latitude. This reinforces the notion that the latitudinal limit of reef development is constrained by factors other than coral growth alone.     

Middle Eocene calcareous algae from Southwestern Kachchh,Gujarat

The Fulra limestone (middle Eocene) exposed in the areas around Jhadwa and Harudi villages, southwestern, Kachchh reveals presence of a rich assemblage of calcareous algae belonging to the Chlorophyceae and Rhodophyceae along with abundant foraminifera. In the present paper, eight species belonging to eight genera of calcareous algae are described. These include Dissocladella longijangensis, Sporolithon keenani, Corallina crossmanni, Arthrocardia sp. Misra et al. 2001, Lithothamnion ishigakiensis, Melobesioideae gen. et spec. indet. 1, Melobesioideae gen. et spec. indet. 2 and Lithoporella melobesioides. Out of these, one taxon belongs to the family Dasycladaceae. Among the remaining taxa, one taxon to the family Sporolithaceae, three taxa to the family Corallinaceae and three to the family Hapalidiaceae. Two coralline species, Corallina crossmanni and Lithothamnion ishigakiensis, are recorded for the first time from India. Another species (Dissocladella longijangensis), though known from other areas of India, is new to the study area.     

Palaeoecological implications of corallinacean red algae and halimedacean green algae from the prang formation of south shillong plateau, meghalaya

The southwestern part of south Shillong plateau (Meghalaya, N-E India), designated as Sylhet Limestone Group is sub-divided into three lithounits i.e., Lakadong, Umlatdoh and Prang formations in ascending order. The Prang Formation is the youngest lithostratigraphic unit of the Sylhet Limestone Group and has been dated as Middle to early Upper Eocene based on the benthic foraminifera studies. Thin section analysis of carbonate rocks from Prang Formation, exposed in the Bholaganj limestone quarry yielded a rich assemblage of calcareous algae. The coralline algal assemblage comprises both non-geniculate and geniculate forms. The green algae are represented by species of Halimeda belonging to the family Halimedaceae. Palaeoecological interpretation based on diversity, growth-form analysis and taphonomic aspects of the algal assemblage indicate that in all probabilities the deposition of Prang Formation occurred in shallow, warm, shelf environment of normal salinity within the transgressive phase.     

Towards a better understanding of magnesium-isotope ratios from marine skeletal carbonates

This study presents magnesium stable-isotope compositions of various biogenic carbonates of several marine calcifying organisms and an algae species, seawater samples collected from the western Dutch Wadden Sea, and reference materials. The aim of this study is to explore the influence of mineralogy, taxonomy and environmental factors (e.g., seawater isotopic composition, temperature, salinity) on magnesium-isotopic (δ²⁶Mg) ratios of skeletal carbonates. Using high-precision multi-collector inductively coupled plasma mass spectrometry, we observed that the magnesium-isotopic composition of seawater from the semi-enclosed Dutch Wadden Sea is identical to that of open marine seawater. We further found that a considerable component of the observed variability in δ²⁶Mg values of marine skeletal carbonates can be attributed to differences in mineralogy. Furthermore, magnesium-isotope fractionation is species-dependent, with all skeletal carbonates being isotopically lighter than seawater. While δ²⁶Mg values of skeletal aragonite and high-magnesium calcite of coralline red algae indicate the absence or negligibility of metabolic influences, the δ²⁶Mg values of echinoids, brachiopods and bivalves likely result from a taxon-specific level of control on Mg-isotope incorporation during biocalcification. Moreover, no resolvable salinity and temperature effect were observed for coralline red algae and echinoids. In contrast, Mg-isotope data of bivalves yield ambiguous results, which require further validation. The data presented here, point to a limited use of Mg isotopes as temperature proxy, but highlight the method’s potential as tracer of seawater chemistry through Earth’s history.