Beringia as an Ice Age genetic museum

Thousands of Late Pleistocene remains are found in sites throughout Beringia. These specimens comprise an Ice Age genetic museum, and the DNA contained within them provide a means to observe evolutionary processes within populations over geologically significant time scales. Phylogenetic analyses can identify the taxonomic positions of extinct species and provide estimates of speciation dates. Geographic and temporal divisions apparent in the genetic data can be related to ecological change, human impacts, and possible landscape mosaics in Beringia. The application of ancient DNA techniques to traditional paleontological studies provides a new perspective to long-standing questions regarding the paleoenvironment and diversity of Late Pleistocene Beringia.     

Origins and antiquity of the island fox (Urocyon littoralis) on California's Channel Islands

The island fox (Urocyon littoralis) is one of few reportedly endemic terrestrial mammals on California's Channel Islands. Questions remain about how and when foxes first colonized the islands, with researchers speculating on a natural, human-assisted, or combined dispersal during the late Pleistocene and/or Holocene. A natural dispersal of foxes to the northern Channel Islands has been supported by reports of a few fox bones from late Pleistocene paleontological localities. Direct AMS ¹⁴C dating of these “fossil” fox bones produced dates ranging from ∼ 6400 to 200 cal yr BP, however, postdating human colonization of the islands by several millennia. Although one of these specimens is the earliest securely dated fox from the islands, these new data support the hypothesis that Native Americans introduced foxes to all the Channel Islands in the early to middle Holocene. However, a natural dispersal for the original island colonization cannot be ruled out until further paleontological, archaeological, and genetic studies (especially aDNA [ancient DNA]) are conducted.     

青藏高原边缘地区晚更新世人类遗存与生存模式

自20世纪80年代在青海小柴旦发现旧石器遗存以来,青藏高原边缘地区已经发现了10余处旧石器时代遗址,年代属晚更新世晚期。这些遗存显示,古人类在距今30ka左右开始尝试向这些高海拔地区扩散,但直至距今15ka以后才有更多的人群迁徙至此。通过对这些遗存分布位置、古环境特征、年代测定和石器技术、组合分析,得以管窥古人类对该地区的开发利用过程,并进而阐释史前人类技术发展、生存模式演变和古环境的耦合关系。     

The Berelekh Mammoth “Graveyard”: New Chronological and Stratigraphical Data from the 2009 Field Season

To clarify unanswered questions of site formation, geology and the archaeology of the Berelekh geoarchaeological complex, a special survey was undertaken in 2009 of the area surrounding the site. Several geological units have been revealed. By establishing the spatial and temporal relationship of these deposits—as well as their age—we have reconstructed the formation history of the Berelekh bone bed. The mammoth bone deposit belongs to a paleochannel. Radiocarbon dating of mammoth remains at Berelekh demonstrates rapid accumulation during the Bølling warming. Human involvement in its formation is, at best, questionable, since there is no real overlap between the radiocarbon dates associated with past human activity, and those of the mammoth bone bed. This study confirms that humans used mammoth bone remains after the bone bed was deposited. Culturally, the Berelekh “site” does not have any relationship to the so‐called “Dyuktai culture.” Instead, the Berelekh archaeological finds (side notched stone pendants) show certain similarities to non‐microblade terminal Pleistocene assemblages found from Yenisei to Kamchatka. Additionally, the Berelekh complex presents a clear analogy with lithics found in Eastern Beringia. Teardrop‐shaped incomplete bifaces found in the assemblage are comparable to the Chindadn points of Alaska. The nature of this “Chindadn connection” is intriguing but it is the only visible cultural link between Western and Eastern Beringia.     

Late Pleistocene steppe lion Panthera leo spelaea (Goldfuss, 1810) footprints and bone records from open air sites in northern Germany – Evidence of hyena-lion antagonism and scavenging in Europe

Bone remains and a trackway of Pantheraichnus bottropensis nov. ichg. ichnsp. of the Late Pleistocene lion Panthera leo spelaea (Goldfuss, 1810) have been recovered from Bottrop and other open air sites in northern Germany. Some of these bones are from open air hyena den sites. A relative high proportion of lion bones (20%) exhibit bite, chew or nibble marks, or bone crushing and nibbling caused by a large carnivore. Repeated patterns of similar bone damage have been compared to bone remains found at hyena dens in gypsum karst areas and cave sites in northern Germany. Ice Age spotted hyenas have been the main antagonists and the main scavengers on lion carcasses. The remains appear to have been imported often by hyenas into their communal dens, supporting the theory of strong hyena-lion antagonism, similar to the well documented antagonism between modern African lions and spotted hyenas. Most of the lion bones from the open air hyena den at Bottrop are probably a result of such antagonism, as are the rare remains of these carnivores found within large hyena prey bone accumulations along the Pleistocene rivers. The Emscher River terrace also has the largest quantity of hyena remains from open air river terrace sites in northern Germany. Their cub remains, and incomplete chewed prey bones from mammoths and woolly rhinoceroses, typical of hyena activity, underline the character of these sites as cub-raising and communal dens, where their prey was accumulated along the riverbanks in a similar manner to modern African hyenas.     

Cave bears and paleolithic artifacts in Yarimburgaz Cave,Turkey: Dissecting a palimpsest

This article addresses the taphonomic issues surrounding artifact-bear associations in Yarimburgaz Cave, northwestern Turkey. Our purpose is to evaluate the circumstances of bone assemblage formation in the Middle Pleistocene deposits of the cave, identifying the agencies responsible for the accumulations and the causes of bone damage before and following deposition. Pleistocene species representation, bone damage, body part representation, and mortality data indicate that cave bear remains in Yarimburgaz are unrelated to hominid use of the same site. None of the observations on the cave bear remains contradicts the expectations developed from modern wildlife accounts of modern bear behavior; these accumulations resulted from mortality normally associated with hibernation over many generations of den use. Nonursid carnivores also played roles in bone collection and/or modification. We conclude that at least three distinct biological processes contributed to the formation of the Middle Pleistocene faunas: hibernating bears, bone-collecting carnivores such as wolves, and hominids in descending order of importance. The Yarimburgaz faunas represent palimpsests or overlays of many short-term depositional events, the close spatial associations of which are explained by slow or uneven sedimentation rates inside the cave. Although occupations by hominids appear to have been ephemeral in nature, hominids ultimately discarded nearly 1700 stone artifacts in Yarimburgaz Cave. The hominids did not discard many ungulate bones in the same circumstances. These observations suggest that hominid foraging efforts focused on resources other than large game while at the cave. © 1996 John Wiley & Sons, Inc.     

Late Pleistocene paleoecology of arctic ground squirrel (Urocitellus parryii) caches and nests from Interior Alaska's mammoth steppe ecosystem, USA

Botanical analyses of fossil and modern arctic ground squirrel (Urocitellus parryii) caches and nests have been used to reconstruct the past vegetation from some parts of Beringia, but such archives are understudied in Alaska. Five modern and four fossil samples from arctic ground squirrel caches and nests provide information on late Pleistocene vegetation in Eastern Beringia. Modern arctic ground squirrel caches from Alaska's arctic tundra were dominated by willow and grass leaves and grass seeds and bearberries, which were widespread in the local vegetation as confirmed by vegetation surveys. Late Pleistocene caches from Interior Alaska were primarily composed of steppe and dry tundra graminoid and herb seeds. Graminoid cuticle analysis of fossil leaves identified Calamagrostis canadensis, Koeleria sp. and Carex albonigra as being common in the fossil samples. Stable carbon isotopes analysis of these graminoid specimens indicated that plants using the C₃ photosynthetic pathways were present and functioning with medium to high water-use efficiency. Fossil plant taxa and environments from ground squirrel caches in Alaska are similar to other macrofossil assemblages from the Yukon Territory, which supports the existence of a widespread mammoth steppe ecosystem type in Eastern Beringia that persisted throughout much of the late Pleistocene.     

Paleolithic burnt bone horizons from the Swabian Jura: Distinguishing between in situ fireplaces and dumping areas

Hohle Fels Cave is one of several Upper Paleolithic sites on the eastern extension of the Swabian Alb in southwestern Germany. Several phases of excavations have been conducted since 1870. The archaeological inventories comprise a broad range of artifacts produced from stone, bone, teeth, ivory, and antler. The site has also yielded Paleolithic rock art, sculpture, and engravings. The Pleistocene inventories unearthed so far belong to the Magdalenian, Gravettian, and Aurignacian; Middle Paleolithic horizons may well be uncovered as excavations continue. The chronology of three main occupation phases has been established with numerous ¹⁴C measurements. The Magdalenian dates to about 13,000 yr B.P., and the richest Gravettian deposit contains a burnt bone‐layer extending over ca. 12 m² that dates to about 29,000 yr B.P. The underlying Aurignacian horizons predate 30,000 yr B.P. The Gravettian burnt bone layer (archaeological layer IIcf) is a key horizon because of its distinct composition and color and its large horizontal extension. Layer IIcf has been examined using a combination of analytical techniques including micromorphological analysis of thin sections with polarizing light microscopy and scanning electron microscopy for fabric and identification of mineral and biogenic constituents. The bone mineral composition was analyzed by Fourier transform infrared spectroscopy (FTIR) and electron probe microanalytical techniques. These investigations in tandem with the field observations indicate intense burning of bones as fuel. The micromorphological and contextual archaeological data suggest that layer IIcf is a secondary dump rather than an in situ accumulation. This deposit and other Paleolithic burnt deposits in the Swabian Jura appear to reflect intense periods of winter occupation in these caves. © 2003 Wiley Periodicals, Inc.     

Context and environment in taphonomic analysis: Examples from Alaska's Porcupine River Caves

Field investigations of caves along Alaska's Porcupine River document three major mechanisms which modify bone in patterns similar to alterations produced by man: (1) carnivore fracture; (2) rodent gnawing; and (3) rock fall and rubble scarring. A late Wisconsin faunal assemblage composed of Equus sp., Rangifer tarandus, Ovis dalli, Bison sp., proboscidean, numerous small mammal species, birds, and fish is well documented. This faunal assemblage suggests a mosaic environment of grassland-tundra-forest in the immediate vicinity of these caves and implies that the late Wisconsin environment in north-central Alaska may have been characterized by a number of microenvironments and colder, dryer, steppe conditions. Taphonomic data which have historically been interpreted to support human occupation of eastern Beringia during the Pleistocene are critically examined and the context of these discoveries (not the specimens themselves) provides the test essential to document the antiquity of man in North America prior to 12,000 yr ago.     

Toward the definition of criteria for the recognition of artificial bone alterations

Cutting, fracturing, flaking, and polishing of bones and other osseous materials may in some instances be interpreted as evidence of former human activity. Such interpretations must avoid confusion with the wide variety of natural processes that alter bones. Reliable criteria are prerequisite to sound inferences based on bone, regardless of whether they have been redeposited or recovered from primary assemblages. Criteria must be defined by means of actualistic studies (neotaphonomy) and experiments that demonstrate causal relationships between patterns of alteration and the processes that produce them. The criteria can be empoyed in the interpretation of fossil bones (paleotaphonomy) on the basis of uniformitarian principles. At the present time, such relationships are relatively well understood in the case of cut marks on bones, and modest progress has been made in investigating fractured and flaked bones. Polished bones are little understood and difficult to interpret.     

Electron spin resonance dating of Quaternary bone material from Tasmanian caves—A comparison with ages determined by aspartic acid racemization and C14

Abstract

Fourteen bone samples are analysed to test the usefulness of equivalent dose (ED) determinations by electron spin resonance (ESR) as a rapid method of determining relative age and making an estimate of absolute age. ED values are compared with eight aspartic acid dates and two C¹⁴ dates. The latter are dates on charcoal found in close association with bone at archaeological sites. For samples less than 25 000 years old an excellent correlation is obtained when ED values are compared with dates obtained by the other two methods. The relationship suggests that ED values can be converted to estimates of bone age by assuming a mean annual dose rate of 0.1 rad/yr. Age determinations provide little evidence to support earlier suggestions that elements of the Late Pleistocene megafauna survived until the end of the Pleistocene. Bone material at some sites in the‐Florentine Valley and near Montagu appears to be much older than had previously been believed. Only one site (Main Cave, Montagu) containing megafaunal elements appears to be terminal Pleistocene in age but the possibility of reworking of megafauna material from nearby older sites cannot be excluded. ESR dating has considerable potential as an exploratory dating tool but can only be applied to dense, unaltered bone samples. Attempts to analyse five samples from Kutikina Cave in Western Tasmania were unsuccessful because of post‐depositional contamination of the bone.     

The U.P. Mammoth Site,Carbon County,Wyoming, USA: More Questions than Answers

In the summer of 1960, mammoth bones were discovered by a dragline operator in southern Wyoming at the Union Pacific (U.P.) Mammoth site. Although subsequent archaeological work during 1960 and 1961 identified artifacts in association with the mammoth remains, many authors have since questioned the nature of that association. Also, little has been published about the site other than a brief article in National Geographic Magazine in 1962. In this paper, we present additional information on the U.P. Mammoth site including stratigraphic profiles from the first author's geoarchaeological work in 1961, stratigraphic and spatial location of bones and artifacts derived from the original field notes, and new radiocarbon dates. Although the precise stratigraphic provenance for many artifacts and skeletal elements remains unclear, a compelling argument can be made for spatial and stratigraphic association of the mammoth remains with the artifact assemblage suggesting some kind of human interaction with the animal.     

Amino acid racemization in mammalian bones and teeth from La Chaise-de-Vouthon (Charente), France

A controlled test of the amino acid dating method as it is currently being applied to bones and teeth was attempted for the time range 100-250 ka, beyond that of ¹⁴C, at the archeological site of La Chaise-de-Vouthon (Charente, France). In Bourgeois-Delaunay, mammal fossils associated with Paleolithic artifacts were collected from layers dated at 101 ± 12 to 114 ± 7 ka by ²³⁰Th/²³⁰ dating of the over- and underlying stalagmitic floors. Racemization ratios for most amino acids were significantly lower than for comparably aged materials from other European sites. Regardless of the rate constants used, most amino acid dates were incorrect. Large associated errors spanned the late Pleistocene, making the dates useless for discriminating archaeological or geological events. Stratigraphic correlations using any tissue were also problematic. Relative interacid ratios varied with age, indicating that diagenetic alteration had affected racemization. Therefore, amino acid relative and absolute dating cannot be accurately applied to mid-Pleistocene bones or teeth.     

Geological perspectives on the Monte Verde archeological site in Chile and pre-Clovis coastal migration in the Americas

Discovery of the Monte Verde archeological site in Chile overturned the previous consensus that the first Americans into the New World from Asia were the makers of Clovis projectile points, and rejuvenated the hypothesis that migration through the Americas occurred largely on portions of the Pacific continental shelf exposed by Pleistocene drawdown in eustatic sea level. The postulate of travel along a paleoshoreline now hidden underwater is an attractive means to posit pre-Clovis human movement southward from Beringia to Chile without leaving traces of migration onshore. Geologic analyses of the Pleistocene paleoenvironment at Monte Verde and of the morphology of the potential migration route along the continental shelf raise questions that have not been fully addressed. The periglacial setting of Monte Verde may call its antiquity into question and the narrowness of the Pacific continental shelf of the Americas makes it unlikely that people could travel the length of the Americas without impacting ground still onshore and no farther inland than Monte Verde itself. Geological perspectives on Monte Verde and coastal migration jointly suggest that the Clovis-first hypothesis for peopling the New World may have been abandoned prematurely.     

Moraine chronosequence of the Donnelly Dome region, Alaska

We present ¹⁰Be exposure ages from moraines in the Delta River Valley, a reference locality for Pleistocene glaciation in the northern Alaska Range. The ages are from material deposited during the Delta and Donnelly glaciations, which have been correlated with MIS 6 and 2, respectively. ¹⁰Be chronology indicates that at least part of the Delta moraine stabilized during MIS 4/3, and that the Donnelly moraine stabilized ∼ 17 ka. These ages correlate with other dates from the Alaska Range and other regions in Alaska, suggesting synchronicity across Beringia during pulses of late Pleistocene glaciation. Several sample types were collected: boulders, single clasts, and gravel samples (amalgamated small clasts) from around boulders as well as from surfaces devoid of boulders. Comparing ¹⁰Be ages of these sample types reveals the influence of pre/post-depositional processes, including boulder erosion, boulder exhumation, and moraine surface lowering. These processes occur continuously but seem to accelerate during and immediately after successive glacial episodes. The result is a multi-peak age distribution indicating that once a moraine persists through subsequent glaciations the chronological significance of cosmogenic ages derived from samples collected on that moraine diminishes significantly. The absence of Holocene ages implies relatively minor exhumation and/or weathering since 12 ka.     

A groundwater hypothesis for the origin of “fire areas” on the Northern Channel Islands,California

Pleistocene areas of red sediments and carbonized vegetation on the Northern Channel Islands, California, have in the past been interpreted as caused by fires of either natural or human origin. Some are associated with darkened mammoth and bird fossils, and these fossils have been considered as having been burned by early man. Reevaluation of these so-called “fire areas” indicates that the above phenomena are the result of low-temperature (≤100°C), nonheating processes occurring in groundwater. Evidence for this conclusion is derived from field observations on fossil carbonized vegetation, and the geology of the areas. Additional evidence derives from experiments on the red sediments and fossil wood, X-ray diffraction analyses, magnetic analyses, studies on the clay minerals smectite and illite, and the demonstration that fossil mammoth bone contains sufficient Fe and Mn to account for their discoloration. Much of the carbonization of vegetation probably occurred in water rather than in fire. Radiocarbon dates from the islands will probably need to be reevaluated. These data provide evidence contrary to the concept of the occurrence of significant fires, either natural or set by early man, on the Northern Channel Islands.     

内蒙古鄂尔多斯乌兰木伦遗址 2010年1期试掘及其意义

乌兰木伦遗址于2010年发现并进行了首期试掘,在7个文化层位中获得石制品1500余件、动物化石千余件以及人类用火遗迹。初步观察和研究表明,该遗址以石片石器为特点的石制品类型十分丰富,工具组合具有明显的旧石器中期的特点,且与欧洲的莫斯特文化存在不少相似之处; 以披毛犀为代表的哺乳动物化石属于华北晚更新世萨拉乌苏动物群; 沉积相和哺乳动物、软体动物化石的性质表明当时人类生活于较为温暖、湿润的气候环境中; 化石总体比较破碎,一些碎骨化石上可见明显的石器切割痕迹,同时发现有大量骨片,带剥片痕的骨核化石和若干骨器标本以及烧骨,反映出当时人类的肉食、熟食习惯等利用动物骨骼的生活行为方式。经 14 C与光释光结合测定乌兰木伦遗址年代为距今70~30ka,属于旧石器时代中期。乌兰木伦遗址的发现、试掘与初步研究,为我国北方旧石器中期文化的研究增添了新的内涵,同时也为现代人起源和中西方文化交流研究提供了新的线索。     

Geology of the dry creek site; a stratified early man site in Interior Alaska

The Dry Creek archeologic site contains a stratified record of late Pleistocene human occupation in central Alaska. Four archeologic components occur within a sequence of multiple loess and sand layers which together form a 2-m cap above weathered glacial outwash. The two oldest components appear to be of late Pleistocene age and occur with the bones of extinct game animals. Geologic mapping, stratigraphic correlations, radiocarbon dating, and sediment analyses indicate that the basal loess units formed part of a widespread blanket that was associated with an arctic steppe environment and with stream aggradation during waning phases of the last major glaciation of the Alaska Range. These basal loess beds contain artifacts for which radiocarbon dates and typologic correlations suggest a time range of perhaps 12,000–9000 yr ago. A long subsequent episode of cultural sterility was associated with waning loess deposition and development of a cryoturbated tundra soil above shallow permafrost. Sand deposition from local source areas predominated during the middle and late Holocene, and buried Subarctic Brown Soils indicate that a forest fringe developed on bluff-edge sand sheets along Dry Creek. The youngest archeologic component, which is associated with the deepest forest soil, indicates intermittent human occupation of the site between about 4700 and 3400 ¹⁴C yr BP.     

旧石器时代古人类敲骨取髓行为的确认——以马鞍山遗址为例

敲骨取髓是旧石器时代古人类获取脂肪的必要途径和手段。对于这一行为的辨识不能仅根据某一孤立的特征判定,而要综合长骨的断裂面形态、骨骼表面与断裂面之间的夹角、断裂面质地、管状骨横剖面的周长以及敲砸痕与敲砸疤在长骨表面的分布等一系列特征组合来确定。本文结合西方动物考古及埋藏学的实证研究（包括实验和民族学观察）方法与数据,对比分析了马鞍山遗址动物骨骼组合中长骨碎片的破裂性质及敲砸疤的分布状况,确认该区域古人类敲骨取髓行为的存在,并初步探讨了他们对长骨骨髓的利用方式与程度。     

On the possible utilization of Camelops by early man in North America

Camelops was a major faunal element in late Wisconsin biotic communities over much of North America. Interpretations of possible human association with Camelops are often based on poorly evaluated evidence. Ideal standards for acceptable evidence are compared here to the actual evidence that has been advanced. Of 25 fossil assemblages examined, 2 might be examples only of geological contemporaneity of humans and Camelops; 2 might indicate behavioral association of humans and Camelops bones; and 2 might indicate actual human utilization of Camelops (killing and/or butchering). Camelops bones interpreted as artifacts are similar to modern specimens affected by noncultural processes.