山地垂直带谱信息系统应用分析与技术改进

山地垂直带信息系统虽实现了山地垂直带谱的首次数字集成,但系统中还存在标识不清、垂直带颜色不易变动、大量带谱显示时可视程度较差等问题。本文以山地垂直带谱空间可视化为研究对象,探讨系统的技术改进,包括山地垂直带谱图形数据的立体标识、翻转页面、颜色可定制、数据轴可操作等功能的实现。山地垂直带信息系统空间可视化功能的进一步完善,不仅可以有效地管理山地垂直带谱数据,而且可方便直观地显示山地垂直带谱数据,有效地集成山地垂直带谱数据并且充分地发掘山地垂直带谱空间变化规律,也为建立界面友好的世界山地垂直带信息系统奠定了基础。     

中国山地垂直带信息系统的设计与开发

本文收集了大量翔实的山地垂直带资料,建立了我国山地垂直带数据库,创建了山地垂直带谱"数字引擎".使垂直带谱数字化成为可能。采用ArcObject组件开发技术及Visual Basic,自主开发中国山地垂直带信息系统(1.0)。该系统是一个面向对象(山地垂直带谱)的地理信息系统;以山地垂直带"数字引擎"为系统内核,在功能上实现了山地垂直带分布与地理空间区域的联接,同时实现了大范围内选择多个区域进行数据分析的功能。系统界面友好,易操作,显示直观、明了,为自然地理和生态学家者提供了一个比较丰富的山地垂直带谱数据集和有效的分析工具。标志着山地垂直带谱研究进入了数字时代。     

Digital extraction of altitudinal belt spectra in the West Kunlun Mountains using SPOT-VGT NDVI and SRTM DEM

In this paper, a digital identification method for the extraction of altitudinal belt spectra of montane natural belts is presented. Acquiring the sequential spectra of digital altitudinal belts in mountains at an acceptable temporal frequency and over a large area requires extensive time and work if traditional methods of field investigation are to be used. Such being the case, often the altitudinal belts of a whole mountain or the belts at a regional scale are represented by single points. However, single points obviously cannot accurately reflect the spatial variety of altitudinal belts. In this context, a digital method was developed to extract the spectra of altitudinal belts from remote sensing data and SRTM DEM in the West Kunlun Mountains. By means of the 1km resolution SPOT-4 vegetation 10-day composite NDVI, the horizontal distribution of altitudinal belts were extracted through supervised classification, with a total classification accuracy of 72.23%. Then, a way of twice-scan was used to realize the automatic transition of horizontal maps to vertical belts. The classification results of remote-sensing data could thus be transformed automatically to sequential spectra of digital altitudinal belts. The upper and lower lines of the altitudinal belts were then extracted by vertical scanning of the belts. Relationships between the altitudinal belts based on the montane natural zones concerning vegetation types and the geomorphological altitudinal belts were also discussed. As a tentative method, the digital extraction method presented here is effective at digitally identifying altitudinal belts, and could be helpful in rapid information extraction over large-scale areas.     

基于垂直带谱的太白山区山地植被遥感信息提取

遥感数据因其全覆盖的优势被广泛应用于山地植被信息的调查和研究。为了实现山区植被类型的高精度提取,本文以太白山区为实验区,结合山地植被的垂直地带性分布规律,利用太白山植被垂直带谱、高分辨率遥感影像（GF1/GF2/ZY3）和1:1万的数字表面模型（Digital Surface Model, DSM）数据,进行了多层次、多尺度的影像分割,构建了具有植被垂直带谱信息的地形约束因子,并据此进行样本选择和面向对象的分类,分类总精度达92.9%,kappa系数达到0.9160。该方法相比于未辅以垂直带谱信息的分类,总精度提高了10%。研究结果表明,分类过程中加入具有垂直带谱信息的地形约束因子,能显著地提高样本选择的效率和准确率,为后续的植被分类提供了精度的保证。通过人机交互的方式,将垂直带谱知识应用到分类中,可以有效地提高山地植被分类的精度。     

Digital Spectra and Analysis of Altitudinal Belts in Tianshan Mountains, China

Based on the framework of the geo-info spectra of montane altitudinal belts, this paper firstly reviews six classification systems for the spectra of mountain altitudinal belts in China and considers that detailed regional study of altitudinal belts is the key for reaching standardization and systemization of mountain altitudinal belts. Only can this further identify and resolve problems with the study of altitudinal belts. The factors forming the spectra of altitudinal belts are analyzed in the Tianshan Mountains of China, and a digital altitudinal belt system is constructed for the northern flank, southern flank, the heartland, and Ili valley in the west. The characteristics of each belt are revealed with a summarization of the pattern of areal differentiation of altitudinal belts.     

Regional-scale identification of three-dimensional pattern of vegetation landscapes

The altitudinal pattern of vegetation is usually identified by field surveys,however,these can only provide discrete data on a local mountain.Few studies identifying and analyzing the altitudinal vegetation pattern on a regional scale are available.This study selected central Inner Mongolia as the study area,presented a method for extracting vegetation patterns in altitudinal and horizontal directions.The data included a vegetation map at a 1∶1 000 000 scale and a digital elevation model at a 1∶250 000 scale.The three-dimensional vegetation pattern indicated the distribution probability for each vegetation type and the transition zones between different vegetation landscapes.From low to high elevations,there were five vegetation types in the southern mountain flanks,including the montane steppe,broad-leaved forest,coniferous mixed forest,montane dwarf-scrub and sub-alpine shrub-meadow.Correspondingly,only four vegetation types were found in the northern flanks,except for the montane steppe.This study could provide a general model for understanding the complexity and diversity of mountain environment and landscape.     

山地垂直带谱数字识别的技术实现和图谱构建

山地垂直带谱是地学信息图谱的一个重要组成部分。"带谱数字识别"就是利用现代数字方法和数据,对客观存在的山地垂直自然带谱进行提取,获取比较完备的山地垂直带连续图谱模式。这是从传统山地垂直带研究走向地学信息图谱研究的重要步骤,有助于使我们对山地垂直谱的认识上升到地学信息图谱的高度。本文探索MATLAB语言快速、准确地实现各种识别算法,结合VB.NET构建的用户操作界面,实现带谱的信息提取、分类集成和多样化表达。山地垂直自然带谱的数字识别分为3种不同的模式:山系单侧、山体单峰和山体多峰;依据各自不同的识别算法,可以得到山地连续的带谱模式。本文全面介绍了数字带谱、带谱数字识别的含义,以及数字识别模型的原理、算法和原型系统等实现过程。     

Comparison of the spatio-temporal dynamics of vegetation between the Changbai Mountains of eastern Eurasia and the Appalachian Mountains of eastern North America

The Changbai Mountains and the Appalachian Mountains have similar spatial contexts. The elevation, latitude, and moisture gradients of both mountain ranges offer regional insight for investigating the vegetation dynamics in eastern Eurasia and eastern North America. We determined and compared the spatial patterns and temporal trends in the normalized difference vegetation index (NDVI) in the Changbai Mountains and the Appalachian Mountains using time series data from the Global Inventory Modeling and Mapping Studies 3rd generation dataset from 1982 to 2013. The spatial pattern of NDVI in the Changbai Mountains exhibited fragmentation, whereas NDVI in the Appalachian Mountains decreased from south to north. The vegetation dynamics in the Changbai Mountains had an insignificant trend at the regional scale, whereas the dynamics in the Appalachian Mountains had a significant increasing trend. NDVI increased in 55% of the area of the Changbai Mountains and in 95% of the area of the Appalachian Mountains. The peak NDVI occurred one month later in the Changbai Mountains than in the Appalachian Mountains. The results revealed a significant increase in NDVI in autumn in both mountain ranges. The climatic trend in the Changbai Mountains included warming and decreased precipitation, and whereas that in the Appalachian Mountains included significant warming and increased precipitation. Positive and negative correlations existed between NDVI and temperature and precipitation, respectively, in both mountain ranges. Particularly, the spring temperature and NDVI exhibited a significant positive correlation in both mountain ranges. The results of this study suggest that human actives caused the differences in the spatial patterns of NDVI and that various characteristics of climate change and intensity of human actives dominated the differences in the NDVI trends between the Changbai Mountains and the Appalachian Mountains. Additionally, the vegetation dynamics of both mountain ranges were not identical to those in previous broader-scale studies.     

The potential contribution of wildlife sanctuary to forest conservation: A case study from binsar wildlife sanctuary

Forest vegetation of a protected area(Binsar Wildlife Sanctuary) in Kumaun region(west Himalaya) was analysed for structure,composition and representativeness across three different altitudinal belts,lower(1,600-1,800 m a.s.l.),middle(1,900-2,100 m a.s.l.) and upper(2,200-2,400 m a.s.l.) during 2009-2011 using standard phytosociological methods.Four aspects(east,west,north and south) in each altitudinal belt were chosen for sampling to depict maximum representation of vegetation in the sanctuary.Population structure and regeneration behaviour was analysed seasonally for two years to show the establishment and growth of tree species.A total of 147 plant species were recorded from the entire region of which 27 tree species were selected for detailed study.Highest number was recorded at upper(18 species),and lowest at lower altitudinal belt(15 species).The relative proportion of species richness showed higher contribution of tree layer at each altitudinal belt.The population structure,based on the number of individuals,revealed a greater proportion of seedling layer at each altitudinal belt.The relative proportion of seedlings increases significantly along altitudinal belts(p<0.05) while opposite trends were observed in sapling and tree layers.The density of sapling and seedling species varied non-significantly across seasons(p>0.05).The density values decreased in summer and increased during rainy season.As far as the regeneration status is concerned,middle and upper altitudinal belts showed maximum number of species with fair regeneration as compared to lower altitudinal belt.Overall density diameter distribution of tree species showed highest species density and richness in the smallest girth class and decreased in the succeeding girth classes.This study suggests that patterns of regeneration behaviour would determine future structural and compositional changes in the forest communities.It is suggested that the compositional changes vis-à-vis role of ‘New’ and ‘Not regenerating’ species need priority attention while initiating conservation activities in the sanctuary.This study calls for exploring other less explored Wildlife Sanctuaries in the Himalaya and across the world,to achieve overall biodiversity status in these protected areas and thus to justify their role in conserving biodiversity in the region.     

Woody species diversity and forest structure from lowland to montane forest in Hyrcanian forest ecoregion

Alborz Mountains host Caspian Hyrcanian forest ecoregion along the northern slopes and forest steppe ecoregion in highlands. Hyrcanian forest covers the southeastern part of Caucasus biodiversity hotspot and is of great biogeographic importance. Altitudinal pattern and correlation between woody species biodiversity (DIV), forest structure ((stem density (DEN), mean basal area (MBA) and mean height class (MHC)) and disturbance (DIS) were explored along 2,400 m altitudinal gradient in Hyrcanian relict forest, Central Alborz Mountains. Vegetation changes from lowland forest (LoF) to mid- altitude forest (MiF) and montane forest (MoF) in this area. The altitudinal gradient was divided into twelve 200 m elevational belts. Point centered quarter method (PCQM) with 96 sampling points and 83 vegetation samples by plot method (PM) were used to record field data. Shannon-Wiener index and Pearson coefficient were used for diversity and correlation analysis. The results showed that DEN decreased linearly, MBA and MHC showed relatively hump shaped and DIS showed a reverse hump shaped pattern of change along altitudinal gradient. Woody species diversity decreased non-steadily from LoF to MoF. Transitional vegetations of Carpinus-Fagus and Fagus-Quercus represented higher diversity of woody taxa compared to adjacent homogenous communities. Significant correlation was observed between altitude and all parameters: DEN with MBA, DIS and DIV; MBA with DIS; MHC with DIS along with DIV; and DIS with DIV at the study area scale. Surprisingly,correlation between studied parameters differed within each vegetation type. Altitude probably acts as a proxy for human and environmental driving forces in this area. Stability of warm and wet condition, season length, soil depth along with forest accessibility probably influences the altitudinal pattern of the studied parameters. Disturbance affects forest structure and consequently diversity; especially in lowlands. The obtained results recommend using both forest biodiversity and mensuration data in management process of forest ecosystems.     

秦巴山地山体基面高度的提取及分布

秦巴山地是中国的南北分界线,也是黄河和长江的分水岭,其山体效应的定量化影响秦巴山地山体垂直带的分布格局、非地带性因素的作用强度和机理,以及中国暖温带和北亚热带的具体位置的确定。山体基面高度是影响山体效应最重要和关键的地形因子,其定量化和数字化提取是秦巴山地山体效应定量化研究的重要内容。本研究针对秦巴山地山体效应的定量化研究,使用30 m分辨率的STRM-1数据,分别基于山体特征线和流域分区2种方法提取了秦巴山地的山体基面高度分区,并根据地形起伏度和坡度,确定基面范围,计算了山体基面高度值。结果表明：① 基于山体特征线的方法将秦巴山地分为93个基面高度分区,基于流域分区的方法将秦巴山地分为209个基面高度分区,根据2种分区结果提取的基面高度值相差不大且均体现了秦巴山地地势的特点;② 秦巴山地山体基面高度从东向西呈阶梯状递增的趋势;③ 从南到北,秦巴山地的东段和中段均呈先增高后降低的趋势,即从大巴山向北至汉江谷地降低,再向北至秦岭升高;④ 山地的不同侧翼的山体基面高度不同,秦岭南坡的基面高度（1000~1809 m）明显高于北坡（850~1300 m）。秦巴山地山体基面高度与其植被带分布上限联系密切,实现山体基面高度的数字化提取,为山体效应的定量化研究提供了重要的技术支持。     

欧亚大陆山地垂直带数字集成系统的设计与应用

在欧亚大陆山地垂直带880个带谱点数据采集的基础上,设计和开发了山地垂直带数字集成系统。其功能：(1)垂直带谱可视化显示功能：实时生成垂直带地理分布图、垂直带堆积柱状图、上下限高度随经纬度变化曲线;(2)垂直带上下限提取和绝对高度和相对高度转化功能：提取山地垂直自然带分布上限和下限高度,进行垂直带的宽度和垂直带上限海拔...     

Omni-directional distribution patterns of montane coniferous forest in the Helan Mountains of China

Slope aspect has significant effect on the development and distribution of montane forest,especially in arid and semiarid regions.This paper,using SPOT5 images and 1:50,000 DEM,digitally extracts and analyzes the spatial information of montane coniferous forest(mainly Qinghai spruce),and thereby explores how the upper and lower limits and the altitudinal range of coniferous forest vary and how the area of coniferous forest is related with annual insolation with all aspects in the Helan Mountains.The results show that: 1) In the eastern flank,the lower limit of coniferous forest is between 1,600 m and 2,000 m a.s.l.,and the upper limit between 2,800 m and 3,000 m a.s.l.;in the western flank,the lower limit of coniferous forest is between 2,000 m and 2,300 m a.s.l.,and the upper limit between 2,800 m and 3,100 m a.s.l.2) The altitudinal ranges of coniferous forest are 806~1,435 m,438~1,140 m for eastern flank and western flank,respectively.3) The area of coniferous forest takes on a normal distribution with aspect,and it has a close relationship with annual insolation.This distribution model developed in this paper quantitatively reveals the significant effect of slope aspect on the distribution of coniferous forest in arid and semi-arid land.     

Tree population dynamics of three altitudinal vegetation communities on Mount Cameroon (1989�C2004)

Changes in permanent sample plots in the lowland,submontane and montane forests on Mount Cameroon(4,095 m above sea level),an active volcano,are described for 15 years from 1989 to 2004.Throughout the study period,the stocking level of trees with a diameter at breast height(DBH) ≥ 10 cm in the three forests were lower than in pan-tropical stands suggesting a significant impact of volcanic and human-related activities on the vegetation communities on the mountain.Annual mortality rates in the submontane and montane forests were consistent with those reported for comparable altitudinal ranges in the Blue Mountains of Jamaica.The annual mortality rate was higher in the lowland forest than other lowland sites included.Divergence between recruitment and mortality rates was large suggesting that the three vegetation communities have not reached their climax.The seven-year difference in half-life of large trees(with a DBH ≥ 50 cm) in the submontane and montane forests suggests an altitudinal effect on turnover of larger trees that in turn contributes to the frequent small stature of high altitude forests.There was little evidence of an altitudinal effect on species turnover and growth rate.This finding supports generalizations about the zero effect of growth on the stature of high altitude trees.Understanding forest dynamics is crucially important in the management of tropical montane environmentsand in this instance particularly so given the recent creation of the Mount Cameroon National Park.     

Variation of Thornthwaite moisture index in Hengduan Mountains,China

The Thornthwaite moisture index, an index of the supply of water(precipitation) in an area relative to the climatic demand for water(potential evapotranspiration), was used to examine the spatial and temporal variation of drought and to verify the influence of environmental factors on the drought in the Hengduan Mountains, China. Results indicate that the Thornthwaite moisture index in the Hengduan Mountains had been increasing since 1960 with a rate of 0.1938/yr. Annual Thornthwaite moisture index in Hengduan Mountains was between –97.47 and 67.43 and the spatial heterogeneity was obvious in different seasons. Thornthwaite moisture index was high in the north and low in the south, and the monsoon rainfall had a significant impact on its spatial distribution. The tendency rate of Thornthwaite moisture index variation varied in different seasons, and the increasing trends in spring were greater than that in summer and autumn. However, the Thornthwaite moisture index decreased in winter. Thornthwaite moisture index increased greatly in the north and there was a small growth in the south of Hengduan Mountains. The increase of precipitation and decrease of evaporation lead to the increase of Thornthwaite moisture index. Thornthwaite moisture index has strong correlation with vegetation coverage. It can be seen that the correlation between Normalized Difference Vegetation Index(NDVI) and Thornthwaite moisture index was positive in spring and summer, but negative in autumn and winter. Correlation between Thornthwaite moisture index and relative soil relative moisture content was positive in spring, summer and autumn, but negative in winter. The typical mountainous terrain affect the distribution of temperature, precipitation, wind speed and other meteorological factors in this region, and then affect the spatial distribution of Thornthwaite moisture index. The unique ridge-gorge terrain caused the continuity of water-heat distribution from the north to south, and the water-heat was stronger than that from the east to west part, and thus determined the spatial distribution of Thornthwaite moisture index. The drought in the Hengduan Mountains area is mainly due to the unstable South Asian monsoon rainfall time.     

Spatio-temporal variation of Fraction of Photosynthetically Active Radiation absorbed by vegetation in the Hengduan Mountains,China

The Fraction of Absorbed Photosynthetically Active Radiation(FPAR) is an important indicator of the primary productivity of vegetation. FPAR is often used to estimate the assimilation of carbon dioxide in vegetation. Based on MOD15 A2 H/FPAR data product, the temporal and spatial variation characteristics and variation trend of FPAR in different vegetation types in 2001 to 2018 were analyzed in the Hengduan Mountains. The response of FPAR to climate change was investigated by using Pearson correlation analytical method and partial least squares regression analysis. Results showed that the FPAR in Hengduan Mountains presented an increasing trend with time. Spatially, it was high in the south and low in the north, and it also showed obvious vertical zonality by elevation gradient.The vegetation FPAR was found to be positively correlated with air temperature and sunshine duration but negatively correlated with precipitation. Partial least squares regression analysis showed that the influence of sunshine duration on vegetation FPAR in Hengduan Mountains was stronger than that of air temperature and precipitation.     

Geoscience landscape division and tourism zonation in the mid-southern section of the Hengduan Mountains,eastern Qinghai-Tibet Plateau

The mid-southern section of the Hengduan Mountains is a typical region of mountainous landscape in western China and is the core region of "Shangri-La", a world-famous ecotourism destination. The landscape classification system is an important scientific basis for landscape protection and tourism development in this region. By combining geology and geography and referring to the concepts of "system tract" and "tectonic system" in geology, this paper comes up with grading standards for the geoscience landscape system of this region. Based on the regional stratigraphic structure, tectonic fault marks and geomorphological differentiation, this paper comes up with 2 Grade Ⅰ geoscience landscape system tracts, 8 Grade Ⅱ geoscience landscape systems, 21 Grade Ⅲ geoscience landscape areas, and 165 representative Grade Ⅳ geoscience landscape attractions. According to the main classification methods for the geological heritage and tourism landscapes, the geoscience landscapes can be divided into 4 categories, 16 types and 19 subtypes. On this basis, 23 eco-tourism areas of the mid-southern section of the Hengduan Mountains can be delimited. The study provides a theoretical direction and method reference for the geoscience landscape division and tourism zonation, which has an importantsignificance on the mountain landscape protection and tourism development in the regions of complex geo-environments.     

Alternative stable states in mountain forest ecosystems: the case of European larch(Larix decidua) forests in the western Alps

European larch(Larix decidua) forests of the western Alps form extensivecultural landscapes whose resilience to global changes is currently unknown. Resilience describes the capacity of ecological systems to maintain the same state, i.e., the same function, processes, structure, and composition despite disturbances, environmental changes and internal fluctuations. Our aim is to explore the resilience of larch forests to changes in climate and land use in the western Italian Alps.To do so, we examined whether larch forests can be described as an alternative stable state in mountain forest ecosystems. We used tree basal area data obtained from field forest inventories in combination with topography, forest structure, land use, and climate information. We applied three different probabilistic methods: frequency distributions, logistic regressions, and potential analyses to infer the resilience of larch forests relative to that of other forest types.We found patters indicative of alternative stable states: bimodality in the frequency distribution of the percent of larch basal area, and the presence of an unstable state, i.e., mixed larch forests, in the potential analyses. We also found:(1) high frequency ofpurelarchforestsathighelevation,(2)the probability of pure larch forests increased mostly with elevation, and(3) pure larch forests were a stable state in the upper montane and subalpine belts.Our study shows that the resilience of larch forests may increase with elevation, most likely due to the altitudinal effect on climate. Under the same climate conditions, land use seems to be the main factor governing the dominance of larch forests. In fact, subalpine larch forests may be more resilient, and natural succession after land abandonment, e.g., towards Pinuscembra forests, seems slower than in montane larch forests. In contrast, in the upper montane belt only intense land use regimes characterized by open canopies and forest grazing may maintain larch forests.We conclude that similar approaches could be applied in other forest ecosystems to infer the resilience of tree species.     

Spatial and temporal variation of drought index in a typical steep alpine terrain in Hengduan Mountains

This study describes the spatial and temporal variation of a drought index and makes inferences regarding the environmental factors that influence this variability in the Hengduan Mountains. A drought index is typically used to determine the moisture conditions and the magnitude of water deficiency in a given area. Based on data from 26 meteorological stations over the period 1960-2012, the spatial and temporal variations of the drought index were analyzed using a thin plate smoothing splines method that considered elevation as a covariate. The drought index was estimated based on the potential evapotranspiration (E₀) as defined by the Penman Monteith model modified by FAO (1998). The results of the reported analysis showed that the drought index in the Hengduan Mountains has been decreasing since 1960 at a rate of -0.008/a. This represented a progressive shift from the "sub-humid" class, which typified the wider area in the Hengduan Mountains, toward the "humid" class, which appeared in the Hengduan Mountains areas. The drought index was relatively high in the north and low in the south and the variation of the drought index varied with seasons. The drought index showed increasing trends in summer and autumn and it is greater in autumn than in summer, while it showed a decreasing trend in spring and winter. Drought index is inversely proportional to the soil relative humidity and Normalized Difference Vegetation Index (NDVI).     

Alternative stable states in mountain forest ecosystems: the case of European larch (<Emphasis Type="Italic">Larix decidua</Emphasis>) forests in the western Alps

European larch (Larix decidua) forests of the western Alps form extensive cultural landscapes whose resilience to global changes is currently unknown. Resilience describes the capacity of ecological systems to maintain the same state, i.e., the same function, processes, structure, and composition despite disturbances, environmental changes and internal fluctuations. Our aim is to explore the resilience of larch forests to changes in climate and land use in the western Italian Alps. To do so, we examined whether larch forests can be described as an alternative stable state in mountain forest ecosystems. We used tree basal area data obtained from field forest inventories in combination with topography, forest structure, land use, and climate information. We applied three different probabilistic methods: frequency distributions, logistic regressions, and potential analyses to infer the resilience of larch forests relative to that of other forest types. We found patters indicative of alternative stable states: bimodality in the frequency distribution of the percent of larch basal area, and the presence of an unstable state, i.e., mixed larch forests, in the potential analyses. We also found: (1) high frequency of pure larch forests at high elevation, (2) the probability of pure larch forests increased mostly with elevation, and (3) pure larch forests were a stable state in the upper montane and subalpine belts. Our study shows that the resilience of larch forests may increase with elevation, most likely due to the altitudinal effect on climate. Under the same climate conditions, land use seems to be the main factor governing the dominance of larch forests. In fact, subalpine larch forests may be more resilient, and natural succession after land abandonment, e.g., towards Pinus cembra forests, seems slower than in montane larch forests. In contrast, in the upper montane belt only intense land use regimes characterized by open canopies and forest grazing may maintain larch forests. We conclude that similar approaches could be applied in other forest ecosystems to infer the resilience of tree species.