Ecological tree line history and palaeoclimate – review of megafossil evidence from the Swedish Scandes

The postglacial tree line and climate history in the Swedish Scandes have been inferred from megafossil tree remains. Investigated species are mountain birch (Betula pubescens ssp. czerepanovii), Scots pine (Pinus sylvestris) and grey alder (Alnus incana). Betula and Pinus first appeared on early deglaciated nunataks during the Lateglacial. Their tree lines peaked between 9600 and 9000 cal. a BP, almost 600 m higher than present‐day elevations. This implies (adjusted for land uplift) that early Holocene summer temperatures may have been 2.3°C above modern ones. Elevational tree line retreat characterized the Holocene tree line evolution. For short periods, excursions from this trend have occurred. Between c. 12 000 and 10 000 cal. a BP, a pine‐dominated subalpine belt prevailed. A first major episode of descent occurred c. 8200 cal. a BP, possibly forced by cooling and an associated shift to a deeper and more persistent snow pack. Thereafter, the subalpine birch forest belt gradually evolved at the expense of the prior pine‐dominated tree line ecotone. A second episode of pine descent took place c. 4800 cal. a BP. Historical tree line positions are viewed in relation to early 21st century equivalents, and indicate that tree line elevations attained during the past century and in association with modern climate warming are highly unusual, but not unique, phenomena from the perspective of the past 4800 years. Prior to that, the pine tree line (and summer temperatures) was consistently higher than present, as it was also during the Roman and Medieval periods, c. 1900 and 1000 cal. a BP, respectively.     

Radiocarbon dating of subfossil Scots pine (Pinus sylvestris L.) in the southern Swedish Scandes

Wood samples from above the present-day pine limit were collected from within a limited area in central Sweden. New dates from an intensively investigated part of that area all fall within the period 8550 ± 110 to 4270 ± 90 ¹⁴C years B. P. No evidence at present exists to suggest that any drastic change, or fluctuations, in the pine tree-limit, took place within that period. Some dates support the view that the present-day altitude of the pine tree-limit was established during the historic period and perhaps is still declining.     

Holocene temperature history of northern Iceland inferred from subfossil midges

The Holocene temperature history of Iceland is not well known, despite Iceland's climatically strategic location at the intersection of major surface currents in the high-latitude North Atlantic. Existing terrestrial records reveal spatially heterogeneous changes in Iceland's glacier extent, vegetation cover, and climate over the Holocene, but these records are temporally discontinuous and mostly qualitative. This paper presents the first quantitative estimates of temperatures throughout the entire Holocene on Iceland. Mean July temperatures are inferred based upon subfossil midge (Chironomidae) assemblages from three coastal lakes in northern Iceland. Midge data from each of the three lakes indicate broadly similar temperature trends, and suggest that the North Icelandic coast experienced relatively cool early Holocene summers and gradual warming throughout the Holocene until after 3 ka. This contrasts with many sites on Iceland and around the high-latitude Northern Hemisphere that experienced an early to mid-Holocene “thermal maximum” in response to enhanced summer insolation forcing. Our results suggest a heightened temperature gradient across Iceland in the early Holocene, with suppressed terrestrial temperatures along the northern coastal fringe, possibly as a result of sea surface conditions on the North Iceland shelf.     

Holocene history of Lake Soldatskoje (Kola Peninsula, Russia) inferred from sedimentary diatom assemblages

A sediment core from Lake Soldatskoje, a small tundra lake located in the northern coastal area of the Kola Peninsula and surrounded by numerous archaeological dwelling sites, was analysed for diatom species. The core covers the entire Holocene, i.e. 10000 radiocarbon years. The diatom record has similarities with studies made earlier from tundra lakes of northern Russia and from Arctic lakes in general. The genus Fragilaria was dominant and many other small, benthic, nordic, cold-water diatom species typical of Arctic lakes were common, including Achnanthes minutissima Kützing, A. pusilla (Grunow) De Toni, Cyclotella tripartita Håkansson and Navicula absoluta Hustedt. Multivariable ordinations were used to characterize the changes in the diatom flora. Diatom-based pH and total phosphorus inferences indicate that the lake has become progressively more acidic and poorer in nutrients. Disturbances in the diatom stratigraphy of Lake Soldatskoje around 4000–5500 ¹⁴C yr BP may be related to human activity.     

A high-resolution Late Holocene landscape ecological history inferred from an intramontane basin in the Western Taurus Mountains,Turkey

Late Holocene vegetation and geomorphological history is reconstructed from a 800 cm long high-resolution palynological and sedimentological record sampled from Bereket, a 6.3 km² semi-arid to sub-humid intramontane basin in the Western Taurus Mountains (southwest Turkey). The well-dated Bereket record provides from cal. 360 BC to cal. AD ∼400 a unique record of biennial-to-decadal landscape changes caused primarily by intensive human impacts against a background of global climate variations. During this period, land clearance with multiple fire episodes, intensive agricultural practices and grazing pressure profoundly altered the pre-existing warm mixed forest. Increasing moisture availability since cal. ∼280 BC has acted as a trigger to crop cultivation and mountain-adapted arboriculture starting with Juglans regia during the Beyşehir Occupation Phase. Pollen from olive groves have been recorded above 1400 m a.s.l. only at cal. ∼23 BC and have disappeared definitively at cal. AD ∼294. During this phase, the sediment accumulation rate was extremely high, reflecting landscape instability. From cal. AD 450 to recent times, the area has mainly recorded pasture and minor cultivation activities reflected in stable soils and thin colluvial depths.     

Holocene climatic variations inferred from treeline fluctuations in the White Mountains,California

Remains of dead bristlecone pine (Pinus longaeva Bailey) are found at altitudes up to 150 m above present treeline in the White Mountains. Standing snags and remnants in two study areas were mapped and sampled for dating by tree-ring and radiocarbon methods. The oldest remnants represent trees established more than 7400 y.a. Experimental and empirical evidence indicates that the position of the treeline is closely related to warm-season temperatures, but that precipitation may also be important in at least one of the areas. The upper treeline was at high levels in both areas until after about 2200 B.C., indicating warm-season temperatures about 3.5°F higher than those of the past few hundred years. However, the record is incomplete, relative warmth may have been maintained until at least 1500 B.C. Cooler and wetter conditions are indicated for the period 1500 B.C.-500 B.C., followed by a period of cool but drier climate. A major treeline decline occurred between about A.D. 1100 and A.D. 1500, probably reflecting onset of cold and dry conditions. High reproduction rates and establishment of scattered seedlings at high altitudes within the past 100 yr represents an incipient treeline advance, which reflected a general climatic warming beginning in the mid-19th century that has lasted until recent decades in the western United States. This evidence for climatic variation is broadly consistent with the record of Neoglacial advances in the North American Cordillera, and supports Antevs' concept of a warm “altithermal age” in the Great Basin.     

Palaeoenvironmental history of the Holocene volcanic crater lake Lago d'Averno (central southern Italy) inferred from aquatic mollusc deposits

The mollusc record from Lago d'Averno, central southern Italy, provides a detailed 1600‐yr record of changes in water quality in response to bradyseismic movements and salinity fluctuations. Bradyseismic vertical land movements and human impact in Roman times led to several transgressions from the Mediterranean Sea, 1 km distant, making the lake a valuable resource for documenting the effect of episodic marine transgressions of a freshwater lake. Low‐oxygen‐tolerant freshwater molluscs suggest that, at around 900–500 bc the lake had a slowly decreasing medium freshwater quality, resulting from contamination of volcanic origin. Disappearance of the obligate freshwater snails and transgression of low‐salinity‐tolerant marine species indicate that, after 500 bc, continuous subsidence resulted in episodic marine transgressions from the nearby sea. The construction of a canal that connected the lake with the sea, in 37 bc, is marked by a considerable increase in the number of shells and by arrival of brackish‐water‐intolerant marine species. Species diversity increased considerably when the area was partly depopulated towards the end of the Roman Empire around ad 400. When the land was slightly uplifted around ad 600–750, the water quality of the lake became less favourable for marine molluscs. Copyright © 2000 John Wiley & Sons, Ltd.     

Holocene storminess inferred from sediments of two lakes on Adak Island,Alaska

The abundance of sedimentary organic material from two lakes was used to infer past Holocene storminess on Adak Island where frequent storms generate abundant rainfall and extensive cloud cover. Andrew and Heart Lakes are located 10 km apart; their contrasting physical characteristics cause the sedimentary organic matter to respond differently to storms. Their records were synchronized using correlated tephra beds. Sedimentation rates increased between 4.0 and 3.5 ka in both lakes. Over the instrumental period, Andrew Lake biogenic-silica content (BSi) is most strongly correlated with winter sunlight availability, which influences photosynthetic production, and river input, which influences the dilution of BSi by mineral matter. Heart Lake BSi is likely affected by wind-driven remobilization of sediment, as suggested by correlations among BSi, the North Pacific Index, and winter storminess. The results indicate relatively stormy conditions from 9.6 to 4.0 ka, followed by drying between 4.0 and 2.7 ka, with the driest conditions from 2.7 to 1.5 ka. The stormiest period was between AD 500 and 1200, then drying from 1150 to 1500 and more variable until 1850. This record of Holocene storminess fills a major gap at the center of action for North Pacific wintertime climate.     

A dendrochemical study of Pinus sylvestris from Siljansfors Experimental Forest,central Sweden

Inductively coupled plasma mass spectrometry (ICP-MS) was used to investigate changes in trace element concentration in two high resolution sequences of tree rings from central Sweden. Individual annual growth increments from 1800–2002 to 1930–2002 were sampled from two Scots pine (Pinus sylvestris) trees from the Siljansfors Experimental Forest. The aims of the study were: to test the viability of conventional solution induction ICP-MS as a technique for investigating the multi-elemental chemistry of long tree ring sequences at annual resolution, and, to test this specifically with a view to detecting changes in elemental concentrations of Swedish tree rings contemporary with the major (and relatively proximal) Icelandic eruption of Askja (1875). It was found that despite a time consuming sample preparation process, it was possible to use conventional ICP-MS for multi-elemental analysis of a long sequence of tree rings at annual resolution. Although promising data were produced, no truly conclusive concentration anomaly could be detected in the sequence to indicate the impact of the Askja eruption on environmental chemistry. Overall findings underlined the complexity of the tree/environment interaction and the cautious approach to data interpretation essential for any dendrochemical study.     

Holocene environmental and climatic changes inferred from Wulungu Lake in northern Xinjiang, China

Sedimentological, geochemical and palynological data from Wulungu Lake in northern Xinjiang, China, are used to reconstruct environmental and climate changes since 9550 cal yr BP. High abundance of Sparganium and Poaceae, low Md (median diameter) and δ¹³C_organic values indicate aridity between 9550 and 6730 cal yr BP. High Md and δ¹³C_organic values, and the prevalence of desert-steppe and steppe vegetation between 4200 and 560 cal yr BP, indicate that effective moisture increased after 6730 cal yr BP, peaking at 4200 and 560 cal yr BP. Low Md values, a negative excursion of δ¹³C_org, and the transition from steppe to desert vegetation since 560 cal yr BP reflect a decrease in effective moisture during the latest Holocene. Late Holocene human activities were indicated by sharp increase in the abundance of Pediastrum then. Variations in carbonate contents indicate that temperature was generally high between 9550 and 7740 cal yr BP, low between 7740 and 6730 cal yr BP, intermediate between 6730 and 560 cal yr BP, and low during the last 560 yr. Regional comparison indicates that the Asian monsoon did not extend to Wulungu Lake and westerlies were the main factor in determining the moisture availability during the Holocene.     

Holocene humidity fluctuations in Sweden inferred from dendrochronology and peat stratigraphy

Tree-ring and peat stratigraphy data were examined back to 5000 BC in order to identify and compare humidity changes in Fennoscandia. The temporal variation in distribution of Scots pine ( Pinus sylvestris L.) was used as a measure of past lake-level fluctuations in central Sweden. The chronology, which spans 2893 BC–AD 1998 with minor gaps in AD 887–907 and 1633–1650 BC and with additional floating chronologies back to 4868 BC, was cross-dated and fixed to an absolute timescale using a chronology from Torneträsk, northern Sweden. The peat stratigraphy from the Stömyren peat bog, south-central Sweden, was transformed into humification indices to evaluate humidity changes during the past 8000 years. The peat chronology is established by four tephra datings and eight ¹⁴C datings. Synchronous periods of drier conditions, interpreted from regeneration and the mortality pattern of pine, tree-ring chronology and peat humification, were recognized at c. 4900–4800 BC, 2400–2200 BC, 2100–1800 BC, 1500–1100 BC, AD 50–200, AD 400–600 and AD 1350–1500. Possible wetter periods were encountered at 3600–3400 BC, 3200–2900 BC, 2200–2100 BC, 1700–1500 BC, 1100–900 BC, 100 BC-AD 50, AD 200–400, AD 750–900 and AD 1550–1700. The wet and dry periods revealed by the tree rings and peat stratigraphy data indicate considerable humidity changes in the Holocene.     

Late Holocene high resolution palaeoclimatic reconstruction inferred from Sebkha Mhabeul, southeast Tunisia

Relations between climate change and landscape evolution during the last two millennia in southeastern coastal Tunisia have been documented using high-resolution reconstruction of flood history and fire activity in the Sebkha Mhabeul core. The age model, based on tephrochronology, indicates that the core extends from Roman to modern times and encompasses the well-defined climatic periods of the last two millennia. This record provides a first palaeoecological/palaeoclimatic high resolution reconstruction in North Africa using a cross-disciplinary approach with both physical (grey-scale intensity, quartz particles) and biological (charcoal and pollen) indicators. The flood history shows four wet/dry cycles (ca. AD 550-950, 950-1300, 1300-1570 and 1570-1870) of different duration. Major hydrological instabilities are concentrated during the Medieval Climate Anomalies and the early Little Ice Age, between AD 1000 and 1550. Direct correlation between climate and fire cannot be established suggesting that the fire history of the Sebkha environment is mainly influenced by human activity. This study demonstrates the great value of sebkhas as palaeoenvironmental archives.     

A Holocene dust record in arid central Asia inferred from Lake Barkol,northwest China

Grain-size distributions and detrital minerals were investigated for modern dust and for lacustrine sediment in Lake Barkol region, northwest China. Characteristics of the modern dust and lacustrine sediments, principle component analysis, and changes of quartz contents all suggest grain size 45–138 μm in the core sediment is the dust sensitive component. It indicates that high dust flux climate occurred at mid-Holocene (5.5–4.5 cal ka BP), corresponding to the high dust flux record in northwest Pacific and the drought interval of deserts in northwest China during the mid-Holocene. The strong dust activity is in contradiction with the climatic optimum recorded by the lacustrine evidence in Xinjiang. Different responses of the dust source region and watershed to the regional climate may have led to the inconsistency between dust and climate records during the mid-Holocene in arid central Asia.     

Deformation history inferred from magnetic fabric in the southwestern Okcheon metamorphic belt, Korea

Magnetic fabric and rock-magnetic studies have been carried out for the non-fossiliferous, low- to medium-grade metasedimentary rocks in the southwestern part of the Okcheon metamorphic belt (OMB). Two major metamorphic events in the study area were previously recognized: regional metamorphism (M₁) in the late Carboniferous to early Permian and contact metamorphism (M₂) due to the intrusion of granite in the middle Jurassic. The metamorphic grade of the study area increases from the biotite zone in southeast through the garnet zone to the staurolite zone towards the northwest. Magnetic fabrics of the study area are generally well defined and can be characterized according to the metamorphic zones. Magnetic foliation is the dominant magnetic fabric in the biotite zone, while magnetic lineation prevails in both garnet and staurolite zones. We interpret the metamorphism-related deformation history of the study area based on magnetic fabrics, magnetic mineralogy and previously reported metamorphic evolution as follows. Penetrative NW-dipping cleavage, represented by magnetic foliation, was formed in the study area by prevailing NW–SE shortening event during the M₁ regional metamorphism in the late Carboniferous–early Permian. This shortening event is interpreted to be associated with the collisional event between the North and South China blocks. Cleavages dipping steeply to the southeast in the staurolite zone are attributed to the pressure exerted from the intrusion of Jurassic granite in the northwestern area.     

Rapid change in early Holocene environments inferred from Lake Pupuke,Auckland City,New Zealand

Sediment cores from Lake Pupuke in Auckland City, New Zealand, contain a high‐resolution millennial to centennial‐scale record of changing climate and catchment hydrology spanning the past ca. 10 000 years. Here, we focus on the period between 9500 ± 25 and 7000 ± 155 cal. yr BP during which grain size, diatom palaeoecology, biogenic silica concentrations, sediment elemental and carbon isotope geochemistry reflect changes in sediment sources and lake conditions, with a significant event commencing at ca. 8240 cal. yr BP, commensurate with a lowering of lake level, faster erosion rates and increased sediment influx with a duration of ca. 360 yrs. However, the changes in the lake are not reflected in the terrestrial vegetation, where the pollen record indicates that podocarp forest dominated the Auckland region, with apparent environmental stability during this part of the early Holocene. The synchronous change in most of the proxies between ca. 8240 and 7880 cal. yr BP at Lake Pupuke indicates the presence of a sustained episode of relatively low lake level and concomitant increased rate of erosion in the early Holocene that appears to be at least partly coeval with the 8200 cal. yr BP meltwater event proposed for the North Atlantic region. Copyright © 2008 John Wiley & Sons, Ltd.     

The biogeographical role of Pinus forest on the Northern Spanish Meseta: a new Holocene sequence

The aim of this paper is to reconstruct the variation in the vegetation during the Holocene on the Northern Meseta through pollen analysis in order to determine the roles played by Pinus and Quercus in the vegetation dynamics. A new pollen sequence, Camporredondo, in the centre of the Duero River basin, Northern Meseta, Spain, is presented. It covers the period from ca 11,250 to 1630 cal yr BP. The Camporredondo sequence suggests that a forested landscape existed from the start of the Holocene with values of approximately 90% of arboreal pollen mainly derived from pine forests in the surroundings. Shortly after 8600 cal yr BP pine forests continued to dominate but the landscape became more open while stands of deciduous and evergreen Quercus developed. Around 4500 years ago, coinciding with the start of a period of less rainfall, the lacustrine areas in these territories began to silt up, favouring hydrophytic taxa such as Salix or Cyperaceae. A comparison with other Holocene pollen sequences from the central-eastern part of the Northern Meseta confirms the presence of pine forests in the whole region since the early Holocene. We discuss the variations of the general vegetation pattern in terms of the local or regional conditions at each site. Throughout the Holocene, Pinus forests dominated at least in the eastern half of the Duero River basin. The typical Holocene substitution of conifers by broadleaf species as recorded in other parts of the Iberian Peninsula and Europe, never takes place in the study area. The results of the present study suggest that the current view of the potential vegetation in the Spanish Northern Meseta should be reviewed.     

Holocene paleoenvironmental changes inferred from diatom assemblages in sediments of Kusawa Lake, Yukon Territory, Canada

The southwest Yukon Territory, Canada, is an important region for recovering sensitive records of Holocene paleoclimatic change. More information is needed, however, to constrain the timing of the major Holocene climatic transitions, and to understand associated impacts on different ecosystems. For example, paleolimnological studies have focused on small lakes and ponds, but the history of large lakes has received little study. We analyzed diatom assemblages, species richness, valve concentrations, and biogenic silica, in the sediments of Kusawa Lake (60°16.5'N; 136°10.9'W; 671 m a.s.l.) to reconstruct the responses of this large (surface area = 142 km²), deep (Z_max = 135 m) freshwater ecosystem to Holocene climatic transitions. Diatoms colonized the lake soon after ice retreat, around 11,000 cal yr BP; assemblages throughout the record were dominated by planktonic types. Diatom concentrations and biogenic silica were high during the Holocene Thermal Maximum between 10,700 and 7300 cal yr BP, then began to decrease in response to cooling associated with orbitally driven reductions in insolation. Diatom assemblages shifted towards taxa with lower surface water temperature optima after 8300 cal yr BP, perhaps in response to abrupt and progressive cooling. Our study confirms that diatom assemblages in large lakes are sensitive to regional-scale paleoclimatic changes.     

Monsoonal changes inferred from the Middle to Late Holocene sediments off Landfall Island,North Andaman

In this paper, sediment texture, clay mineral composition, and δ ¹⁸O data on Globigerinoides ruber of a sediment core collected from a water depth of 250 m near Landfall Island, Bay of Bengal, is presented to understand paleoenvironmental shifts during the Mid–Late Holocene period. From the sediment core, five organic carbon-rich samples were radiocarbon dated and the reservoir-corrected ages range from 6,078 to 1,658 years BP. The marine sediment core is represented dominantly by clayey silt with incursions of coarser components that occur around 6,000, 5,400, and 3,400 years BP. The sedimentation of the coarser detritus is due to strengthened southwest monsoon (SWM) since 6,500 to 5,400 years BP. Clay minerals are represented by smectite, illite, kaolinite, and chlorite in varying amounts. High kaolinite content and K/C ratio indicate intense SWM and strong bedrock weathering from the hinterland (~6,500–5,400 years BP). Incidence of smectite (48.82 to 25.09 %) and chlorite/illite (C/I) ratio (0.56 to 0.28) indicate an overall weakened southwest monsoon since 6,000 to 2,000 years BP with a brief incursion of extremely reduced SWM around 4,400 to 4,200 years BP. This is corroborated with the oxygen isotope on G. ruber that reveals a significant shift in the isotopic values ~4,300 years BP (?3.39?‰), indicating weakening in SWM. Subsequently, fluctuations in the intensity of SWM are observed since 2,000 years to present.     

Holocene vegetation history of the Okanogan Valley,Washington

Haploxylon pine(s) and Artemisia dominated the initial vegetation in front of the receding Okanogan Lobe until ca. 10,000 yr B.P., as revealed by two pollen records in north-central Washington. After 10,000 yr B.P. the macroclimate became warmer throughout the Okanogan drainage as diploxylon pines and Artemisia increased. The Mount Mazama eruption at ca. 6700 yr B.P. is recorded as two stratigraphically separate and petrographically distinct tephra units at Bonaparte Meadows. While there are apparent short-term changes in the vegetation coincident with the ashfall(s), Artemisia continues to dominate the Okanogan Valley until ca. 5000 yr B.P. By 4700 yr B.P. the modern vegetation, dominated by Pseudotsuga menziesii, had become established around Bonaparte Meadows.     

Holocene variations in the Asian monsoon inferred from the geochemistry of lake sediments in central Tibet

We present a record of monsoon variations for the early and middle Holocene that is inferred from the geochemistry of sediment cores from Ahung Co, a lake in central Tibet. The resolution of this record is better than 50 yr and the age model is derived from radiocarbon ages of terrestrial charcoal, which eliminates errors associated with the lake hard-water effect. We made down-core geochemical measurements of % carbonate, % organic carbon, C/N and δ¹³C of bulk organic matter, δ¹³C and δ¹⁸O of carbonate, and % dolomite. Proxy calibration and modern water-balance reconstruction show that these are proxies for lake depth and the amount of monsoon precipitation. We find that lake level and monsoon precipitation have been decreasing at Ahung Co since the early Holocene (∼7500 cal yr B.P.). Superimposed on this trend are rapid declines in monsoon rainfall at 7000-7500 and 4700 cal yr B.P. and seven century-scale wet-dry oscillations. The cores do not contain sediment from the last ∼4000 yr. Surface sediments from the lake accumulated during the 20th century, however. From this, we argue that lake levels have risen again recently following a late Holocene dry period.