A New Fossil Hangingfly (Mecoptera: Cimbrophlebiidae) from the Early Cretaceous of China

A new species, Cimbrophlebia rara sp. nov., in the family Cimbrophlebiidae（Mecoptera） is described and illustrated. This specimen was collected from the Early Cretaceous Yixian Formation at Liutiaogou in Inner Mongolia, China. This is the first record of a cimbrophlebiid from the Jehol biota. A key to all species of Cimbrophlebia is given. A preliminary review of published taxa data indicates that from the Middle Jurassic to the Early Cretaceous, Mecoptera became less diverse and less abundant in northeastern China at familiar level（from 11 to 6）, generic level（from 32 to 8） and specific level（from 44 to 14）.     

New Jurassic Pseudopolycentropodids from China (Insecta: Mecoptera)

Abstract: In this paper two new species of fossil Pseudopolycentropus Handlirsch, 1906 are described: Pseudopolycentropus janeannae sp. nov. and P. novokshonovi, sp. nov. All of them were recovered from the Middle Jurassic non-marine sedimentary strata of northeastern China. The new material from China reveals that the early diversification of pseudopolycentropodids was well underway by the Middle Jurassic.     

New Species of Cretolimonia and Mesotipula (Diptera: Limoniidae) from the Middle Jurassic of Northeastern China

Two new species, Cretolimonia excelsa sp. nov. and Mesotipula gloriosa sp. nov., in Limoniidae of Diptera are described and illustrated, based on three fossil specimens with bodies and complete wings. These specimens were collected from the latest Middle Jurassic Jiulongshan Formation at Daohugou, Inner Mongolia, China. These are the first fossil records of these two genera from China.     

New Genus and Species of Rhyacophilidae (Insecta: Trichoptera) from the Middle Jurassic of China

One new genus with a new fossil species,Declinimodus setulosus gen.et sp.nov.which was found in the Middle Jurassic Jiulongshan Formation of Daohugou,Inner Mongolia,China,are described and illustrated.The new species is assigned to the Rhyacophilidae according to buttonshaped of the 2nd segment of maxillary palp,and the forked R1 （in the forewing,located near apex）.These new discovery are the earliest fossil records of Rhyacophilidae,which providing important evident for the study of the origin and early evolution of Trichoptera.     

A New Species of the Family Juraperlidae(Insecta:Grylloblattida)from the Middle Jurassic of China

<正>Juraperla grandis sp.nov.(Grylloblattida:Juraperlidae) is described from the Daohugou locality (Middle Jurassic,China).Its wing venation is very similar to that of Juraperla daohugouensis Huang and Nel,2007 (Grylloblattida:Juraperlidae),in the particular occurrence of a supplementary longitudinal vein in the area between the anterior wing margin and ScP.The larger size of the new specimen,the lower number of CuA1 branches,and the occurrence of two rows of cells between MP and CuA in forewings justify the erection of a new species,J.grandis sp.nov..The material also preserved hind wing and body structures,described for the first time in Juraperlidae.     

A New Mesopsychid (Mecoptera) from the Middle Jurassic of Northeastern China

The family Mesopsychidae Tillyard,1917 presently consists of ten described genera from the Early Permian to the Early Cretaceous of Australia,China,Kyrgyzstan,Russia,South Africa,Tajikistan,and Ukraine.Herein,a new genus and a new species of fossil mesopsychid,Epicharmesopsyche pentavenulosa gen.et sp.nov.,is described from the Middle Jurassic Jiulongshan Formation of Inner Mongolia,China,supplementing the family-level diversity.The well-preserved,new material reveals many characters,including antennae,wing venation,shape and genitalia that increase our morphological understanding of the family,although unfortunately the mouthparts and most of the legs are not preserved.Diagnosis of the Mesopsychidae is emended to include a four-or five-branched MP in both the fore-and hind wings.This is the first documentation of a wing coupling structure in the Mesopsychidae,consisting of three to four frenula bristles on the humeral lobe at the base of the costal margin of both hind wings.Asymmetrical shape and size of the left and right wings on a specimen of E.pentavenulosa gen.et sp.nov.,seems to be a common condition for mesopsychid taxa from northeastern China.     

A New Species of Liadotaulius (Insecta: Trichoptera) from the Middle Jurassic of Daohugou,Inner Mongolia

Diverse caddis flies are recently collected from the Middle Jurassic Daohugou fauna in Ningcheng County, Inner Mongolia, northern China, and different types of caddis cases have been reported in the same locality. A new species of caddisfly, Liadotaulius daohugouensis sp. nov. is described and illustrated; the establishment is based on a well-preserved isolated forewing collected from the Middle Jurassic Jiulongshan Formation of Daohugou. The family status of Liadotaulius is discussed on the basis of new material and is referred to the family Philopotamidae. The Daohugou fauna continues to provide new data for studying the early evolution of Trichoptera.     

A New Axymyiid Genus with Two New Species from the Middle Jurassic of China (Diptera: Nematocera: Axymyiidae)

The Axymyiidae is one of the small families of the suborder Nematocera within the Diptera.Up to date,three genera and seven species of extant axymyiids have been described from the Holarctic Region,including Canada,China,Hungary,Japan,Russia,and USA,with three genera and three species of fossil taxa described from the Middle Jurassic of Daohugou,northeastern China.A new genus Raraxymyia gen.nov.and two new species,R.parallela and R.proxima gen.et sp.nov.from the Middle Jurassic Jiulongshan Formation of Daohugou Village,Inner Mongolia,China,are herein described based on their well-preserved wings and body characters.A revised key to the genera of axymyiids is provided.     

A New Fossil Scorpionfly (Insecta: Mecoptera: Holcorpidae) with Extremely Elongate Male Genitalia from Northeastern China

Since Holcorpa maculosa was first described in 1878, it has attracted lots of attention due to its extremely elongate abdominal segments from the sixth to the eighth. Fossil records of family Holcorpidae are very rare, comprising two species in one genus represented by three fossil specimens, resulting in limited knowledge on the structure and function of holcorpid’s unique male organ, as well as its evolutionary relationship with other scorpionflies. Herein we describe a new genus with a new species, Conicholcorpa stigmosa gen. et sp. nov., of the Holcorpidae from the Middle Jurassic Jiulongshan Formation near Daohugou Village, Inner Mongolia, China. This is the first holcorpid documented from the Daohugou locality, albeit two previously reported species with extremely elongate male organs but deemed as Family Incertae sedis. C. stigmosa gen. et sp. nov. is the earliest holcorpid hitherto, extending its existence from the Early Eocene to the Middle Jurassic by 114 million years and providing one more early case of presenting exaggerated male body parts for sexual display and/or selection. Our analysis supports the sister relationship between Holcorpidae and Orthophlebiidae. Our results not only enhance our knowledge of the morphology of Holcorpidae but also elucidate the relationship of known species of this family.     

A new species of Eomeropidae (Insecta: Mecoptera) from the Middle Jurassic of China

The Eomeropidae is a species-poor family with only 11 known fossil species from the Early Jurassic to Palaeogene. A new species of fossil eomeropid, Tsuchingothauma gongi sp. nov., is described based on a well-preserved wing from the Middle Jurassic Daohugou deposits of Inner Mongolia, China. Our new species is distinguished from the type species T. shihi mainly in having very numerous crossveins and cells; more longitudinal veins: 13 and 11 terminal branches in radial sector and medial field, CuA with 3 terminal branches and A with 7 terminal branches.     

Two New Species of Protocedroxylon Gothan (Pinaceae) from the Middle Jurassic of Eastern Inner Mongolia, NE China

Two new coniferous wood taxa, Protocedroxylon zhangii sp. nov. and P. zhalantunense sp. nov., are described from the Middle Jurassic Wanbao Formation in Zhalantun City, Inner Mongolia, NE China. The new discovery represents the first record of petrified wood in the Wanbao Formation, and contributes to further understanding the floral composition, especially that of the forest, of the Wanbao Formation. Anatomically, the two new species are characterized by having a mixed type of radial pitting and Abietineentüpfelung. In extant conifers, Abietineentüpfelung is considered to be characteristic of the wood of Pinaceae. The new discovery further indicates that the first occurrence of Abietineentüpfelung in conifer-like wood should be earlier than the Kimmeridgian. The palaeolatitudinal distribution pattern of Protocedroxylon indicates that the genus is a palaeobiogeographically consistent group, which was probably restricted to wetter and cooler temperate climates. In combination with distinct growth rings, the occurrence of Protocedroxylon suggests that a cool temperate and seasonal climate might have prevailed during the sedimentation period of the Wanbao Formation in eastern Inner Mongolia, NE China. Such a view is also in agreement with the palaeoclimatic conditions indicated by the plant megafossils of the Wanbao Formation.     

A new caraboid larva from the Middle Jurassic of China (Insecta: Coleoptera)

A new species of caraboid larva (Coleoptera, Adephaga), Carabilarva gongi sp. nov., is described based on a well-preserved specimen from the Middle Jurassic Daohugou deposits (in the upper part of Jiulongshan Formation) of Inner Mongolia, China. It is the fourth caraboid larva from the Mesozoic, and differs from the other three Mesozoic species mainly in having a longer body with shorter mandibles. Scanning electron microscopy (SEM) and energy dispersive X-ray spectroscopy (EDS) show that this Daohugou fossil is preserved as carbonaceous compressions that lost the micro-structure, such as macrochaetae.     

Two New Fossil Lacewing Species from the Middle Jurassic of Inner Mongolia, China (Neuroptera: Grammolingiidae)

Two new species of lacewings are described and illustrated from the Jiulongshan Formation (Middle Jurassic) of Inner Mongolia, China: Leptolingia calonervis sp. nov. and Litholingia ptesa sp. nov. (Grammolingiidae). The species can be distinguished by the following features: the outermost branch of MP2 on the hind wing did not reach the margin of wing in Leptolingia calonervis sp. nov.; the second branch of Rs forked deeply while other branches of Rs forked much later in Litholingia ptesa sp. nov..     

A New Elaterid from the Middle Jurassic Daohugou Biota (Coleoptera: Elateridae: Protagrypninae)

A new genus and species of fossil elaterid namely Clavelater ningchengensis gen. et sp. nov. assigned to the subfamily Protagrypninae based on one specimen from the Middle Jurassic Jiulongshan Formation near Daohugou Village, Ningcheng County, Inner Mongolia, northeastern China is described and illustrated. This specimen displays a pair of large antennae that easily differs from those of most Mesozoic elaterids. There are around 36 species within 20 genera referred to elateroids described from the Early Jurassic to Miocene of China, with another three species within three genera reviewed, but the taxonomic assignment of at least seven species within three genera is doubted. The systematic position of fossil elateriform beetles reported from China is reviewed and the evolution of Mesozoic elateroids briefly discussed.     

New Mesozoic Mesopsychidae(Mecoptera)from Northeastern China

<正>The Mesozoic family Mesopsychidae Tillyard,1917 presently consists of seven described genera and 13 species from the mid-Triassic to the Early Cretaceous of Australia,South Africa and Eurasia.In the present paper one new genus and three new species of fossil mesopsychids are described that add significant distributional and stratigraphic extensions to the family.This finding documents the first formal record of fossil Mesopsychidae in China.Both Lichnomesopsyche gloriae gen.et sp.nov.and L.daohugouensis gen.et sp.nov. were found from the Middle Jurassic Jiulongshan Formation of Inner Mongolia,whereas Vitimopsyche kozlovi sp. nov.,of mid Early Cretaceous age,was collected from the Jehol Biota of northern Hebei Province.This new,well-preserved material from China reveals complete and previously unknown body features,including head, antennae,mouthparts,legs and abdomen.The delicate and long proboscides of these new taxa indicate that they were feeding on externally exposed,nutrient-rich fluids of gymnospermous ovulate fructifications,and incapable of piercing surface epidermis-attributable principally to the absence of stylets.These proboscides originated, perhaps multiply,among basal Mecoptera and are functionally and structurally convergent with equivalent mouthparts borne by fossil and extant Diptera,Lepidoptera,Neuroptera and Coleoptera.     

A New Species of Phoenicopsis Subgenus Windwardia (Florin) Samylina (Czekanowskiales) from the Middle Jurassic of Inner Mongolia, China

A new species of Phoenicopsis subgenus Windwardia(Florin) Samylina, Phoenicopsis neimengguensis sp.nov., is recognized from the Middle Jurassic Zhaogou Formation in the Shiguai Basin, a Jurassic coal-bearing sedimentary basin in Inner Mongolia, China. Linear leaves are in bundles of six to eight and are attached on a short shoot. The leaf is amphistomatic with one or two irregular rows of longitudinally oriented stomata in each intercostal zone on the upper epidermis and three or four(occasionally six) rows of stomata in each intercostal zone on the lower epidermis. Each stomata is surrounded by four to six subsidiary cells with unevenly developed papillae. The anticlinal walls of epidermal cells are straight. Cuticular thickenings or papillae are present on the outer surface of the periclinal walls. This is the first time that Phoenicopsis subgenus Windwardia(Florin) Samylina is recognized from the Middle Jurassic of Inner Mongolia, China. The occurrence of the new species is noteworthy because it extends the geological and geographic distribution of Phoenicopsis subg. Windwardia in China and Eurasia. The discovery is significant as it increases our knowledge of Phoenicopsis subg. Windwardia by providing new data on morphological and epidermal features.     

蚊蝎蛉科(昆虫纲:长翅目)的研究历史及发展

本文回顾了世界蚊蝎蛉科化石的研究进展,列出已发表的化石种类名录、分布及年代,共28属52种,分布于美洲、澳大利亚、欧洲和亚洲,但其中又以德国、前苏联和中国最多.简要探讨了蚊蝎蛉科的古地理和迁移方面的问题.认为欧亚大陆可能是蚊蝎蛉科的起源地,亚洲是它们的分化扩散中心,并在始新世迁移扩散到其它各洲.分析了蚊蝎蛉科化石研究中存在的一些问题和今后努力的方向.     

New Isophlebioid Dragonflies from the Middle Jurassic of Inner Mongolia, China (Insecta: Odonata: Isophlebioptera:Campterophlebiidae)

Three new species of fossil dragonflies assigned to Sinokaratawia Nel, Huang and Lin in family Campterophlebiidae, i.e. S. daohugouica sp. nov., S. magica sp. nov. and S. gloriosa sp. nov., and new materials of male S. prokopi Nel, Huang and Lin, 2007 are described from the Middle Jurassic of Daohugou, Inner Mongolia, China. An emended diagnosis of genus Sinokaratawia was proposed.     

New Isophlebioid Dragonflies from the Middle Jurassic of Inner Mongolia,China(Insecta:Odonata:Isophlebioptera:Campterophiebiidae)

Three new species of fossil dragonflies assigned to Sinokaratawia Nel,Huang and Lin in family Campterophlebildae,i.e.S,daohugouica sp.nov.,S.magica sp.nov.and S.gloriosa sp.nov.,and new materials of male S.prokopi Nel,Huang and Lin,2007 are described from the Middle Jurassic of Daohugou,Inner Mongolia,China.An emended diagnosis of genus Sinokaratawia was proposed.     

New Jurassic Fossil True Bugs of the Pachymeridiidae(Hemiptera:Pentatomomorpha)from Northeast China

Four new fossil genera and species of true bugs from the family Pachymeridiidae, Beipiaocoris multifurcus, Bellicoris mirabilis, Nitoculus regiUus, and Viriosinervis stolidus, are described. New specimens were collected from the Middle and Upper Jurassic non-marine sedimentary strata from the Jiulongshan and Yixian Formations of northeast China. The species Karatavocoris asiatica Becker-Migdisova, 1963, which was considered to be a member of the family Coreidae is transferred to the Pachymeridiidae. A new map of all known and newly discovered fossil pachymeridiid localities is given. The diagnosis of the family is modified.