The importance of plant macrofossils in the reconstruction of Lateglacial vegetation and climate: examples from Scotland,western Norway,and Minnesota,USA

Lateglacial and early Holocene (ca 14–9000 ¹⁴C yr BP; 15–10,000 cal yr BP) pollen records are used to make vegetation and climate reconstructions that are the basis for inferring mechanisms of past climate change and for validating palaeoclimate model simulations. Therefore, it is important that reconstructions from pollen data are realistic and reliable. Two examples of the need for independent validation of pollen interpretations are considered here. First, Lateglacial-interstadial Betula pollen records in northern Scotland and western Norway have been interpreted frequently as reflecting the presence of tree-birch that has strongly influenced the resulting climate reconstructions. However, no associated tree-birch macrofossils have been found so far, and the local dwarf-shrub or open vegetation reconstructed from macrofossil evidence indicates climates too cold for tree-birch establishment. The low local pollen production resulted in the misleadingly high percentage representation of long-distance tree-birch pollen. Second, in the Minnesotan Lateglacial Picea zone, low pollen percentages from thermophilous deciduous trees could derive either from local occurrences of the tree taxa in the Picea/Larix forest or from long-distance dispersal from areas further south. The regionally consistent occurrence of low pollen percentages, even in sites with local tundra vegetation, and the lack of any corresponding macrofossil records support the hypothesis that the trees were not locally present. Macrofossils in the Picea zone represent tundra vegetation or Picea/Larix forest associated with typically boreal taxa, suggesting it was too cold for most thermophilous deciduous trees to grow. Any long-distance tree pollen is not masked by the low pollen production of tundra and Picea and Larix and therefore it is registered relatively strongly in the percentage pollen spectra.Many Lateglacial pollen assemblages have no recognisable modern analogues and contain high representations of well-dispersed ‘indicator’ taxa such as Betula or Artemisia. The spectra could have been derived from vegetation types that do not occur today, perhaps responding to the different climate that resulted from the different balance of climate forcing functions then. However, the available contemporaneous plant-macrofossil assemblages can be readily interpreted in terms of modern vegetation communities, suggesting that the pollen assemblages could have been influenced by mixing of locally produced pollen with long-distance pollen from remote vegetation types that are then over-represented in situations with low local pollen production. In such situations, it is important to validate the climate reconstructions made from the pollen data with a macrofossil record.     

Full-glacial — late-glacial palaeoclimate of the Southern Andes: evidence from pollen,beetle and glacial records

Palaeoecological studies carried out in the Chilean Lake District and Chilotan Archipelago (41°–43°S) record full-glacial and late-glacial pollen assemblages beginning just after 21000 and beetle assemblages after 18000, both sets extending until 10000 ¹⁴C yr BP. Pollen records indicate that Subantarctic Parkland, the vegetation of the early millennia of record, changed after about 14000 yr BP to become open woodland and later North Patagonian Evergreen Forest. Assemblages of plants and beetles, responding more or less in unison to a strong rise in temperature (≥ 6°C), behaved in accord at around 14000 until 13000–12500 yr BP, the beetle fauna displaying a marked increase in obligate forest types. During full-glacial conditions (17400–16100 and 15300 and 14400 yr BP) and in the late-glacial interval (after about 13000 yr BP), however, climate evidently coerced populations dissimilarly, the pollen sequence showing an increase in plant taxa indicative of colder climate, whereas the beetle fauna underwent little or no variation. Contrasting climate modes implied by plants and beetles may be attributed to differential responses to apparent low-order temperature changes (≤ 2–3°C).     

Quaternary vegetation history of the Mississippi embayment

Nonconnah Creek, located in the loess-mantled Blufflands along the eastern wall of the Lower Mississippi Alluvial Valley in Tennessee displays a sedimentary sequence representing the Altonian Substage through the Woodfordian Substage of the Wisconsinan Stage. The site has a biostratigraphic record for the Altonian and Farmdalian Substages that documents warm-temperate upland oak-pine forest, prairie, and bottomland forest. At 23,000 yr B.P., white spruce and larch migrated into the Nonconnah Creek watershed and along braided-stream surfaces in the Mississippi Valley as far as southeastern Louisiana. The pollen and plant-macrofossil record from Nonconnah Creek provides the first documentation of a full-glacial locality in eastern North America for beech, yellow poplar, oak, history, black walnut, and other mesic deciduous forest taxa. During the full and late glacial, the Mississippi Valley was a barrier to the migration of pine species, while the adjacent Blufflands provided a refuge for mesic deciduous forest taxa. Regional climatic amelioration, beginning about 16,500 yr B.P., is reflected by increases in pollen percentages of cooltemperate deciduous trees at Nonconnah Creek. The demise of spruce and jack pine occurred 12,500 yr B.P. between 34° and 37° N in eastern North America in response to postglacial warming.     

Synchronization of late-glacial vegetation changes at Crystal Lake, Illinois, USA with the North Atlantic Event Stratigraphy

Late-glacial (17–11 cal ka BP) pollen records from midwestern North America show similar vegetation trends; however, poor dating resolution, wide-interval pollen counts, and variable sedimentation rates have prevented the direct correlation with the North Atlantic Event Stratigraphy as represented in the Greenland ice-core records, thus preventing the understanding of the teleconnections and mechanisms of late-Quaternary events in the Northern Hemisphere. The widespread occurrence of late-glacial vegetation and climates with no modern analogs also hinders late-glacial climate reconstructions. A high-resolution pollen record with a well-controlled age model from Crystal Lake in northeastern Illinois reveals vegetation and climate conditions during the late-glacial and early Holocene intervals. Late-glacial Crystal Lake pollen assemblages, dominated by Picea mariana and Fraxinus nigra with lesser amounts of Abies and Larix, suggest relatively wet climate despite fluctuations between colder and warmer temperatures. Vegetation changes at Crystal Lake are coeval with millennial-scale trends in the NGRIP ice-core record, but major shifts in vegetation at Crystal Lake lag the NGRIP record by 300–400 yr. This lag may be due to the proximity of the Laurentide ice sheet, the ice sheet's inherent slowness in response to rapid climate changes, and its effect on frontal boundary conditions and lake-effect temperatures.     

Late Quaternary vegetational changes on the east side of Yellowstone park,Wyoming

A bog pond 4 km east of Yellowstone Lake has a pollen record starting with an Artemisia spruce assemblage, implying alpine vegetation. A layer of volcanic ash, dated as 14,360 ± 400 BP (probably Glacier Peak or Mt. St. Helens J), occurs within the zone, which terminates at 11,630 ± 180 BP. The rest of the pollen sequence is dominated by lodgepole pine, with reappearance of spruce pollen in modest quantities about 4500 BP, according to dating provided by a layer of Mt. Mazama ash (6600 BP). The present vegetation of the area is marked by forests of lodgepole pine with some stands of spruce and fir. The pollen sequence suggests that the upper treeline before 11,600 y. a. was perhaps 500 m lower than today. The climate then became warmer and/or drier than today (Altithermal interval). About 4500 y. a., a slight climatic reversal took place, roughly contemporaneous with the regrowth of glaciers in the western mountains (neoglaciation).     

Expanded response-surfaces: a new method to reconstruct paleoclimates from fossil pollen assemblages that lack modern analogues

Pollen-based paleoclimatic interpretations of late-glacial to early Holocene climates (17–9 ka) in Midwestern North America are hampered by samples that lack modern analogues. Unresolved questions include the magnitude and direction of temperature seasonality (i.e. were these climates more or less seasonal than present) and the temporal changes in precipitation. Central to the no-analogue problem is the truncation of modern pollen-climate relationships for abundant late-glacial taxa such as Fraxinus. Here we present a new method called the expanded response-surface (ERS) method, developed to reconstruct climates from no-analogue pollen assemblages and applied to a high-resolution late-glacial pollen record from Crystal Lake, Illinois to test hypotheses about late-glacial climates. The key assumptions central to the ERS method are: (1) plant species and pollen abundances follow symmetrical unimodal distributions along climatic gradients, (2) taxa with truncated distributions in modern climate space occupy a subset of their fundamental niche, and (3) expansion of truncated distributions by mirroring around the distributional mode recovers the portion of the fundamental niche not realized in the modern climate space. With the ERS method, we expanded modern pollen-climate relationships by mirroring pollen abundances for each taxon around a mode defined with respect to four climate axes (mean winter temperature, mean summer temperature, mean winter precipitation, and mean summer precipitation). The ERS method reconstructed past temperatures and precipitation during the height of no-analogue conditions (14 160–12 370 cal yr BP) for 37% of the Crystal Lake samples where techniques that employed only modern observational data found matches for only 13% of the fossil samples. The total climate space of the expanded taxa set allowed analogue matches under more seasonal-than-present climates with higher-than-present precipitation. The ERS climate reconstructions for the height of no-analogue conditions indicated cooler-than-present summer and winter temperatures, similar-to-present seasonal range in temperatures, higher-than-present winter precipitation, and similar-to-present summer precipitation. These results thus suggest that high moisture availability helped drive the formation of the Midwestern no-analogue communities with high Fraxinus nigra abundances, but do not show higher-than-present temperature seasonality notwithstanding the higher-than-present insolation seasonality at this time. During the no-analogue late-glacial interval, Picea mariana, F. nigra, and Larix stands probably grew on low-lying, poorly drained soils in the Crystal Lake region; whereas Abies, Picea glauca, Quercus, and Ostrya/Carpinus grew on upland positions with better soil drainage.     

Southern Westerlies during the last glacial maximum

Vegetation and climate over approximately the past 13,000 yr are reconstructed from fossil pollen in a 9.4-m mire section at Caleta Róbalo on Beagle Channel, Isla Navarino (54°56′S, 67°38′W), southern Tierra del Fuego. Fifty surface samples reflecting modern pollen dispersal serve to interpret the record. Chronologically controlled by nine radiocarbon dates, fossil pollen assemblages are: Empetrum-Gramineae-Gunnera-Tubuliflorae (zone 3b, 13,000–11,850 yr B.P.), Gramineae-Empetrum-assorted minor taxa (zone 3a, 11,850-10,000 yr B.P.), Nothofagus-Gramineae-Tubuliflorae-Polypodiaceae (zone 2, 10,000–5000 yr B.P.), Nothofagus-Empetrum (zone 1b, 5000-3000 yr B.P.), and Empetrum-Nothofagus (zone 1a, 3000-0 yr B.P.). Assemblages show tundra under a cold, dry climate (zone 3), followed by open woodland (zone 2), as conditions became warmer and less dry, and later, with greater humidity and lower temperatures, by closed forest and the spread of mires (zone 1). Comparisons drawn with records from Antarctica, New Zealand, Tasmania, and the subantarctic islands demonstrate broadly uniform conditions in the circumpolar Southern Hemisphere. The influences of continental and maritime antarctic air masses were apparently considerable in Tierra del Fuego during cold late-glacial time, whereas Holocene climate was largely regulated by interplay between maritime polar and maritime tropical air.     

A late Pleistocene long pollen record from Lake Urmia, NW Iran

A palynological study based on two 100-m long cores from Lake Urmia in northwestern Iran provides a vegetation record spanning 200 ka, the longest pollen record for the continental interior of the Near East. During both penultimate and last glaciations, a steppe of Artemisia and Poaceae dominated the upland vegetation with a high proportion of Chenopodiaceae in both upland and lowland saline ecosystems. While Juniperus and deciduous Quercus trees were extremely rare and restricted to some refugia, Hippophaë rhamnoides constituted an important phanerophyte, particularly during the late last glacial period. A pronounced expansion in Ephedra shrub-steppe occurred at the end of the penultimate late-glacial period but was followed by extreme aridity that favoured an Artemisia steppe. Very high lake levels, registered by both pollen and sedimentary markers, occurred during the middle of the last glaciation and late part of the penultimate glaciation. The late-glacial to early Holocene transition is represented by a succession of Hippophaë, Ephedra, Betula, Pistacia and finally Juniperus and Quercus. The last interglacial period (Eemian), slightly warmer and moister than the Holocene, was followed by two interstadial phases similar in pattern to those recorded in the marine isotope record and southern European pollen sequences.     

Abrupt Climatic Changes during the Last Glaciation Based on Pollen Analysis of the Abric Romani, Catalonia, Spain

The Abric Romani continental pollen record provides evidence of abrupt climate variations during the last glaciation. The pollen record reveals a pattern of high-frequency variations similar to that in ice cores as shown by oxygen isotopes or dust content. Analyses of 14 travertine samples yield coherent U-series ages in correct stratigraphic order, ranging from 40,000 to 70,000 yr. According to changes in the composition and structure of the vegetation, five paleoclimatic phases have been differentiated; the oldest, ranging from 70,200 to 65,500 yr ago, records thermophilous taxa which correspond to the last warm events of isotopic stage 5. The next phase, from 65,500 to 56,800 yr, records a cold, humid climate that correlates with isotope stage 4. During the third phase, from 56,800 to 49,500 yr, relatively warm and cold events alternate. The fourth phase, from 49,500 to 46,200 yr, records a cold, dry climate. The fifth phase, ranging from 46,200 to 40,800 yr, records a milder phase that is similar in appearance to the beginning of the Holocene.     

Late-glacial and Holocene vegetation and climatic history of the Cass Basin, central South Island, New Zealand

Lithology, pollen, macrofossils, and stable carbon isotopes from an intermontane basin bog site in southern New Zealand provide a detailed late-glacial and early Holocene vegetation and climate record. Glacial retreat occurred before 17,000 cal yr B.P., and tundra-like grassland–shrubland occupied the basin shortly after. Between 16,500 and 14,600 cal yr B.P., a minor regional expansion of forest patches occurred in response to warming, but the basin remained in shrubland. Forest retreated between 14,600 and 13,600 cal yr B.P., at about the time of the Antarctic Cold Reversal. At 13,600 cal yr B.P., a steady progression from shrubland to tall podocarp forest began as the climate ameliorated. Tall, temperate podocarp trees replaced stress-tolerant shrubs and trees between 12,800 and 11,300 cal yr B.P., indicating sustained warming during the Younger Dryas Chronozone (YDC). Stable isotopes suggest increasing atmospheric humidity from 11,800 to 9300 cal yr B.P. Mild (annual temperatures at least 1°C higher than present), and moist conditions prevailed from 11,000 to 10,350 cal yr B.P. Cooler, more variable conditions followed, and podocarp forest was completely replaced by montane Nothofagus forest at around 7500 cal yr B.P. with the onset of the modern climate regime. The Cass Basin late-glacial climate record closely matches the Antarctic ice core records and is in approximate antiphase with the North Atlantic.     

Paleoecological evidence for abrupt cold reversals during peak Holocene warmth on Baffin Island, Arctic Canada

A continuous record of insect (Chironomidae) remains preserved in lake sediments is used to infer temperature changes at a small lake in Arctic Canada through the Holocene. Early Holocene summers at the study site were characterized by more thermophilous assemblages and warmer inferred temperatures than today, presumably in response to the positive anomaly in Northern Hemisphere summer insolation. Peak early Holocene warmth was interrupted by two cold reversals between 9.5 and 8 cal ka BP, during which multiple cold-stenothermous chironomid taxa appeared in the lake. The earlier reversal appears to correlate with widespread climate anomalies around 9.2 cal ka BP; the age of the younger reversal is equivocal but it may correlate with the 8.2 cal ka BP cold event documented elsewhere. Widespread, abrupt climate shifts in the early Holocene illustrate the susceptibility of the climate system to perturbations, even during periods of enhanced warmth in the Northern Hemisphere.     

长江下游巢湖9870cal.aB.P.以来孢粉记录的环境演变

根据对巢湖490cm湖泊沉积物柱样7个AMS¹⁴C年龄的测定以及98个孢粉样品的分析,植物种类分属于86个（科）属,可以划分为6个孢粉组合带:孢粉带Ⅰ（9870～6040cal.aB.P.）代表了末次冰期之后全新世温暖气候到来之前气候转暖的早全新世过渡时期,气候呈现温和略干的特点,其中亚带Ⅰ-1（9870～7700cal.aB.P.）,Ⅰ-2（7700～6250cal.aB.P.）和Ⅰ-3（6250～6040cal.aB.P.）分别对应温暖湿润→温暖较湿→温和干燥的气候波动;孢粉带Ⅱ（6040～4860cal.aB.P.）代表中全新世温暖湿润期,水热配置条件最佳;孢粉带Ⅲ（4860～2170cal.aB.P.）体现中全新世后期温和干燥的气候,约2170cal.aB.P.干旱程度达到最高;孢粉带Ⅳ（2170～1040cal.aB.P.）反映巢湖流域由干燥向湿润气候的转型,气候总体上温和湿润;孢粉带Ⅴ（1040～200cal.aB.P.）反映了晚全新世巢湖流域温凉稍湿的气候;孢粉带Ⅵ（200cal.aB.P.至今）则体现巢湖流域处在相对温暖湿润的时期。植被类型演替大体为:以壳斗科的落叶、常绿属种为主的落叶阔叶、常绿阔叶混交林→以落叶栎类、栗属、青冈属和栲/石栎属为主的落叶阔叶、常绿阔叶混交林→以落叶栎类占绝对优势的落叶阔叶、常绿阔叶混交林→以禾本科为主的草地→以禾本科、蒿属和蓼属等为主的草丛。     

神农架大九湖15.753kaB.P.以来的孢粉记录和环境演变*

根据对大九湖297cm厚的泥炭地层10个AMS ¹⁴ C年龄的测定以及148块孢粉样品的分析,发现每块样品的孢粉(浓度)含量都很高,植物种类繁多,分属于137(科)属,可划分为6个孢粉组合带,植被类型演替大体为:含少量常绿树种的针阔叶混交林—含常绿阔叶树种的落叶阔叶林—常绿落叶阔叶林—含常绿阔叶树种的落叶阔叶林—含少量常绿树种的针阔叶混交林。孢粉带Ⅰ(15.753~11.280kaB.P.)记录了该区域晚冰期及其向全新世过渡时期冷暖波动频繁的气候条件,其中亚带Ⅰ -1(15.753~15.549kaB.P.),Ⅰ -3(14.936~14.505kaB.P.)和Ⅰ -5(12.655~11.280kaB.P.)可能分别对应最老仙女木期、老仙女木和新仙女木冷期,而亚带Ⅰ -2(15.549~14.936kaB.P.)和Ⅰ -4(14.505~12.655kaB.P.)分别对应博令和阿勒罗德暖期。孢粉带Ⅱ(11.280~9.218kaB.P.)代表了由晚冰期冷湿气候向全新世温暖气候转换的早全新世缓慢升温的过渡时期。孢粉带Ⅲ(9.218~7.530kaB.P.)代表中全新世前期的气温波动上升期,7.7kaB.P.前后达到最高温。孢粉带Ⅳ(7.530~4.051kaB.P.)代表中全新世适宜期,水热配置条件最佳。孢粉带Ⅴ(4.051~0.911kaB.P.)体现晚全新世温干的气候。孢粉带Ⅵ(0.911~0kaB.P)反映了大九湖温凉稍湿的气候。     

Late Wisconsin vegetational and climatic history at Kylen Lake, northeastern Minnesota

Kylen Lake, located within the Toimi drumlin field, is critically positioned in relation to Late Wisconsin glacial advances, for it lies between the areas covered by the Superior and St. Louis glacial lobes between 12,000 and 16,000 yr B.P. The pollen and plant-macrofossil record suggests the presence of open species-rich “tundra barrens” from 13,600 to 15,850 yr B.P. Small changes in percentages of Artemisia pollen between 14,300 and 13,600 yr B.P. appear to be artifacts of pollen-percentage data. Shrub-tundra with dwarf birch, willow, and Rhododendron lapponicum developed between 13,600 and 12,000 yr B.P. Black and white spruce and tamarack then expanded to form a vegetation not dissimilar to that of the modern forest-tundra ecotone of northern Canada. At 10,700 B.P. spruce and jack pine increased to form a mosaic dominated by jack pine and white spruce on dry sites and black spruce, tamarack, and deciduous trees such as elm and ash on moist fertile sites. At 9250 yr B.P. red pine and paper birch became dominant to form a vegetation that may have resembled the dry northern forests of Wisconsin today. The diagram terminates at 8410 ± 85 yr B.P. Climatic interpretation of this vegetational succession suggests a progressive increase in temperature since 14,300 yr B.P. This unidirectional trend in climate contrasts with the glacial history of the area. Hypotheses are presented to explain this lack of correspondence between pollen stratigraphy and glacial history. The preferred hypothesis is that the ice-margin fluctuations were controlled primarily by changes in winter snow accumulation in the source area of the glacier, whereas the vegetation and hence the pollen stratigraphy were controlled by climatic changes in front of the ice margin.     

The response of vegetation at the end of the last glacial period (MIS 3 and MIS 2) in littoral areas of NW Iberia

This paper describes and discusses the palaeobotanical data obtained from organic levels of two exposed deposits on the Atlantic shore of the northwestern Iberian Peninsula. Radiocarbon dating assigns these levels to a period of marine regression at the end of MIS 3 and the beginning of the Last Glacial Maximum. The pollen record shows an initial predominance of tree taxa (mainly deciduous, including the presence of Fagus far from its current limit), followed by an episode of partial forest retraction related to the end of MIS 3 and the beginning of the Last Glacial Maximum. However, the presence of numerous tree taxa in the record, even during the cold intervals associated with Heinrich events, points to the existence of sheltered refugia for these species during this period, in keeping with the conclusions of recent reviews of the vegetation dynamics of this region for the Lateglacial and the Holocene.     

A 35,000 Year Vegetation and Climate History from Potato Lake, Mogollon Rim, Arizona

A new record from Potato Lake, central Arizona, details vegetation and climate changes since the mid-Wisconsin for the southern Colorado Plateau. Recovery of a longer record, discrimination of pine pollen to species groups, and identification of macrofossil remains extend Whiteside's (1965) original study. During the mid-Wisconsin (ca. 35,000-21,000 yr B.P.) a mixed forest of Engelmann spruce (Picea engelmannii) and other conifers grew at the site, suggesting a minimum elevational vegetation depression of ca. 460 m. Summer temperatures were as much as 5°C cooler than today. During the late Wisconsin (ca. 21,000-10,400 yr B.P.), even-cooler temperatures (7°C colder than today; ca. 800 m depression) allowed Engelmann spruce alone to predominate. Warming by ca. 10,400 yr B.P. led to the establishment of the modern ponderosa pine (Pinus ponderosa) forest. Thus, the mid-Wisconsin was not warm enough to support ponderosa pine forests in regions where the species predominates today. Climatic estimates presented here are consistent with other lines of evidence suggesting a cool and/or wet mid-Wisconsin, and a cold and/or wet late-Wisconsin climate for much of the Southwest. Potato Lake was almost completely dry during the mid-Holocene, but lake levels increased to near modern conditions by ca. 3000 yr B.P.     

Diatom responses to limnological and climatic changes at Ribains Maar (French Massif Central) during the Eemian and Early Würm

High-resolution diatom analysis was carried out to assess the limnological and climatic changes that took place at Ribains maar (French Massif Central) during the Late Pleistocene (∼131–∼105 ka BP), with a focus on the Eemian interglacial in particular. Numerical analyses were used to show that most of the variability in the fossil diatom assemblages was due to climate independently from the changes in the lake catchment vegetation (as represented by pollen data). Diatom-based quantitative reconstructions of the past limnological conditions, as well as a comprehensive literature review on the auto-ecological requirements for the principal diatom taxa, were used to interpret the record. An absolute time-scale for the sequence was derived by matching the major pollen shifts with the radiometrically dated changes in oxygen isotopes observed in Italian stalagmites. This study shows that at Ribains maar, the transition from the Riss (=Saalian) Glacial to the Eemian interglacial was marked by a gradual increase in the contribution of spring-blooming diatom species, indicating a longer growing season and milder winter/spring conditions at that time. A short cooling event interrupts this trend and may correspond to a stadial. At the start of the Eemian a peak in benthic taxa and the suppression of spring-blooming flora probably reflects the effects of deglaciation on the catchment. During the Eemian interglacial itself three main phases were distinguished within the diatom record. The first phase (∼8000 years in duration) was dominated by Stephanodiscus minutulus, which suggests that intense mixing in the water-column took place during spring. The pollen record was simultaneously dominated by Quercus and Corylus that typify this phase as the climatic optimum of the Eemian. The second phase, almost equal in duration to the first phase (∼7000 years), is generally dominated by Cyclotella taxa and suggests a less productive lake and much reduced period of spring mixing compared with the first phase. In the pollen diagram this corresponds to an interval dominated by Carpinus–Picea–Abies that indicates a cooler and wetter climate. The third and last phase of the Eemian, ∼2000 year long, saw the return to Stephanodiscus-dominated assemblages, indicating a warming that may correspond to the Dansgaard–Oeschger event 25 identified in the Greenland ice-core record. In the early stage of the Würm Glacial (=Weichselian), assemblages in the Melisey I stadial (∼3000 year long) were dominated by either Aulacoseira subarctica or Asterionella formosa, which suggest colder spring conditions than during the late Eemian, but not as cold as the ones indicated by the pollen record. Stephanodiscus spp. again dominate during the Saint-Germain Ia interstadial (∼5000 year long) suggesting a return to the conditions that prevailed before the Melisey stadial, in agreement with the pollen record. The record ends with the Montaigu cold event, which is characterised by a Pinus peak in the pollen record, and corresponds to a large abundance of A. subarctica in the diatom sequence. Throughout the Eemian the abundance of Stephanodiscus spp., which is thought to be driven by winter conditions, show cyclic fluctuations that most likely match the cooling events identified in a pollen record from Germany. Variation in insolation throughout the Eemian may have been the driving factor behind the species succession observed in the diatom sequence. While this study demonstrates that diatom analysis of lake sediment can provide very detailed information on long-term climate change, a review of the few other diatom investigations published on European Eemian deposits shows that this technique has been so far seldom used to its full potential in this context in central and southern Europe.     

Late-glacial vegetation associated with caribou and mastodon in central Indiana

The Christensen Mastodon Site, located in central Indiana, contains a rich assemblage of vertebrates (including mastodon, caribou, and giant beaver), invertebrates, and plant macrofossils in situ in lake and bog sediments of late-glacial age. Studies of pollen and plant macrofossils suggest the existence of open, white spruce-dominated boreal forests from >; 14,000 yr B.P. to ca. 13,000 yr B.P. The regional decline of spruce, local occurrence of black spruce, white spruce, and larch, immigration of many hardwood taxa (e.g., ash, oak, elm), and the initiation of bog development are recorded beginning about 13,000 yr B.P. Recent reconstructions of late-glacial and early postglacial vegetational changes provide a context for understanding the disappearance of mastodons. The dramatic and rapid restriction of boreal forests along the retreating ice margin from 11,000 to 9000 yr B.P. may have caused a substantial reduction of mastodon populations. A diminished population would be more susceptible to small-scale, stochastic events such as short-term extremes of weather, outbreaks of disease, or predation pressure from paleoindian hunters.     

Multiproxy evidence for terrestrial and aquatic ecosystem responses during the 8.2 ka cold event as recorded at Højby Sø, Denmark

A sediment succession from Højby Sø, a lake in eastern Denmark, covering the time period 9400–7400 cal yr BP was studied using high-resolution geochemistry, magnetic susceptibility, pollen, macrofossil, diatom, and algal pigment analysis to investigate responses of the terrestrial and aquatic ecosystems to the 8.2 ka cold event. A reduced pollen production by thermophilous deciduous tree taxa in the period c. 8250–8000 cal yr BP reveal that the forest ecosystem was affected by low temperatures during the summer and winter/early-spring seasons. This finding is consistent with the timing of the 8.2 ka cold event as registered in the Greenland ice cores. At Højby Sø, the climate anomaly appears to have started 200–250 yr earlier than the 8.2 ka cold event as the lake proxy data provide strong evidence for a precipitation-induced distinct increase in catchment soil erosion beginning around 8500 cal yr BP. Alteration of the terrestrial environment then resulted in a major aquatic ecosystem change with nutrient enrichment of the lake and enhanced productivity, which lasted until c. 7900 cal yr BP.     

Vegetation and climate changes in northwestern Russia during the Lateglacial and Holocene inferred from the Lake Ladoga pollen record

The new pollen record from the upper 12.75 m of a sediment core obtained in Lake Ladoga documents regional vegetation and climate changes in northwestern Russia over the last 13.9 cal. ka. The Lateglacial chronostratigraphy is based on varve chronology, while the Holocene stratigraphy is based on AMS ¹⁴C and OSL dates, supported by comparison with regional pollen records. During the Lateglacial (c. 13.9–11.2 cal. ka BP), the Lake Ladoga region experienced several climatic fluctuations as reflected in vegetation changes. Shrub and grass communities dominated between c. 13.9 and 13.2 cal. ka BP. The increase in Picea pollen at c. 13.2 cal. ka BP probably reflects the appearance of spruce in the southern Ladoga region at the beginning of the Allerød interstadial. After c. 12.6 cal. ka BP, the Younger Dryas cooling caused a significant decrease in spruce and increase in Artemisia with other herbs, indicative of tundra‐ and steppe‐like vegetation. A sharp transition from tundra‐steppe habitats to sparse birch forests characterizes the onset of Holocene warming c. 11.2 cal. ka BP. Pine forests dominated in the region from c. 9.0 to 8.1 cal. ka BP. The most favourable climatic conditions for deciduous broad‐leaved taxa existed between c. 8.1 and 5.5 cal. ka BP. Alder experiences an abrupt increase in the local vegetation c. 7.8 cal. ka BP. The decrease in tree pollen taxa (especially Picea) and the increase in herbs (mainly Poaceae) probably reflect human activity during the last 2.2 cal. ka. Pine forests have dominated the region since that time. Secale and other Cerealia pollen as well as ruderal herbs are permanently recorded since c. 0.8 cal. ka BP.