二叠系-三叠系界线全球层型剖面——回顾和进展

综合评述了5年来对二叠系—三叠系界线(PTB)全球层型剖面——煤山D剖面的研究进展。简要介绍剖面的研究历史、地理和地质背景,叙述了世界上最完整的二叠纪长兴期至三叠纪格里斯巴赫期的牙形石和菊石带,并简短讨论了Otoceras的年代。叙述了煤山与其他剖面长兴期至格里斯巴赫期磁性地层学的对比。介绍了δ13C和δ34S的化学地层学最新研究进展,特别是在PTB附近多个δ13C负漂移的发现。两项重要的分子地层学研究显示了基础生态系统的深刻变化——微生物生态系的灾变和在二叠纪—三叠纪生态系之交严重缺氧事件期间透光带的静海相环境。文章报道了自2001年以来PTB年代测定的进展。PTB之下14cm处火山成因的粘土层25层的年代,现在定为(252·4±0·3)Ma(Mundil等,2004)。文章还讨论了事件地层学,认为尽管在25-26层有一个主事件,煤山PTB附近的事件是多幕式的,并且开始于事件层25-26层之前。因此,PTB大绝灭主因是外星撞击导致的可能性是不大的。     

Study on a Terrestrial Permian—Triassic Boundary Section—Zhejue Section, Weining County,Guizhou Province, China

The Zhejue Section is an excellent terrestrial permian-Triassic boundary (TPTB) section not only for its convenient accessibility and good outcrop, but also for its abundant fossils. In addition, there are two claystone beds that were formed by events across the Permian-Triassic boundary (PTB) at this section. The claystones provide a bridge for high-resolution correlation between marine and terrestrial PTBs. Another advantage is that the PTB at the Zhejue Section can be correlated with that of littoral and neritic facies by tracing estwards. Synthesized stratigraphic studies, including biostratigraphy, eventostratigraphy )susceptibilities and carbon isotopes across the TPTB, and microspherules found in the boundary claystones) and sequence stratigraphy, are carried out at this section.     

地层单位与全球界线层型:概念、术语、有关规定与研究实例

全球二叠系-三叠系界线(PTB)层型及点("金钉子")定位我国浙江省长兴县煤山D剖面,极大地推动了PTB及二叠系和三叠系研究进展。在简要介绍地层单位与全球界线层型与点建立的相关概念、术语和有关规定、地层划分的主要种类及其地层单位术语的基础上,以全球二叠系—三叠系界线层型及点(中国浙江省长兴县煤山D剖面)为研究实例,综述了该金钉子剖面牙形石生物地层和年代地层研究现状。煤山D剖面构建了目前全球最完整的PTB地层牙形石带序列,为研究二叠纪末大灭绝后生物残存和复苏过程提供了重要的时间坐标系,也为相关的地质事件研究奠定了坚实的基础。     

Ophiceras(蛇菊石)在滇东卡以头组的发现及其意义

Ophiceras(蛇菊石)在滇东海陆交互相卡以头组下部(25层下部)的首次发现,是近期海陆交互相和陆相二叠系-三叠系界线(PTB)研究的重要进展之一。该化石的发现为滇东卡以头组地层时代的确定提供了更加确凿的依据。在煤山D剖面,29-37层为Ophiceras(蛇菊石)的顶峰带,因此,可将滇东密德B剖面的25层与煤山D剖面29层对比,那么滇东地区卡以头组主体(甚至全部)的时代可划归早三叠世。     

南盘江盆地北部二叠纪末放射虫动物群与古水深

古-中生代之交,不同古地理背景中生物灭绝的表现不同.有关放射虫动物群研究,一方面,提供古地理背景等重要信息;另一方面,为生物多样性演变的研究提供基础资料.二叠纪-三叠纪之交盆地相放射虫的研究对揭示不同沉积相生物灭绝和古环境演变具有重要意义.贵州惠水以南克脚二叠系-三叠系界线剖面位于南盘江盆地北部,为盆地相沉积.对克脚剖面大隆组和大冶组底部放射虫动物群进行了初步统计、类群组合特征分析,结合放射虫的保存状态,对克脚剖面二叠纪末古水深进行了详细研究,认为大隆组至大冶组底部地层沉积于浪基面以下,古水深普遍浅于150 m,大隆组上部最大海侵时古水深最浅达200 m左右.      

湖北崇阳二叠纪—三叠纪之交生物灭绝和沉积微相演化

二叠纪—三叠纪之交,湖北崇阳地区处于浅水碳酸盐岩台地环境.二叠纪末的全球事件在该剖面的沉积微相和生物演化上均留下了清楚的记录.二叠纪末生物大灭绝之前,崇阳地区为典型的正常浅海台地环境,生物种类多样,数量丰富,主要生物化石有钙藻、有孔虫、腕足、棘皮类和海绵等.生物大灭绝之后,钙藻、(筳)类、棘皮类、海绵、绝大部分有孔虫开始消失,取而代之的是个体微小的腹足、介形虫和大量的蓝细菌化石.大灭绝界线之上,首先出现的是25 cm厚的纹层状的微生物岩,含较丰富的种类单调的有孔虫化石.之后逐渐相变为花斑状微生物岩和穹隆状微生物岩,厚度分别为6.4,2.3m.不同类型微生物岩在结构构造和生物组成上存在差别.微生物岩沉积结束之后,则相变为浅滩相鲕粒灰岩.共划分出3种沉积相,即开阔台地相、潮坪相和浅滩相.崇阳剖面的生物灭绝和沉积微相变化是二叠纪—三叠纪之交浅水台地环境生物与环境过程的典型代表,为认识二叠纪末浅海沉积相演化和全球事件提供了新材料.     

二叠纪-三叠纪之交全球海平面变化研究

二叠纪-三叠纪界线附近的全球海平面变化是当前沉积学研究的热点和难点问题,其与当时的显生宙最大规模生物灭绝事件存在一定关联,具有重要的研究意义。然而二叠纪-三叠纪界线附近的全球海平面变化存在较多争议,受单剖面或区域范围内相对海平面变化研究程度的制约,在缺乏从沉积学角度的综合对比研究的情况下,可能会影响对全球海平面变化过程与持续时间的判识。综述了二叠纪-三叠纪界线附近的海平面变化研究进展,整合了多位学者的研究剖面、主要观点及认识,梳理了全球海平面变化的主要观点（“上升论”和“下降-上升论”）,包括其各自的发展历程、代表剖面及海平面变化识别特征、海平面上升/下降的原因以及海平面变化与生物灭绝的关系等,并在此基础上,探讨了二叠纪-三叠纪全球海平面变化研究过程中产生争议的原因。本文旨在为二叠纪-三叠纪界线（PTB）附近海平面变化研究提供线索,同时为研究全球PTB地质事件发生的背景及差异性原因提供基础证据。     

湖北黄石二门二叠系/三叠系界线剖面长兴阶牙形刺分带及有机碳同位素的变化

湖北黄石二门二叠系/三叠系界线剖面出露完好,原始沉积连续,由乐平统龙潭组保安段灰黑色硅质岩、大隆组黑色硅质岩—硅质泥岩和下三叠统大冶组黑灰色泥岩以及泥岩夹灰岩组成,为介于典型浅水碳酸盐岩型与深海硅质岩型二叠系/三叠系界线剖面之间的半深海剖面。长兴阶可划分为Clarkinasubcarinata—Clarkinawangi及Clarkinachangxingensis两个牙形刺带。后者又可划分为3个亚带,自下而上依次为Clarkinachangxingensischangxingensis—Clarkinadeflecta亚带、Clarkinameishanensis亚带及Waning—Clarkina亚带,完全可以与二叠系/三叠系界线的全球层型剖面与点进行对比。另外,二叠纪末牙形刺的生态演化呈阶段性绝灭模式;有机碳同位素值在二叠纪/三叠系界线处出现明显的负偏移,指示了二叠纪/三叠纪之交生物绝灭后海水表层水原始产率的降低以及大气和海水中CO2含量的增加。     

东秦岭陕西聂家沟剖面二叠纪—三叠纪之交沉积特征及其古环境意义*

东秦岭陕西镇安聂家沟地区二叠纪—三叠纪之交发育一套海相碳酸盐岩沉积序列,较完整地记录了二叠纪末生物大灭绝事件前后浅海碳酸盐岩台地相生物群落演替及环境变化等信息,是研究二叠纪末生物大灭绝事件前后生态环境变化与沉积响应的理想区域。通过碳酸盐岩微相分析方法,对陕西镇安聂家沟剖面二叠系—三叠系界线附近的碳酸盐岩生物组合和微相类型进行了详细分析,共识别出11个微相类型: 斑点状凝块石、叠层石灰岩、鲕粒—纹层状叠层石灰岩、有孔虫颗粒灰岩、藻—海百合泥粒灰岩、含有被包壳和被磨蚀骨屑颗粒的粒泥灰岩、鲕粒颗粒灰岩、集合颗粒灰岩、含鲕粒的粒泥灰岩、泥晶灰岩和泥岩。根据碳酸盐岩微相特征及沉积相标志,在二叠系—三叠系界限附近划分出台地边缘、开阔台地和局限台地3种沉积相,其反映了二叠纪—三叠纪之交频繁的沉积相带变更的特点。该剖面碳酸盐岩微相反映的古生态和生物群落演替特征与中国华南同时期其他剖面具有较好的一致性,即二叠纪末生物大灭绝事件之后,早三叠世正常浅水碳酸盐岩台地生物类型和丰度极低,仅含有少量的双壳类、海百合等,灭绝事件界线附近以微生物碳酸盐岩等特殊微生物沉积构造占主导,之后微生物岩等消失,又出现了以双壳类为首的生物碎屑石灰岩。研究结果为了解二叠纪末生物大灭绝事件中生物和环境的协同演化提供了新的材料和认识。     

川南先锋地区二叠系-三叠系界线附近碳酸盐微相及沉积演化

根据川南先锋地区二叠系-三叠系界线地层的沉积特征,结合钻孔岩相及其组合特征,识别出等斜碳酸盐缓坡和局限台地2种沉积相,包括高能生屑浅滩、内缓坡、潮间带等7种亚相。同时采用微相分析的方法,对碳酸盐岩的生物组合、颗粒成分以及支撑类型等微相特征进行了研究,将界线地层沉积归纳为14种微相类型,并由此建立了该时期的沉积相序及其演化。研究区产出一层微生物岩,显微镜观察显示:微生物岩主要是叠层石;在它们内部发现了大量的蓝藻球粒、钙化鞘和微亮晶包覆层等微生物成因的显微组构。对ZK1501孔所采集的0.58 m K9灰岩岩芯连续磨片后进行生物组合分析,在距底6 cm地层中发现了丰富的三叶虫、腹足类和叶枝藻化石,其上出现小型化的介形虫(小于200 μm),表明二叠纪末期生物主灭绝界线应位于飞仙关组底面之上6 cm处,而二叠系-三叠系界线应位于小型化介形虫层的中下部。     

东秦岭陕西聂家沟剖面二叠纪—三叠纪之交沉积特征及其古环境意义*

东秦岭陕西镇安聂家沟地区二叠纪—三叠纪之交发育一套海相碳酸盐岩沉积序列,较完整地记录了二叠纪末生物大灭绝事件前后浅海碳酸盐岩台地相生物群落演替及环境变化等信息,是研究二叠纪末生物大灭绝事件前后生态环境变化与沉积响应的理想区域。通过碳酸盐岩微相分析方法,对陕西镇安聂家沟剖面二叠系—三叠系界线附近的碳酸盐岩生物组合和微相类型进行了详细分析,共识别出11个微相类型: 斑点状凝块石、叠层石灰岩、鲕粒—纹层状叠层石灰岩、有孔虫颗粒灰岩、藻—海百合泥粒灰岩、含有被包壳和被磨蚀骨屑颗粒的粒泥灰岩、鲕粒颗粒灰岩、集合颗粒灰岩、含鲕粒的粒泥灰岩、泥晶灰岩和泥岩。根据碳酸盐岩微相特征及沉积相标志,在二叠系—三叠系界限附近划分出台地边缘、开阔台地和局限台地3种沉积相,其反映了二叠纪—三叠纪之交频繁的沉积相带变更的特点。该剖面碳酸盐岩微相反映的古生态和生物群落演替特征与中国华南同时期其他剖面具有较好的一致性,即二叠纪末生物大灭绝事件之后,早三叠世正常浅水碳酸盐岩台地生物类型和丰度极低,仅含有少量的双壳类、海百合等,灭绝事件界线附近以微生物碳酸盐岩等特殊微生物沉积构造占主导,之后微生物岩等消失,又出现了以双壳类为首的生物碎屑石灰岩。研究结果为了解二叠纪末生物大灭绝事件中生物和环境的协同演化提供了新的材料和认识。     

The prelude of the end-Permian mass extinction predates a postulated bolide impact

The mass extinction at the Permian–Triassic Boundary (PTB) is said to have been abrupt and probably caused by an extraterrestrial impact. However, evidence from the Global Stratotype Section and Point (GSSP) of the base of the Induan at Meishan, China, shows that the biotic crisis began prior to the level, in beds 25 and 26 at which the postulated impact event occurred. Evidence of such an earlier biotic crisis occurs in other sections in South China, and in central and western Tethyan regions. This event is characterized by the extinction of a range of faunas, including corals, deep-water radiolarians, most fusulinids and pseudotirolitid ammonoids, and many Permian brachiopods. In all sections, this extinction level is usually a few decimeters to meters below that of the main mass extinction in the event beds (25 and 26) at Meishan, and their correlatives elsewhere. This earlier extinction event happened before the postulated bolide impact at the level of beds 25 and 26, and constrains interpretation of the mechanisms that brought about this greatest mass extinction. Electronic supplementary material Supplementary material is available in the online version of this article at and is accessible for authorized users.     

Aproposed Area for Study of Accessory Section and Point of Terrestrial Permian—Triassic Boundary

After the establishment of the global stratotype section and point (GSSP) of the Permian-Triassic boundary (PTB), the definition of the accessory section and point (ASP) of the terrestrial Permian-Triassic boundary (TPTB) is now on the agenda. However, all good TPTB sections so far known have the following shortcoming:(1) the exact TPTB horizon is difficult to define paleontologically with high-resolution, and (2) accurate correlation between marine and terrestrial PTBs in hard to attain. In order to enhance the understanding of the nature of the global life crisis in both the marine and terrestrial environments across the Paleozoic-Mesozoic transition, these shortcomings need to be addressed. In western Guizhou and eastern Yunnan, Southwest China, some fossiliferous PTB sections which include marine, paralic and terrestrial are well-developed, allowing bed-to-bed correlation of the PTB sequences. Fortunately, the marine PTB sequence in this area is almost the same as found at the Meishan Section, where the GSSP of the PTB is located, which may provide a reliable auxiliary marker for high-resolution demarcation of the TPTB. These features fund in estern Guizhou and eastern Yunnan make this area a good place to study the ASP of the TPTB, so we propose to study the ASP of the TPTB in this area.     

黔西威宁岔河陆相二叠系-三叠系界线粘土岩研究--基于X射线衍射、扫描电镜分析

利用X射线衍射、扫描电镜等分析方法,对黔西威宁岔河陆相剖面二叠系-三叠系界线附近的粘土岩进行了研究.在界线粘土岩中,粘土矿物组合为伊-蒙混层矿物 蒙脱石 绿泥石;碎屑物质中有六方双锥石英和锆石,无球粒;多种化学成分在界线粘土岩中出现了异常.据此得出岔河剖面的二叠系-三叠系事件地层界线确定在第66f层比较合适;第66f和68a这两层粘土岩为此剖面的界线粘土岩;自第66f层底部至第68a层顶部这一段地层为二叠系-三叠系的界线层(组);界线粘土岩的成因可能是由于火山喷发、火山灰降落伴随正常沉积物沉积形成的.     

A Review on the Global Stratotype Section and Point of the Permian-Triassic Boundary

1 Introduction The Global Stratotype Section and Point (GSSP) of the Permian-Triassic Boundary (PTB) has been ratified by IUGS in 2001 (Yin et al., 2001). It is defined at the base of the Hindeodus parvus horizon, i.e. the base of Bed 27c of Meishan Section D, Changxing County, Zhejiang Province, South China. The PTB is important because it is not only an erathem boundary but also a great turning point of geological history symbolized by profound global changes and the strongest …     

Evidence for wildfire in the Meishan section and implications for Permian-Triassic events

Polynuclear aromatic hydrocarbons (PAHs) and black carbon (BC) in sediments are powerful tools in the identification of the combustion process throughout geologic history. In this study, combustion-derived PAHs and BC were carefully investigated in sediments from the Global Stratotype Section and Point of the Permian-Triassic (P/Tr) boundary in Meishan, China. Quantitative analyses of combustion-derived PAHs and BC demonstrate anomalously high concentrations in the boundary event beds that coincide with the mass extinction horizon. The prevalence of parent polynuclear aromatics (e.g., phenanthrene) in PAHs, together with non-metric multidimensional scaling analysis, confirms that the PAHs are mainly derived from vegetation burning instead of having a coal and/or oil origin. BC detected in sediment occurs in various forms from large irregular charcoal particles to fine aciniform soot, with an equivalent reflectance of up to 3.5%. The results strongly suggest that a wildfire occurred during the P/Tr boundary, which served as one of the possible triggers of mass extinction on land. The wildfire occurrence indicates that the O₂ concentration of the atmosphere during (or before) the P/Tr mass extinction was probably >17%. The temporal coincidence of the mass extinction with intensive volcanic eruption, marine anoxia and wildfire events in the region of the Meishan section provides new insight into the mechanisms of the P/Tr biotic crisis. Our results show that wildfires could have played an important role in the collapse of the ecosystem in the Meishan P/Tr events.     

贵州威宁金钟二叠系-三叠系界线剖面生物地层及其对比

二叠纪—三叠纪之交发生了显生宙最大的生物灭绝事件,同时发生了复杂的环境变化.由于陆相二叠系-三叠系界线附近生物化石记录没有海相记录完整,同时也缺乏其他可靠的年代地层标志,所以如何标定陆相剖面的二叠纪—三叠纪界线一直是研究热点和难点.贵州威宁金钟地区二叠纪—三叠纪之交位于康滇古陆的东侧,毗邻海陆过渡相区,可以与同期的浅海碎屑岩相、海陆过渡相、陆相剖面进行由海到陆“追索式”的生物地层对比,是我国研究海、陆相二叠系-三叠系界线及对比的关键研究区之一.研究区的金钟二叠系-三叠系界线剖面中化石丰富,共发现双壳类化石4属6种,叶肢介2属4种,植物化石9属及腕足类化石1属,为海、陆相地层对比提供了桥梁.依据叶肢介Euestheria gutta-Palaeolimnadia xuanweiensis组合、植物Annalepis-Peltaspermum组合以及双壳类Pteria ussurica variabilis-Neoschizodus orbicularis组合的生物化石时代归属讨论,识别出金钟剖面的二叠系-三叠系界线应该在卡以头组下部.通过与邻区不同相剖面的对比,提出双壳类Pteria ussurica variabilis-Neoschizodus orbicularis组合以及陆相的叶肢介Euestheria gutta-Palaeolimnadia xuanweiensis组合可以作为三叠纪最早期的识别和对比标志.      

Timing and magnitude of tetrapod extinctions across the Permo-Triassic boundary

A review of the tetrapod (amphibian and amniote) record across the Permo-Triassic boundary (PTB) indicates a global evolutionary turnover of tetrapods close to the PTB. There is also a within-Guadalupian tetrapod extinction here called the dinocephalian extinction event, probably of global extent. The dinocephalian extinction event is a late Wordian or early Capitanian extinction based on biostratigraphic data and magnetostratigraphy (the extinction precedes the Illawara reversal), so it is not synchronous with the end-Guadalupian marine extinction. The Russian PTB section documents two tetrapod extinction events, one just before the dinocephalian extinction event and the other at the base of the Lystrosaurus assemblage. However, generic diversity across the latter extinction remains essentially the same despite a total evolutionary turnover of tetrapod genera. The Chinese and South African sections document the stratigraphic overlap of Dicynodon and Lystrosaurus. In the Karoo basin, the lowest occurrence of Lystrosaurus is in a stratigraphic interval of reversed magnetic polarity, which indicates it predates the marine-defined PTB, so, as previously suggested by some workers, the lowest occurrence of Lystrosaurus cannot be used to identify the PTB in nonmarine strata. Correlation of the marine PTB section at Meishan, southern China, to the Karoo basin based primarily on magnetostratigraphy indicates that the main marine extinction preceded the PTB tetrapod extinction event. The ecological severity of the PTB tetrapod extinction event has generally been overstated, and the major change in tetrapod assemblages that took place across the PTB was the prolonged and complex “replacement” of therapsids by archosaurs that began before the end of the Permian and was not complete until well into the Triassic. The tetrapod extinctions are not synchronous with the major marine extinctions at the end of the Guadalupian and just before the end of the Permian, so the idea of catastrophic causes of synchronous PTB extinctions on land and sea should be reconsidered.     

Timing and magnitude of tetrapod extinctions across the Permo-Triassic boundary

A review of the tetrapod (amphibian and amniote) record across the Permo-Triassic boundary (PTB) indicates a global evolutionary turnover of tetrapods close to the PTB. There is also a within-Guadalupian tetrapod extinction here called the dinocephalian extinction event, probably of global extent. The dinocephalian extinction event is a late Wordian or early Capitanian extinction based on biostratigraphic data and magnetostratigraphy (the extinction precedes the Illawara reversal), so it is not synchronous with the end-Guadalupian marine extinction. The Russian PTB section documents two tetrapod extinction events, one just before the dinocephalian extinction event and the other at the base of the Lystrosaurus assemblage. However, generic diversity across the latter extinction remains essentially the same despite a total evolutionary turnover of tetrapod genera. The Chinese and South African sections document the stratigraphic overlap of Dicynodon and Lystrosaurus. In the Karoo basin, the lowest occurrence of Lystrosaurus is in a stratigraphic interval of reversed magnetic polarity, which indicates it predates the marine-defined PTB, so, as previously suggested by some workers, the lowest occurrence of Lystrosaurus cannot be used to identify the PTB in nonmarine strata. Correlation of the marine PTB section at Meishan, southern China, to the Karoo basin based primarily on magnetostratigraphy indicates that the main marine extinction preceded the PTB tetrapod extinction event. The ecological severity of the PTB tetrapod extinction event has generally been overstated, and the major change in tetrapod assemblages that took place across the PTB was the prolonged and complex “replacement” of therapsids by archosaurs that began before the end of the Permian and was not complete until well into the Triassic. The tetrapod extinctions are not synchronous with the major marine extinctions at the end of the Guadalupian and just before the end of the Permian, so the idea of catastrophic causes of synchronous PTB extinctions on land and sea should be reconsidered.     

Paleomagnetic Constraints on the Permian-Triassic Boundary in Terrestrial Strata of the Karoo Supergroup, South Africa: Implications for Causes of the End-Permian Extinction Event

The closure of the Palaeozoic witnessed the greatest biotic crisis in earth history. Surprisingly little is known about the effects and timing of the terrestrial counterpart of the well-described End-Permian mass extinction from known marine successions worldwide. In the present study, reliable paleomagnetic results were obtained from a PT boundary section in the terrestrial Karoo Basin of South Africa. Permo-Triassic aged mudstones from a locality in the Eastern Cape Province yielded two magnetic chrons, reverse followed by normal (with the boundary possibly close to the reversal). This extends to results from a previous study: thereby jointly identifying a R/N/R polarity pattern for this boundary interval. The PTB interval is constrained below the red mudstones of the Beaufort Group at the present locality and within reverse-magnetised green mudstone, implying a diachronic relation between the marine and terrestrial End-Permian mass extinction events.