A new gigantic lacewing species (Insecta: Neuroptera) from the Lower Cretaceous of Brazil confirms the occurrence of Kalligrammatidae in the Americas

Makarkinia kerneri sp. nov. is described from the Lower Cretaceous Crato Formation of Brazil, based on an incomplete but very well-preserved hind wing (Neuroptera: Kalligrammatidae). The previously presumed attribution of the genus is confirmed, based on the diagnostic characters of its wing venation (e.g., dense crossvenation; the anteriorly directed branches of MP) and the presence of a distinct eye-spot on the wing. Makarkinia is the only American representative and youngest known genus of Kalligrammatidae. With an estimated wing length of 100–160 mm, it has the largest wings amongst all fossil and extant Neuroptera.     

Pelecinid wasps (Insecta, Hymenoptera, Proctotrupoidea) from the Cretaceous of Russia and Mongolia

Thirteen new species referable to four genera, of which one is new, from the Cretaceous of Russia and Mongolia are established herein and assigned to the family Pelecinidae. Among the four genera, Protopelecinus gen. nov., including four new species, is referred to the subfamily Pelecininae, while Iscopinus Kozlov, including three new species, Eopelecinus Zhang, Rasnitsyn and Zhang, including five new species, and Scorpiopelecinus laetus sp. nov. are assigned to the subfamily Iscopininae. Of these new taxa, eight, namely Protopelecinus regularis, P. furtivus, Iscopinus simplex, ?I. suspectus, Eopelecinus exquisitus, E. scorpioideus, E. rudis and Scorpiopelecinus laetus, are from the Lower Cretaceous Zaza Formation of Baissa, Transbaikalia, Russia; two, E. minutus and E. fragilis, are from the basal Lower Cretaceous Tsagan-Tsab Formation of Khutel-Khara, Mongolia; two, P. dubius and P. deformis, are from the Lower Cretaceous (Aptian?) of Bon Tsagan, Mongolia; and one, I. separatus, is from the Upper Cretaceous (Cenomanian) Ola Formation of Obeshchayushchiy, Russia. A key to fossil pelecinid wasps is provided and a morphological analysis shows that the Pelecinidae might be paraphyletic with respect to the Proctotrupidae. The Chinese insect fauna from both the Yixian and Laiyang formations is dominated by Eopelecinus and Sinopelecinus whereas the Siberian + Mongolian fauna from the Zaza and Tsagan-Tsab formations is dominated by Eopelecinus and Iscopinus. Hence, Eopelecinus is common to both. The differences between the two faunas are likely to be the result of geographical variation in populations.     

The first fossil hilarimorphid fly (Diptera: Brachycera)

Although recent molecular phylogenetic analyses suggest a great antiquity, going into the Jurassic, for the small modern fly family Hilarimorphidae, no fossil was attributed to this group. The first fossil hilarimorphid Cretahilarimorpha lebanensis gen. et sp. nov., is described, based on a specimen from the Lower Cretaceous Lebanese amber. Its external morphology is analysed and compared with that of Hilarimorpha, unique modern hilarimorphid genus. The differences concern the wing venation, antenna and especially mouthparts. Cretahilarimorpha has very elongate mouthparts, adapted for nectar feeding or (less probably) for predation on other insects. Several other Lower Cretaceous lineages have developed similar elongate mouthparts, viz. nemestrinid and xylomyiid flies, and the Mecoptera: Aneuretopsychina, probably adapted to still unknown deep nectar-producing flowers. A checklist of species belonging to the Hilamorphidae is given.     

Consideration of the genus Ginkgoites Seward and a redescription of two species from the Lower Cretaceous of Germany

Genera used for fossil ginkgoalean leafy shoots, especially Baiera F. Braun,Ginkgoites Seward and Ginkgo L., are discussed both in their historical context and in the light of recent discoveries. Use of Ginkgoites Seward as a form- or organ-genus for fossil leaves lacking associated reproductive structures is readopted. Two species, Ginkgoitesbrauniana (Dunker) comb. nov. and Ginkgoites pluripartita (Schimper) Seward, from the Lower Cretaceous Wealden facies of Germany, are rediagnosed and neotypes selected. The occurrence of G. pluripartita in the Lower Cretaceous of western Greenland, Montana, U S A, Canada and Siberia is confirmed.     

Palaeoleptus burmanicus n. gen., n. sp., an Early Cretaceous shore bug (Hemiptera: Palaeoleptidae n. fam.) in Burmese amber

Palaeoleptus burmanicus gen. et sp. nov. (Hemiptera: Leptopodomorpha: Palaeoleptidae fam. nov.) is described from Early Cretaceous Burmese amber. The fossil is characterized by nearly completely coreaceous wings with only a small membranous area at the distal tip, and a unique wing venation consisting of eight closed cells, including four obliquely orientated toward the embolar wing margin and two large vertically-positioned cells extending nearly to the apical wing margin. Additional characters are large, compound eyes, three pairs of cephalic trichobothria, four antennal segments all similar in texture, spines on the profemur, ocelli positioned on a tubercle and a rostrum extending to the mesocoxae, with the second segment bearing four pairs of spines. The female fossil contains an asymmetrical subgenital plate orientated toward the left side of the body, indicating a side-by-side mating behavior as occurs in extant Leptopodomorpha.     

A new species of Cyrtophyllitinae (Insecta: Ensifera) from the Cretaceous China

A new fossil species of Cyrtophyllitinae, Vitimoilus ovatus sp. nov., is described from the Dabeigou Formation and Yixian Formation of the Lower Cretaceous of China. It is clearly placed in Vitimoilus Gorochov, 1996 according to the following characters: R forking into RA and RP distally, M forking into MA and MP distally; CuA fused with CuPaα distal of the middle of the wing length; cross-veins in basal part of CuPb–CuPaβ area non-strongly curved.     

A new genus of Trichomyiinae (Diptera: Psychodidae) from Upper Cretaceous amber of New Jersey

Xenotrichomyia newjerseyiensis a new genus and species belonging to Trichomyiinae within Psychodidae is characterized, described, illustrated and its taxonomic position is discussed. It shares some similarities with Trichomyiinae of group B, but retains some archaic features, such as normal and non-excentric flagellomeres, and possesses an autapomorphy in wing venation that distinguish it from all remaining Trichomyiinae. This discovery represents the first Trichomyiinae (Psychodidae) from the Upper Cretaceous New Jersey amber, and adds more evidence on the very high palaeodiversity of the psychodids since at least the Lower Cretaceous. An updated check list of fossil Trichomyiinae is given.     

New Aetheogrammatidae (Insecta: Neuroptera) from the Lower Cretaceous Yixian Formation,China

Two new genera and three new species of the Mesozoic family Aeotheogrammatidae are described from the Lower Cretaceous of Yixian Formation (Liaoning Province, China): Cyclicogramma rotundum gen. et sp. nov. and Aetheogramma bistriatum sp. nov. from Huangbanjigou, and Curtogramma ovatum gen. et sp. nov. from Dawangzhangzi. The hind wing of Cyclicogramma rotundum gen. et sp. nov. differs from that of other species by its smaller size, its overall shape, RP bearing only two oblique radial branches, and its configurations of CuP, AA1, and AA2. Aetheogramma bistriatum sp. nov. is closely related to A. speciosum Ren and Engel, 2008, but distinguished from the latter by its distinctive configuration of RP1, and the narrower and longer AA2 space in the hind wing. Curtogramma ovatum gen. et sp. nov. is distinguished from all other species of the family by the forewing with M forking very far from the wing base.     

New Evidence for a Cretaceous Age for a Mesozoic Nonmarine Bivalve Assemblage from Paekto-dong, Sinuiju City, The Democratic People’s Republic of Korea

The Sinuiju Formation in Paekto-dong, Sinuiju City in the Democratic People's Republic of Korea has yielded Mesozoic nonmarine bivalve fossils, which is the first occurrence of such in the DPRK. Based on these fossil specimens, a new Cretaceous bivalve assemblage, the Arguniella yanshanensis-Sphaerium anderssoni Assemblage is erected. This assemblage includes Arguniella yanshanensis, A. lingyuanensis and Sphaerium anderssoni and can be compared with the Jehol Biota. The age of the Sinuiju Formation is also clarified and on the basis of the bivalves and the presence of a Eosestheria–Ephemeropsis–Lycoptera(E–E–L) assemblage, the formation is not Upper Jurassic, but Lower Cretaceous in age.     

Diverse stigmaphronid wasps in Early Cretaceous amber from Spain (Hymenoptera: Ceraphronoidea: Stigmaphronidae)

A diverse fauna of wasps of the extinct parasitoid family Stigmaphronidae (Ceraphronoidea) are recorded in Early Cretaceous (Lower Albian) amber from Spain. Seven new species in five genera are described and figured based on 51 specimens, representing more material than in all the world’s other amber deposits combined. New species include: Elasmophron mari sp. nov., Libanophron sugaar sp. nov., Hippocoon basajauni sp. nov., Burmaphron jentilak sp. nov., B. sorginak sp. nov., B. iratxoak sp. nov., and Tagsmiphron olentzero sp. nov. The significance of the fauna is discussed and compared with that of other Cretaceous amber deposits, in particular the tremendous richness of the Spanish fauna is contrasted with the complete absence of stigmaphronids in the slightly younger and nearby French amber. Whether this stark difference represents particularly favorable conditions for these parasitoids, or their hosts, in the Cretaceous Spanish archipelago, or whether it is owing to taphonomic factors is discussed.     

A new genus and species of snipe fly (Diptera: Rhagionidae) in Lebanese Cretaceous amber

The rhagionid Lebanoleptis huangi gen. et sp. nov. is described from the Lower Cretaceous amber of Lebanon. It is characterized, illustrated, and compared with the recent and fossil genera having the same particular absence of wing vein M3. Other Mesozoic flies with similar venation and currently attributed to the Rhagionidae are briefly discussed and should be revised.     

Paleogeographic evidence on the Jurassic tectonic history of the Pontides: new paleomagnetic data from the Sakarya continent and Eastern Pontides

The Jurassic paleogeographic position of the Pontides is not well studied because of insufficient paleomagnetic data. For this reason, a paleomagnetic study was carried out in order to constrain the paleolatitudinal drift of the Turkish blocks during the Jurassic period. A total of 32 sites were sampled from volcanic and volcanoclastic rocks of the Lower/Middle Jurassic Kelkit formation (Eastern Pontides), Mudurnu formation (Sakarya continent) and Upper Jurassic–Lower Cretaceous Ferhatkaya formation exposed around Amasya region (Eastern Pontides). Rock magnetic experiments demonstrate that the main ferromagnetic mineral is pseudo-single-domain titanomagnetite in these rocks. Paleomagnetic analysis revealed two main components of the natural remanent magnetization during stepwise thermal and alternating field demagnetization. The first component is a low-coercivity (unblocking temperature) component with a direction sometimes similar to that of the earth’s present field or a viscous component. The second component, which is interpreted as the characteristic remanent magnetization (ChRM) direction, has low to high coercivity properties between 20 and 100 mT or unblocking temperatures between 300 and 580°C. A positive fold test at the 95% level of confidence proved that the ChRM of the sites is primary. Paleomagnetic directions calculated for the Kelkit formation in the Eastern Pontides have a mean direction of D = 334.8°, I = 49.7°, α ₉₅ = 7.1° after tilt-correction. A mean direction of D = 332.2°, I = 48.5°, α ₉₅ = 14.6° was obtained from the volcanoclastic rocks of the Mudurnu formation, and D = 324.3°, I = 43.3°, α ₉₅ = 9.5° was calculated for the Upper Jurassic–Lower Cretaceous limestones/Ferhatkaya formation of the Amasya region. The Jurassic rocks in the Eastern Pontides and Mudurnu region are considered to represent products of the rifted Neo-Tethys ocean, while the Upper Jurassic–Lower Cretaceous sediments in Amasya are related to basin-filling materials. The data suggest that the Kelkit formation was formed at 30.5°N paleolatitude and the equivalent Mudurnu formation at 29.5°N paleolatitude. The paleolatitude of the Eastern Pontides indicates that this rifting block was separated from Eurasia by a marginal basin instead of being a part of Eurasia. The lower paleolatitude of the Amasya region at 24.8°N in the Upper Jurassic to Lower Cretaceous clearly indicates southward drift of the Turkish blocks during the Jurassic to Lower Cretaceous period together with the motion of Eurasia.     

New genus and species of Cerophytidae (Insecta: Coleoptera) from the Lower Cretaceous of Baissa (Siberia)

One new genus and two new species of fossil Cerophytidae, Baissophytum convexus gen. et. sp. nov. and Baissophytum amplus sp. nov. are described and illustrated from Zaza Formation (Lower Cretaceous), Eravnensky (Bauntovsky) District, Buriatskay Autonomous Republic (Transbaikalia), Russia.     

翼龙类翼膜的折叠，褶皱以及物质特性

Preservation of wing membranes in pterosaurs is only reported from a few localities in the world, namely from the Karatau Hills, Kazakhstan (Upper Jurassic), from the Solnhofen Lithographic Limestone, Germany (Upper Jurassic), the Daohugou Beds of China (Upper Jurassic or Lower Cretaceous) and the Crato and Santana Formations, Chapada do Araraipe, NE Brazil (Lower Cretaceous).     

First false jewel beetle (Coleoptera: Schizopodidae) from the Lower Cretaceous of China

A new false jewel beetle, Mesoschizopus elegans n. gen. n. sp., is described and illustrated based on a well-preserved impression fossil from the Lower Cretaceous Yixian Formation of China. It represents a definitive Mesozoic fossil belonging to the recent small buprestoid family Schizopodidae, which is now endemic to the western North America. Mesoschizopus is placed in Schizopodidae based on its overall body shape and size, short prosternum in front of procoxae, wide metanepisternum, and wing venation. The discovery of the first schizopodid from the Early Cretaceous not only highlights the antiquity of the small family, but also provides an example of a now narrowly distributed group whose ancestral groups were probably more widespread in the Mesozoic.     

Taxonomy, facies relationships and palaeobiology of bakevelliid bivalves from the Lower Cretaceous of west-central Argentina

Cretaceous bakevelliids in Argentina are restricted to the Lower Cretaceous. They are recorded in shallow marine deposits in two Mesozoic basins located in the Andes foothills in Patagonia. In the Austral Basin (44°–55°S) there is only one genus, Gervillella, represented by a single specimen from the Barremian. In the Neuquén Basin (30°–40°S) two genera, Gervillaria and Gervillella, occur, ranging from the Berriasian to the Lower Barremian. In the Pilmatué Member of the Agrio Formation two species are identified: Gervillaria alatior (Imlay) and Gervillella aviculoides (J. Sowerby). The former is also recorded in Mexico while the latter is more widespread, occurring in Europe, Asia, and Africa. In the Bajada del Agrio section they come from eight tabular shell beds located near and on top of shallowing-upwards cycles. They were abundant only in four of these levels. These shell beds are interpreted as having been deposited in the mid-ramp (middle of cycles), and upper mid-ramp (top of cycles). A functional analysis of shells revealed two life habits: epibyssate and endobyssate.     

New Schizolepis Fossils from the Early Cretaceous in Inner Mongolia, China and its Phylogenetic Position

Three Schizolepis species collected from the Lower Cretaceous layer of the Huolinhe Basin,Inner Mongolia,China are described.These fossils are Schizolepis longipetiolus Xu XH et Sun BN sp.nov.,which is a new species,Schizolepis cf.heilongjiangensis Zheng et Zhang,and Schizolepis neimengensis Deng.The new species is a well-preserved female cone,slender and cylindrical in shape.The seed–scale complexes have long petioles and are arranged on the cone axis loosely and helically.The seed scales are divided into two lobes from the base.Each lobe is semicircular or elongate ligulate in shape,widest at the middle or the lower middle part,with an obtuse or bluntly pointed apex.The inner margin is almost straight and the outer margin is strongly arched.On the surface of the lobe,there are longitudinal and somewhat radial striations from the base to the margin.The seed is borne on the adaxial surface at the base or middle of each lobe.Schizolepis was established in 1847,and,although more than twenty species have been discovered and reported,its phylogenetic position is controversial because of the imperfection of fossils.Most authors have considered there to be a close evolutionary relationship between Schizolepis and extant Pinaceae.Here,we analyze characteristics and compare Schizolepis with Picea crassifolia Kom,which is morphologically most similar to Schizolepis.The results indicate that the genus probably has a distant evolutionary relationship with extant Pinaceae.A detailed statistical analysis of the global paleogeographic distribution of Schizolepis showed that all the fossils of this genus appeared in strata ranging from the Upper Triassic to the Lower Cretaceous in the North Hemisphere,being rare in the Upper Triassic and Lower Jurassic,but being very common from the Middle Jurassic to the Lower Cretaceous,and particularly abundant in the Lower Cretaceous.According to the statistical results,we speculate that the genus originated in Europe in the Late Triassic then spread from Europe to Asia between the Late Triassic and the Late Jurassic.In the Early Cretaceous most species existed in China’s three northeastern Provinces and the Inner Mongolia Autonomous Region and adjacent areas.Combining the paleogeographic distribution of the genus with ancient climatic factors,we deduced that Schizolepis began to decline and became extinct in the Early Cretaceous,and the reason for its extinction is closely related to the icehouse climate during the Early Cretaceous.     

Ascalochrysidae – a new lacewing family from the Mesozoic of China (Insecta: Neuroptera: Chrysopoidea)

Ascalochrysa megaptera gen. et sp. nov. is described from the Yixian Formation (Lower Cretaceous) of China. It is treated as belonging to the new family Ascalochrysidae fam. nov., closely related to Mesochrysopidae but easily distinguished from it in the hindwing venation features: e.g., numerous subcostal crossveins; branches of Rs widely, irregularly spaced and deeply branched; crossvenation rich, irregular; convex vein-like fold (‘M5’) before concave CuA present. The presence of well-developed ‘M5’ in the hind wing is interpreted to be plesiomorphic condition within the order. Ascalochrysidae are considered the sister group of Mesochrysopidae (s.l.).     

辽宁北部秀水盆地秀D1井孢粉组合及其地层意义

孢粉化石采自辽宁省北部秀水盆地的秀D1井,分析、鉴定和系统研究结果表明,来自秀D1井井深62.1~1089.7m的孢粉化石自下而上划分为3个孢粉组合,下部孢粉组合(井深703.85~1089.7m)以Osmundacidites-Klukisporites-Podocarpidites为代表,地质时代为晚侏罗世堤塘期(Tithonian),中部孢粉组合(井深381.5~699.3m)以Densoisporites-Cicatricosisporites-Piceaepollenites为代表,地质时代为早白垩世贝里阿斯期(Berriassian),上部孢粉组合(井深62.1~339m)以Cicatricosisporites-Impardecispora-Pinuspollenites为代表,地质时代为早白垩世凡兰吟期—欧特里夫期(Valanginian-Hauterivian)。秀D1井钻遇地层所含孢粉组合特征在区域上可以与冀北地区大北沟组,辽宁西部地区下白垩统义县组、九佛堂组,松辽盆地东南缘下白垩统火石岭组、沙河子组所产的孢粉组合对比。含孢粉组合地层时代的确定解决了井柱地层的划分,以及与区域地层的对比关系,同时为区域地层古生物研究提供了翔实的基础资料。