New records of coleoid cephalopod jaws from the Upper Cretaceous of Hokkaido,Japan, and their paleobiogeographic and evolutionary implications

Seven coleoid jaws recovered from Santonian to lower Campanian (Upper Cretaceous) strata in Hokkaido, Japan were taxonomically studied. Based on the comparison with the jaws of modern and fossil coleoids, six of the seven jaw fossils are referred to the following two genera and three species, including one possible new species: Nanaimoteuthis jeletzkyi and N. yokotai of the order Vampyromorpha, and Paleocirroteuthis sp. nov. (?) of the order Cirroctopodida. The other single lower jaw is seemingly similar to those of modern octopods and teuthids with respect to the shape of the inner lamella, but its order-level assignment could not be determined because of its imperfect preservation. N. jeletzkyi has been described in the Upper Cretaceous fore-arc basin deposits in Hokkaido (Yezo Group) and Vancouver Island, Canada (Nanaimo Group), whereas N. yokotai occurs only in the Yezo Group. These findings, complemented by previous reports of coleoid jaws, gladii, and phragmocones from the Yezo and Nanaimo Groups, demonstrate that a highly diversified, non-belemnitid coleoid fauna including large teuthids had already appeared during the post-Albian Late Cretaceous, in the North Pacific region.     

First record of a Cretaceous cheilostome bryozoan from Hokkaido, Japan

A small collection of recrystallised, encrusting colonies of a single species from the Mikasa Formation (lower Middle Cenomanian), represents the first record of cheilostome (malacostegan or anascan) bryozoans from Hokkaido, Japan.     

Integrated biostratigraphy and palaeoenvironmental interpretation of the Upper Cretaceous to Paleocene succession in the northern Moldavidian Domain (Eastern Carpathians,Romania)

This study of the upper Maastrichtian to Danian sedimentary succession from the northern part of the Romanian Eastern Carpathians (Varniţa section) aims to establish an integrated biostratigraphy based on calcareous nannofossils, organic-walled dinoflagellate cysts (dinocysts) and foraminiferal assemblages, and to reconstruct the depositional environments of the interval. The stratigraphic record across the studied section is incomplete, considering that an approximately 16 m thick strata interval from the top of the Maastrichtian to lowermost Danian cannot be analyzed due to a landslide covering the outcrop. The upper Maastrichtian is marked by a succession of biostratigraphic events, such as the First Appearance Datum (FAD) of the nannoplankton taxon Nephrolithus frequens and FAD of the dinocyst species Deflandrea galeata and Disphaerogena carposphaeropsis, and the Last Appearance Datum (LAD) of Isabelidinium cooksoniae in the lower part of the section. These bioevents are followed by the LAD of the Dinogymnium spp. and Palynodinium grallator dinocyst markers in the top of the Maastrichtian deposits analyzed. In terms of foraminiferal biostratigraphy, the upper Maastrichtian Abathomphalus mayaroensis Zone is documented in the lower part of the studied section. Some bioevents, such as the bloom of the calcareous dinoflagellate genus Thoracosphaera and the FAD of the organic-walled dinocysts Damassadinium californicum, Senoniasphaera inornata, Xenicodinium lubricum and X. reticulatum suggest an early Danian age for the middle part of the section. From the Danian deposits in the Varniţa section, we describe a new organic-walled dinocyst species, Pentadinium darmirae sp. nov., which is until now the only species of the Pentadinium genus discovered in the Paleocene. The occurrence of the global Danian dinocyst marker Senoniasphaera inornata in the top of the section, suggests an age not younger than middle Danian (62.6 Ma) for the analyzed deposits.The palynofacies constituents, as well as the agglutinated foraminiferal morphogroups, used to reconstruct the depositional environments, show that the late Maastrichtian sediments were deposited in an outer shelf to distal (bathyal) environment, followed by a marine transgression during the Danian.     

Palynology and palynofacies of the Upper Cretaceous succession of the El-Noor-1X borehole,northwestern Egypt

Palynological and palynofacies analyses were carried out on some middle–upper Cretaceous samples from the El-Noor-1X borehole, northern Western Desert, Egypt. Palynological age has lead to a refinement of the original ages suggested by the drilling company. Upper Albian–Lower Cenomanian, Upper Cenomanian, and Turonian–Coniacian were recognized. The palaeoenvironment was interpreted on the basis of the ecological preferences of the palynomorphs. It was fluctuating between marginal to inner-middle shelf environment. Distribution of araucaroid pollen and xerophytes suggests that arid or semi-arid paleclimate prevailed during the deposition of the studied sediments. A warm tropical palaeoclimate is suggested on the basis of abundance of hygrophilous plants. Based on the recovered palynological organic matter, two palynofacies were recognized: palynofacies A for the Bahariya Formation, which suggests kerogen type III, and palynofacies B for the upper Bahariya and Abu Roash Formations, which suggests kerogen type IX. Data gathered from the theoretically estimated vitrinite reflectances, which are based on spore/pollen coloration, and visual pterographic kerogen analysis are used to define the source rock potentialities of the studied sediments.     

Infrazonal biostratigraphy of the Upper Cretaceous in the East European province based on benthic foraminifers,Part 2: Santonian-Maastrichtian

A chart of infrazonal biostratigraphic subdivisions in the Cenomanian-Coniacian succession of the East European paleobiogeographic province is substantiated based on distribution of benthic foraminifers. The suggested chart characterizes successive trend of changes in ecologic assemblages of benthic foraminifers and morphologic evolution of certain agglutinated (Gaudryina, Ataxophragmium, Ataxoorbignyina, Arenobulimina, Novatrix, Voloshinovella, Orbignyina, Bolivinopsis) and secretory (Neoflabellina, Globorotalites, Stensioeina, Osangularia, Eponides, Gavelinella, Pseudovalvulineria, Pseudogavelinella, Brotzenella, Cibicides, Cibicidoides, Angulogavelinella, Falsoplanulina, Anomalinoides, Coryphostoma, Bolivinoides, Praebulimina) foraminiferal genera. The chart includes 23 biostratigraphic units (zones and subzones), most of which are recognizable over the vast territory from the Mangyshlak to southern Baltic areas. It is correlated with the acknowledged belemnite, inoceramid, ammonoid and echinoid zonations. Eight stadia of taxonomic changes in foraminiferal assemblages, which are distinguished in this work, show that principal biotic events took place across the middle-late Santonian and Santonian-Campanian boundaries, in the earliest Campanian, at the early-late Campanian boundary time, during the late and terminal Campanian, and in the mid-early Maastrichtian.     

Infrazonal biostratigraphy of the Upper Cretaceous in the East European province based on benthic foraminifers, Part 1: Cenomanian-Coniacian

A chart of infrazonal biostratigraphic subdivision of Cenomanian-Coniacian deposits in the East European paleobiogeographic province is based on distribution of benthic foraminifers. The suggested chart characterizes successive trend of changes in ecologic assemblages of benthic foraminifers and morphologic evolution of certain agglutinated (Gaudryina, Bolivinopsis, Heterostomella, Arenobulimina, Ataxophragmium, Ataxoorbignyna, Marssonella) and secretory (Globorotalites, Valvulinera, Gyroidinoides, Stensioeina, Osangularia, Berthelina, Pseudovalvulineria, Gavelinella, Cibicides, Praebulimina, Reussella) foraminiferal genera. The chart includes 7 zones and 13 subzones, most of which are recognizable over the vast territory from the Mangyshalk to southern Baltic areas. It is correlated with the acknowledged ammonoid and inoceramid zonations. Five stadia of taxonomic changes in foraminiferal assemblages, which are substantiated in this work, show that principal biotic events took place in the mid-late Cenomanian, during the Cenomanian-Turonian and early-middle Turonian transitions, in the late Turonian, and at the early-middle Coniacian boundary time.     

Petrology of peridotite xenolith-bearing basaltic to andesitic lavas from the Shiribeshi Seamount,off northwestern Hokkaido,the Sea of Japan

The Shiribeshi Seamount off northwestern Hokkaido, the Sea of Japan, is a rear-arc volcano in the Northeast Japan arc. This seamount is composed of calc-alkaline and high-K basaltic to andesitic lavas containing magnesian olivine phenocrysts and mantle peridotite xenoliths. Petrographic and geochemical characteristics of the andesite lavas indicate evidence for the reaction with the mantle peridotite xenoliths and magma mixing between mafic and felsic magmas. Geochemical modelling shows that the felsic end-member was possibly derived from melting of an amphibolitic mafic crust. Chemical compositions of the olivine phenocrysts and their chromian spinel inclusions indicate that the Shiribeshi Seamount basalts in this study was derived from a primary magma in equilibrium with relatively fertile mantle peridotites, which possibly represents the mafic end-member of the magma mixing. Trace-element and REE data indicate that the basalts were produced by low degree of partial melting of garnet-bearing lherzolitic source. Preliminary results from the mantle peridotite xenoliths indicate that they were probably originated from the mantle beneath the Sea of Japan rather than beneath the Northeast Japan arc.     

A new,richly fossiliferous member comprised of tidal deposits in the Upper Cretaceous Maevarano Formation,northwestern Madagascar

A new member of the Upper Cretaceous (Maastrichtian) Maevarano Formation is proposed to accommodate a distinctive succession of strata exposed along the shores of Lac Kinkony in northwestern Madagascar. The new Lac Kinkony Member overlies fully terrestrial sandstones of the Anembalemba Member of the Maevarano Formation, and is capped by marine dolostones of the Berivotra Formation. In the stratotype section, the base of the Lac Kinkony Member consists of siltstone interbeds that host networks of Ophiomorpha. Siltstone facies pass up-section to distinctive white sandstones packed with dolomitic mud matrix that exhibit rhythmic clay drapes, flaser and wavy bedding, and oppositely-oriented ripples developed on the toes of larger foresets. Thin flat interbeds of microgranular dolostone and claystone comprise the uppermost facies of the Lac Kinkony Member, and a laterally traceable ravinement bed mantled by cobbles of rounded dolostone marks the contact with the superjacent Berivotra Formation. Deposits of the Lac Kinkony Member are interpreted to represent siliciclastic and carbonate tidal flats dissected by tidally-influenced rivers. Vertebrate fossils are abundantly preserved in these coastal deposits, and are locally concentrated in microfossil bonebeds that have the potential to yield thousands of small identifiable specimens. In addition to many taxa already known from the Maevarano Formation, the Lac Kinkony Member has yielded a wealth of phyllodontid albuloid fish skull elements, the distal humerus of a new frog taxon, five vertebrae representing two new snakes, a tooth of a possible dromaeosaurid, and a complete skull of a new mammal. The discovery of several new vertebrate taxa from this new member reflects the fact that it samples a previously unsampled nearshore, peritidal paleoenvironment in the Late Cretaceous of Madagascar.     

Integrated chemostratigraphy and biostratigraphy of the Windermere Supergroup, northwestern Canada: implications for Neoproterozoic correlations and the early evolution of animals

The thick, richly fossiliferous succession of the upper Windermere Supergroup, Mackenzie Mountains, northwestern Canada, provides a test of integrated biostratigraphic and chemostratigraphic frameworks in terminal Proterozoic correlation. The C- and Sr-isotopic abundances of lower Keele Formation carbonates approximate those for other pre-Varanger samples, confirming that the simple disc-like fossils of the underlying Twitya Formation predate all known diverse Ediacaran faunas. "Tepee" and Sheepbed carbonates record strong post-glacial isotopic excursions; in contrast, delta13C values for Gametrail through Risky carbonates vary only within the narrow range of about +l% to +2%. A second negative excursion occurs in Ingta Formation carbonates that immediately underlie the paleontologically determined Precambrian-Cambrian boundary. The upper Windermere profile as a whole compares closely with curves determined for other terminal Proterozoic successions. The lowermost diverse Ediacaran assemblages in the Sheepbed Formation correlate chemostratigraphically with the oldest fauna in Namibia, but the two assemblages differ in taxonomic composition. Blueflower assemblages correlate both chemostratigraphically and taxonomically with faunas from Australia, China, Siberia, and elsewhere. Increasing data support the hypothesis that paleontological and geochemical data together provide a reliable means of correlating terminal Proterozoic sedimentary rocks throughout the world.     

Dinocyst biostratigraphy of the Lower Cretaceous in North Siberia

The described scheme of the Lower Cretaceous (Berriasian-Barremian) stratigraphic subdivisions is elaborated based on palynological study of sections in the Khatanga depression, Ust-Yenisei region, Pur-Taz interfluve, and around the Ob River latitudinal segment, the North Siberia. Stratigraphic distribution of microphytoplankton studied in detail is used to distinguish 10 biostratigraphic units in the rank of dinocysts zones. Stratigraphic position of the zones is determined with confidence using data on the Lower Cretaceous reference sections in the Khatanga depression, which were principal ones by constructing the Boreal standard zonation. In majority, boundaries of the dinocysts beds are of a high correlation potential and can be regarded as reliable stratigraphic markers, as they are recognizable not only in Siberia, but also in northern Europe and America.     

Contribution to the biostratigraphy of the Lower Cretaceous in eastern Serbia: Upper Hauterivian Orbitolinidae from the Kamenica 1 section,Kurilovo anticline

Upper Hauterivian deposits in the Kurilovo area, Kamenica 1 section, NE of Niš, are described based of abundant and diverse orbitolinids. So far, the interval was assigned to the Barremian–Aptian on the geological map. Such a new age assignment results from the first detailed study carried out on the orbitolinid fauna contained in the Lower Cretaceous (upper Hauterivian) shallow-water limestones of eastern Serbia. The upper Hauterivian is documented on the basis of two key stratigraphic markers, specifically Valserina primitiva and Paleodictyoconus beckerae.In addition to these late Hauterivian index fossils, the studied section bears orbitolinids having a larger stratigraphic distribution: Cribellopsis neoelongata, Cribellopsis thieuloyi?, Montseciella glanensis, Orbitolinopsis debelmasi, Orbitolinopsis cf. debelmasi, Orbitolinopsis sp., Paleodictyoconus cuvillieri, Paleodictyoconus cf. cuvillieri, Paleodictyoconus cf. beckerae, Paleodictyoconus cf. actinostoma, Paleodictyoconus sp., Paracoskinolina? jourdanensis, Paracoskinolina cf. hispanica, Urgonina alpillensis, Valserina sp. The microfossil assemblage includes other foraminifers such as Charentia cuvillieri, Mayncina bulgarica, Nautiloculina cretacea, Pfenderina globosa, Pseudocyclammina cf. lituus, Pseudolituonella gavonensis, Ammobaculites sp., Bolivinopsis sp., abundant trocholinids, various miliolids, other foraminifers and sparse algae which will be presented separately.     

日本北海道音调津的球状石墨

本文利用光学显微镜、X射线衍射仪、透射电镜等手段对日本北海道音调津球状石墨的成因、结构等进行研究,认为日本北海道音调津球状石墨中除了石墨外,还存在镍黄铁矿﹑黄铜矿﹑磁黄铁矿等金属硫化物矿物。通过透射电子显微镜观察发现音调津石墨中大部分是椭圆状石墨颗粒,高分辨晶格条纹主要是非晶态结构,但是同时也发现了结晶良好的石墨。高结晶度石墨的存在可能是在硫化物Fe-Ni-Cu的触媒作用下形成的。天然球状石墨的微结构和性状研究对天然石墨的形成和开发应用有重要意义,对新型碳材料的制备技术有重要科学意义。     

Upper Cretaceous depositional environments and bivalve assemblages of far-east Asia: the Himenoura Group, Kyushu, Japan

The depositional environments and bivalve assemblages are determined for the Upper Cretaceous Hinoshima Formation of the Himenoura Group, Kamishima, Amakusa Islands, Kyushu, Japan. The Hinoshima Formation is characterized by a thick transgressive succession that varies from incised-valley-fill deposits to submarine slope deposits with high aggradation rates of depositional systems. The incised valley is filled with fluvial, bayhead delta, brackish-water estuary, and marine embayment deposits, and is overlain by thick slope deposits.Shallow marine bivalves are grouped into five fossil assemblages according to species composition: Glycymeris amakusensis (foreset beds of a bayhead delta), Nippononectes tamurai (foreset beds of a bayhead delta), Ezonuculana mactraeformis–Nucula formosa (central bay), Glycymeris amakusensis–Apiotrigonia minor (slope), and Inoceramus higoensis–Parvamussium yubarensis (slope). These bivalve assemblages all represent autochthonous and parautochthonous conditions except for a Glycymeris amakusensis–Apiotrigonia minor assemblage found in debris flow and slump deposits. The life habitats of these bivalves and the compositions of the assemblages are discussed in terms of the ecological history of fossil bivalves of the mid- to Late Cretaceous.     

Late Cenozoic Metallogeny of Southwest Hokkaido, Japan

Abstract: This paper reviews the Miocene to Pleistocene tectonic framework, geology, magmatic style and stress field of southwest Hokkaido, Japan, and compiles deposit form, type, ore and alteration minerals, strike and length of mineralized veins, and associated igneous activity. The late Cenozoic tectonic regime of the Sapporo‐Iwanai ore district is divided into five periods on the basis of the subduction mode of the Pacific plate: Period 1 (15.0–12.1 Ma) oblique‐subduction setting with an orthogonal convergence rate (OCR) of 51–81 mm/y; Period 2 (12.1–6.2 Ma) normal subduction with an OCR of 81–94 mm/y; Period 3 (6.2–3.6 Ma) oblique subduction setting with an OCR of 73–99 mm/y; Period 4 (3.6–1.5 Ma) normal‐subduction setting with an OCR of 99–103 mm/y and Period 5 (1.5–0 Ma) oblique‐subduction setting with an OCR from 99 to 57 mm/y. The hydrothermal deposits in the district include Kuroko deposits of Period 1 and epithermal vein‐type deposits of Periods 2 to 5. The Kuroko deposits were accompanied by submarine monogenetic rhyolite volcanism associated with tholeiitic basalt in the backarc region. In contrast, Late Miocene to Pliocene epithermal vein‐type deposits were associated mainly with polygenetic andesite and monogenetic rhyodacite volcanism of calc‐alkaline series. These different styles of magmatism occurred in an extensional tectonic regime (Period 1), and weakened extensional (Periods 2–3) or neutral tectonic regimes (Periods 4). Periods 2–5 epithermal vein‐type deposits are divided into base‐metal and precious‐metal deposits. The base‐metal deposits are associated mainly with large (>5 km in diameter) polygenetic andesitic volcanoes and subvol‐canic intrusions. The precious‐metal deposits are associated with small (<5 km in diameter) polygenetic or monogenetic volcanoes and/or subvolcanic intrusions of andesite, dacite and rhyolite near the volcanic arc front. This difference in distribution is ascribed to different states of horizontal differential stress. Productive vein‐type deposits, such as Toyoha, Inakuraishi, Ohe and Chitose, formed in the neutral regime with a large horizontal differential stress during Period 4, which may have promoted strike‐slip faulting and non–extrusive, subvol‐canic intrusion. This tectonic regime and stress field resulted from the normal subduction of the Pacific plate with elevated velocity. This observation leads to the conclusion that large metallic deposits in southwest Hokkaido are expected to have formed primarily during Pliocene magmatic‐hydrothermal activity, when the orthogonal convergence rate was highest and strike‐slip faulting was active.     

Petrology of a mantle-derived rhyolite, Hokkaido, Japan

The Miocene Kitami rhyolite, consisting of orthopyroxene and plagioclase-phyric lavas and dikes, occurs on the back-arc side of the Kuril arc with coeval basalts and Fe-rich andesites. Temperatures estimated from orthopyroxene–ilmenite pairs exceed 900°C. Although the whole rock compositions of the Kitami rhyolite correspond to S-type granites (i.e., high K, Al, large ion lithophile elements, and low Ca and Sr), Sr–Nd isotope compositions are remarkably primitive, and similar to those of the coeval basalts and andesites. They are distinct from those of lower crustal metamorphic rocks exposed in the area. Comparison of chondrite-normalized rare earth element (REE) patterns between the rhyolite and the basalts and andesites show that the rhyolite is more light REE enriched, but has similar heavy REE contents than the basalts. All rhyolites show negative Eu anomalies. The geochemical data suggest that did not formed by simple dehydration melting of basaltic rocks or fractional crystallization of basaltic magmas. The features of slab-derived fluids expected from recent high pressure experimental studies indicates that mantle wedge is partly metasomatized with “rhyolitic” materials from subducted slabs; it is more likely that very low degree partial melting of the metasomatized mantle wedge formed the rhyolite magma.     

Upper cretaceous biostratigraphy of the south-central Pyrenees (Lleida,Spain)

Abstract

Biostratigraphical data using larger foraminifera and planktonic foraminifera permitted us to establish the correlation between shallow platform sediments rich in larger foraminifera (Montsec and Serres Marginals thrust sheets) and deeper ones containing planktonic foraminifera (Boixols thrust sheet).

Consequently, the Santa Fe limestones containing Ovalveolina-Praealveolinaassemblage represent the Cenomanian. Early Turonian ( ‘IT~ archaeocretacea and P. helvetica zones) exist in both, Montsec and Boixols thrust sheets and it is constituted by Pithonella limestones. Late Turonian (M. schneegansi zone) is only present in Boixols thrust sheet (Reguard Fm.), the Montsec thrust sheet having an erosive hiatus.

Late Coniacian-Early Santonian (D. Concavata zone) is represented in the Montsec thrust sheet (Cova Limestones) and in the eastern part of the Boixols thrust sheet (St. Corneli Fm.) by two differents facies giving two different microfaunal assemblages; the firts one, characterized by Ophtalmidiidae s.l indicate a restricted lagoonal environment while the second one, characterized by diverses species of complex agglutinated, Fabulariidae, Meandropsinidae and Rotaliidae, represents an open shallow platform. In the Boixols thrust sheet (Anseroles Fm.) dominate the planktonic foraminifera and small benthic.

In the late Santonian (D. asyrnetrica zone) the sea reached as far as Serres Marginales thrust sheet where sediments (Tragó de Noguera unit) are terrigenous and deposited in a very shallow platform. In the Montsec thrust sheet (Montsec marls) the larger foraminifera indicate a platform deeper than that of the Serres Marginals thrust sheet. In the Boixols thrust sheet the sediments are deposited in an outer platform (Herbasavina Fm.) or turbiditic basin (Mascarell Mb.).

During Campanian times the transgresion reaches the maximum. In the Serres Marginals sediments are deposited in a restricted shallow environment rich in Meandropsinidae (Serres Limestones). In the Montsec thrust sheet they are deposited in a platform with patch reefs and shoals (Terradets limestones) and in the Boixols one in an outer platform, talus and/or basin.

During Early Maastrichtian times (C. falsostuarti zone) terrigenous materials arrived in the basin, the rate of sedimentation increased outstripping the subsidence rate and the retreat of the sea to the north. Late Maastrichtian (C. gansseri zone) is only present in the Boixols thrust sheet.     

Migmatite tectonics of the hidaka metamorphic belt, Hokkaido, Japan

The Hidaka metamorphic belt is situated at the junction of the Honshu and Kuril arcs in the axial zone of Hokkaido in northern Japan. Various migmatites, which occupy the core of the metamorphic belt, are classified as lens, sheet, falling star and dome facies on the basis of composition, scale and form as proposed by Harland (1956). Each facies is produced progressively. Movement is first lateral and then upwards at the sheet facies stage, followed by the development of the diapiric falling star and dome facies. Subsequently, the granitic phase starts to form from the lens facies, again within the migmatite sheets, leading to the emplacement of granitic plutons. The movement of the migmatite and granite bodies is controlled by the tangential stress field, as well as by the buoyancy in the gravitational field.     

Integrated biostratigraphy and carbon isotope stratigraphy of the Lower Cretaceous (Barremian to Albian) Adriatic-Dinaridic carbonate platform deposits in Istria, Croatia

The Adriatic-Dinaridic carbonate platform (ADCP) was one of the largest and relatively well preserved Mesozoic platforms in the Mediterranean region (central Tethys). The peninsula Istria, in the northwestern part of the ADCP, is built up predominantly of shallow-water carbonates of the Middle Jurassic (Dogger) to Eocene age and, to a lesser extent, of Paleogene clastic deposits (flysch and calcareous breccia). This study focuses on a Lower Cretaceous (Barremian to Albian) succession of strata at five localities in western Istria. Stratigraphic determinations are based on identification of nine microfossil assemblages (benthic foraminifera and calcareous algae Dasycladales) and on using their taxa as index fossils. The age of strata with these microfossil assemblages, however, is questionable. Most of the age uncertainties are associated with a regional emersion, which occurred on the ADCP during the Aptian or close to the Aptian-Albian transition. It is unclear what portions of the Upper Aptian and/or Lower Albian are missing along this unconformity. A stable isotope study was conducted on homogenous micritic matrix samples in an attempt to resolve some of these uncertainties. Variations in carbon isotope compositions proved useful for stratigraphic correlation between the examined successions of strata, for improving their age determination, and for relating them to other coeval successions that span an important time interval of major oceanographic changes and carbon-cycle perturbations associated with the Early Aptian oceanic anoxic event (OAE 1a).