Late Quaternary vegetation,climate and fire dynamics inferred from the El Tiro record in the southeastern Ecuadorian Andes

In order to study the stability and dynamics of mountain rainforest and paramo ecosystems, including the biodiversity of these ecosystems, the Holocene and late Pleistocene climate and fire variability, and human impact in the southeastern Ecuadorian Andes, we present a high‐resolution pollen record from El Tiro Pass (2810 m elevation), Podocarpus National Park. Palaeoenvironmental changes, investigated by pollen, spores and charcoal analysis, inferred from a 127 cm long core spanning the last ca. 21 000 cal. yr BP, indicate that grass‐paramo was the main vegetation type at the El Tiro Pass during the late Pleistocene period. The grass‐paramo was rich in Poaceae, Plantago rigida and Plantago australis, reflecting cold and moist climatic conditions. During the early Holocene, from 11 200 to 8900 cal. yr BP, subparamo and upper mountain rainforest vegetation expanded slightly, indicating a slow warming of climatic conditions during this period. From 8900 to 3300 cal. yr BP an upper mountain rainforest developed at the study site, indicated by an increase in Hedyosmun, Podocarpaceae, Myrsine and Ilex. This suggests a warmer climate than the present day at this elevation. The modern subparamo vegetation became established since 3300 cal. yr BP at El Tiro Pass. Fires, probably anthropogenic origin, were very rare during the late Pleistocene but became frequent after 8000 cal. yr BP. Copyright © 2007 John Wiley & Sons, Ltd.     

The influence of fine‐scale topography on the impacts of Holocene fire in a Tasmanian montane landscape

Tasmania's montane temperate rainforests contain some of Australia's most ancient and endemic flora. Recent landscape‐scale fires have impacted a significant portion of these rainforest ecosystems. The complex and rugged topography of Tasmania results in a highly variable influence of fire across the landscape, rendering predictions of ecosystem response to fire difficult. We assess the role of topographic variation in buffering the influence of fire in these endemic rainforest communities. We developed a new 14 000‐year (14‐ka) palaeoecological dataset from Lake Perry, southern Tasmania, and compared it to neighbouring Lake Osborne (<250 m distant) to examine how topographic variations influence fire and vegetation dynamics through time. Repeated fire events during the Holocene cause a decline in montane rainforest taxa at both sites; however, in the absence of fire, rainforest taxa are able to recover. Montane temperate rainforest taxa persisted at Lake Perry until European settlement, whilst these taxa were driven locally extinct and replaced by Eucalyptus species at Lake Osborne after 2.5 ka. Contiguous topographic fire refugia within the Lake Perry catchment probably provided areas of favourable microclimates that discouraged fire spread and supported the recovery of these montane temperate rainforests. Copyright © 2019 John Wiley & Sons, Ltd.     

Vegetation and climates in southern Tasmania since the last glaciation

Enclosed basins (glacial and nonglacia) of Tasmania contain the most comprehensive record in Australia of trends in a regional vegetation and climate since the late Pleistocene. Seven pollen sequences, each continuous and extending back at least 10,000 years, are used to reconstruct the history of postglacial vegetation and climate in Southern Tasmania (42°S–43°30′S). Interpretations are supported by a study of the modern pollen rain. Postglacial climates in Tasmania were characterized by a strong west-to-east decrease in precipitation. During the late Pleistocene, climates were markedly colder and drier than at present, and the vegetation was largely devoid of trees. A major rise in temperature between ca. 11,500 and 9500 yr B.P., accompanied by rising effective precipitation, resulted in the expansion of Eucalyptus, then other trees, across Tasmania. This warming trend may have been temporarily reversed during the early postglacial. Dry climates delayed the development of forest in inland eastern Tasmania until after ca. 9500 yr B.P. There is no evidence for a major change in climate since this temperature rise. Two broad phases of development have occurred within the postglacial forests. The first was an early Holocene phase during which Nothofagus cunninghamii cool temperate rain forest developed in western Tasmania and on the slopes of mountains in central and southeastern Tasmania. Eucalyptus sclerophyll forests developed in eastern Tasmania and have remained dominant there since. By ca. 7800 yr B.P. rain-forest communities were established beyond present-day limits. The second phase was a mid to late Holocene phase during which forests and alpine vegetation became more open in structure, leading to the re-expansion of Eucalyptus and shade-intolerant species. During the early to mid Holocene, climates in Southern Tasmania were wetter and (? then) warmer than at present. Maximum and minimum dates for this “optimum” are 8000 and 5000 yr B.P. Since then, climates have become increasingly rigorous, possibly through an increased incidence of inequable “weather types” leading to an increase in the frequency of drought and frost. Structural changes in the postglacial vegetation of Southern Tasmania closely parallel those at equivalent latitudes in New Zealand and Chilean South America, hence are likely to reflect the same primary cause.     

A 23,000-yr pollen record from Lake Euramoo, Wet Tropics of NE Queensland, Australia

A new extended pollen and charcoal record is presented from Lake Euramoo, Wet Tropics World Heritage rainforest of northeast Queensland, Australia. The 8.4-m sediment core taken from the center of Lake Euramoo incorporates a complete record of vegetation change and fire history spanning the period from 23,000 cal yr B.P. to present. The pollen record is divided into five significant zones; 23,000–16,800 cal yr B.P., dry sclerophyll woodland; 16,800–8600 cal yr B.P., wet sclerophyll woodland with marginal rainforest in protected pockets; 8600–5000 cal yr B.P., warm temperate rainforest; 5000–70 cal yr B.P., dry subtropical rainforest; 70 cal yr B.P.–AD 1999, degraded dry subtropical rainforest with increasing influence of invasive species and fire.The process of rainforest development appears to be at least partly controlled by orbital forcing (precession), though more local environmental variables and human activity are also significant factors. This new record provides the opportunity to explore the relationship between fire, drought and rainforest dynamics in a significant World Heritage rainforest region.     

Late‐glacial and Holocene record of vegetation and climate from Cynthia Bay,Lake St Clair,Tasmania

A Late‐glacial–Holocene pollen record was obtained from a 3.96 m sediment core taken from Lake St Clair, central Tasmania. Modern vegetation and pollen analyses formed the basis for interpretation of the vegetation and climate history. Following deglaciation and before ca. 18450 yr BP Podocarpus lawrencei coniferous heath and Astelia–Plantago wet alpine herbfield became established at Lake St Clair. A distinct Poaceae‐Plantago peak occurs between 18450 and 11210 yr BP and a mean annual temperature depression from ca. 6.2°C to 3°C below present is inferred for this period. The marked reduction in Podocarpus and strong increase of Poaceae suggests reduced precipitation levels during the period of widespread deglaciation (ca. 18.5–11 kyr BP). The local Late Pleistocene–Holocene non‐forest to forest biostratigraphical boundary is dated at 11.2 kyr BP. It is characterised by expansion of the subalpine taxa Athrotaxis/Diselma with Nothofagus gunnii, and by the establishment of Nothofagus cunninghamii with Eucalyptus spp. A ‘Phyllocladus bulge’ prior to the expansion of Nothofagus cunninghamii, reported at other Tasmanian sites, is not present at Lake St Clair. Nothofagus cunninghamii cool temperate rainforest peaked at 7800 yr BP, probably under wetter climatic conditions than present. The maximum development of rainforest in the early–middle Holocene may indicate that the temperature was slightly warmer than present, but the evidence is not definitive. The expansion of Eucalyptus spp. and Poaceae after 6000 yr BP may be partly a disclimax effect as a result of Aboriginal burning, but appears also to reflect reduced precipitation. The changes in vegetation and inferred climate can be explained by major changes in synoptic patterns of southern Australia and the adjacent southwest Pacific. Copyright © 2000 John Wiley & Sons, Ltd.     

Late Pleistocene and Holocene vegetation and climate records from Lake Kotokel, central Baikal region

Climatically driven Late Pleistocene and Holocene vegetation changes were reconstructed based on pollen records from the sediments of Lake Kotokel and Cheremushka Bog, located on the eastern shore of Lake Baikal. The described paleoenvironmental record has higher resolution than records collected from Lake Baikal and unites individual events identified in prior studies of bottom and onshore cores. Remarkable shifts in landscapes and expansions of index plants are as follows. Forest tundra and/or forest steppe landscape with birch, spruce, Artemisia, and Poaceae prevailed at ca. 50–25 ¹⁴C kyr BP. Tundra and/or steppe vegetation dominated by Artemisia and Poaceae was typical for the Last Glacial Maximum. The expansion of shrub birch and willow occurred at ca. 15.5 ¹⁴C kyr BP. Two peaks of spruce expansion at ca. 47.5–42.4 ¹⁴C kyr BP (Karginian time) and at ca. 14.5–13 ka (Bølling-Allerød warm intervals) suggest that the condition were more humid than today. A slight increase in Artemisia at ca. 11–10.5 ¹⁴C kyr BP (13–12 ka) was indicative of the Younger Dryas event. An expansion of birch forests with fir at ca. 12–6.4 ka suggests higher humidity. The currently dominant Scots and Siberian pine forests with birch expanded since 6.4 ka.     

Vegetation and climate changes in northwestern Russia during the Lateglacial and Holocene inferred from the Lake Ladoga pollen record

The new pollen record from the upper 12.75 m of a sediment core obtained in Lake Ladoga documents regional vegetation and climate changes in northwestern Russia over the last 13.9 cal. ka. The Lateglacial chronostratigraphy is based on varve chronology, while the Holocene stratigraphy is based on AMS ¹⁴C and OSL dates, supported by comparison with regional pollen records. During the Lateglacial (c. 13.9–11.2 cal. ka BP), the Lake Ladoga region experienced several climatic fluctuations as reflected in vegetation changes. Shrub and grass communities dominated between c. 13.9 and 13.2 cal. ka BP. The increase in Picea pollen at c. 13.2 cal. ka BP probably reflects the appearance of spruce in the southern Ladoga region at the beginning of the Allerød interstadial. After c. 12.6 cal. ka BP, the Younger Dryas cooling caused a significant decrease in spruce and increase in Artemisia with other herbs, indicative of tundra‐ and steppe‐like vegetation. A sharp transition from tundra‐steppe habitats to sparse birch forests characterizes the onset of Holocene warming c. 11.2 cal. ka BP. Pine forests dominated in the region from c. 9.0 to 8.1 cal. ka BP. The most favourable climatic conditions for deciduous broad‐leaved taxa existed between c. 8.1 and 5.5 cal. ka BP. Alder experiences an abrupt increase in the local vegetation c. 7.8 cal. ka BP. The decrease in tree pollen taxa (especially Picea) and the increase in herbs (mainly Poaceae) probably reflect human activity during the last 2.2 cal. ka. Pine forests have dominated the region since that time. Secale and other Cerealia pollen as well as ruderal herbs are permanently recorded since c. 0.8 cal. ka BP.     

A record of Lateglacial/Holocene environmental change from a high‐elevation site in the Intermountain West,USA

Records of past vegetation and fire history can be complicated by changes in the depositional environment of a sampling location. However, these changes can alternatively be used as a measure of climate variability. Our study site, ca. 18.0 cal. ka BP record from Little Brooklyn Lake, Wyoming, located near the crest of the Snowy Range, records three moisture states. Initially, the lake was likely a glacier‐fed pond indicated by the presence of Pediastrum algae colonies. Around 13.0 cal. ka BP this pond transitioned to a meadow environment, suggested by the loss of Pediastrum algae colonies and slow sedimentation rates. Meadow conditions were maintained until ca. 5.0 cal. ka BP when Pediastrum algae colony abundance increased,indicating the formation of a shallow lake. From 18.0 to ca. 5.0 cal. ka BP, the pollen record is suggestive of alpine vegetation conditions with relatively high spruce and herbaceous taxa. Low charcoal influx also characterizes the period between 18.0 and 5.0 cal. ka BP. After 5.0 cal. ka BP, the coincidence of the formation of shallow lake and pollen data, indicating a shift to a spruce and fir forest, suggests an increase in effective moisture. Fire remained rare in this basin over the entire record, however, once the lake established sedimentation rates and charcoal influx increased. Copyright © 2013 John Wiley & Sons, Ltd.     

Fire and Vegetation History from the Coastal Rain Forest of the Western Oregon Coast Range

High-resolution charcoal and pollen analyses were used to reconstruct a 4600-yr-long history of fire and vegetation near Taylor Lake in the wettest forests of coastal Oregon. Today, fires in these forests are rare because the season of ignition does not coincide with months of dry fuels. From ca. 4600 to 2700 cal yr B.P. fire episodes occurred at intervals of 140±30 yr while forest vegetation was dominated by disturbance-adapted taxa such as Alnus rubra. From ca. 2700 cal yr B.P. to the present, fire episodes have become less common, occurring at intervals of 240±30 yr, and fire-sensitive forest taxa, such as Tsuga heterophylla and Picea sitchensis, have become more prominent. Fire occurrence during the mid-Holocene was similar to that of the more xeric forests in the eastern Coast Range and suggests that summer drought was widespread. After ca. 2700 cal yr B.P., a decrease in fire episode frequency suggests that cooler conditions and possibly increased summer fog allowed the establishment of present-day Picea sitchensis forests within the watershed. These results provide evidence that fire has been an important disturbance agent in the Coast Range of Oregon, and variations in fire frequency and climate have led to the establishment of present-day forests.     

南海南部末次冰期以来的孢粉记录及其气候意义

本文通过分析南海南部１７９６４柱状样的孢粉记录,重建了该地区２６０００ａＢ．Ｐ．以来的植被和气候变化历史。南海南部末次盛冰期时高山雨林的范围可能是现在的２～３倍,表明气温明显下降;从末次盛冰期到冰消期,巽他陆架一直为热带低地雨林及红树林占据,表明在此期间湿度没有明显变化,但存在着千年级的准周期波动。这种波动一直持续到全新世中期。本地区在２０００ａＢ．Ｐ．以来芒萁孢子大量增加,一些高山雨林的成分重新出现,可能与人类活动所导致的低地植被的破坏有关。     

Abrupt vegetation changes during the last glacial to Holocene transition in mid‐latitude South America

A pollen record from the Huelmo site (ca. 41°30′S) shows that vegetation and climate changed at millennial time‐scales during the last glacial to Holocene transition in the mid‐latitude region of western South America. The record shows that a Nothofagus parkland dominated the landscape between 16 400 and 14 600 ¹⁴C yr BP, along with Magellanic Moorland and cupressaceous conifers. Evergreen North Patagonian rainforest taxa expanded in pulses at 14 200 and 13 000 ¹⁴C yr BP, following a prominent rise in Nothofagus at 14 600 ¹⁴C yr BP. Highly diverse, closed canopy rainforests dominated the lowlands between 13 000 and 12 500 ¹⁴C yr BP, followed by the expansion of cold‐resistant podocarps and Nothofagus at ca. 12 500 and 11 500 ¹⁴C yr BP. Local disturbance by fire favoured the expansion of shade‐intolerant opportunistic taxa between 10 900 and 10 200 ¹⁴C yr BP. Subsequent warming pulses at 10 200 and 9100 ¹⁴C yr BP led to the expansion of thermophilous, summer‐drought resistant Valdivian rainforest trees until 6900 ¹⁴C yr BP. Our results suggest that cold and hyperhumid conditions characterised the final phase of the Last Glacial Maximum (LGM), between 16 400 and 14 600 ¹⁴C yr BP. The last ice age Termination commenced with a prominent warming event that led to a rapid expansion of North Patagonian trees and the abrupt withdrawal of Andean ice lobes from their LGM positon at ca. 147 000 ¹⁴C yr BP. Hyperhumid conditions prevailed between 16 400 and 13 000 ¹⁴C yr BP, what we term the ‘extreme glacial mode’ of westerly activity. This condition was brought about by a northward shift and/or intensification of the southern westerlies. The warmest/driest conditions of the last glacial–interglacial transition occurred between 9100 and 6900 ¹⁴C yr BP. During this period, the westerlies shifted to an ‘extreme interglacial mode’ of activity, via a poleward migration of stormtracks. Our results indicate that a highly variable climatic interval lasting 5500 ¹⁴C years separate the opposite extremes of vegetation and climate during the last glacial‐interglacial cycle, i.e. the end of the LGM and the onset of the early Holocene warm and dry period. Copyright © 2003 John Wiley & Sons, Ltd.     

Holocene fire and vegetation dynamics in a montane forest, North Cascade Range, Washington, USA

We reconstructed a 10,500-yr fire and vegetation history of a montane site in the North Cascade Range, Washington State based on lake sediment charcoal, macrofossil and pollen records. High-resolution sampling and abundant macrofossils made it possible to analyze relationships between fire and vegetation. During the early Holocene (> 10,500 to ca. 8000 cal yr BP) forests were subalpine woodlands dominated by Pinus contorta. Around 8000 cal yr BP, P. contorta sharply declined in the macrofossil record. Shade tolerant, mesic species first appeared ca. 4500 cal yr BP. Cupressus nootkatensis appeared most recently at 2000 cal yr BP. Fire frequency varies throughout the record, with significantly shorter mean fire return intervals in the early Holocene than the mid and late Holocene. Charcoal peaks are significantly correlated with an initial increase in macrofossil accumulation rates followed by a decrease, likely corresponding to tree mortality following fire. Climate appears to be a key driver in vegetation and fire regimes over millennial time scales. Fire and other disturbances altered forest vegetation at shorter time scales, and vegetation may have mediated local fire regimes. For example, dominance of P. contorta in the early Holocene forests may have been reinforced by its susceptibility to frequent, stand-replacing fire events.     

Reconstructing the palaeoenvironments of the early Pleistocene mammal faunas from the pollen preserved on fossil bones

We carried out a systematic investigation on the pollen content of sediment adhering to skeletal elements of large mammals which originate from the long lacustrine record of Leffe (Early Pleistocene of the Italian Alps). Three local faunas were discovered during mining activities along the intermediate part (spanning from 1.5 to 0.95 Ma) of the basin succession. The excellent pollen preservation allowed testing the reproducibility of the pollen signal from single skeletons. A clear palaeoenvironmental patterning, consistent with the ecological preferences of the considered mammal species, emerged from the canonical correspondence analysis of pollen types diagnostic for vegetation communities. Edaphic factors related to seasonal river activity changes and to the development of swamp forests in the riverbanks are significantly associated to the occurrences of Hippopotamus cf. antiquus, whereas finds of Mammuthus meridionalis belong to fully forested landscapes dominated by conifer or mixed forests of oceanic, warm to cool-temperate climate. Rhinoceros habitats include variable forest cover under different climate states. Distinct cool-temperate, partially open vegetation could be recognized for large deer included Cervalces cf carnutorum.A palynostratigraphic correlation between individual spectra and a reference palynostratigraphic record allowed assignment of many fossil remains to a precise stratigraphic position. This procedure also shown that the Leffe local faunas include specimens accumulated under different environmental and climate states, as a consequence of high-frequency climate changes characterizing the Late Villafranchian Early Pleistocene.     

Lateglacial vegetation and palaeoenvironment in W Norway,with new pollen data from the Sunnmøre region

Krüger, L. C., Paus, A., Svendsen, J. I. & Bjune, A. E. 2011: Lateglacial vegetation and palaeoenvironment in W Norway, with new pollen data from the Sunnmøre region. Boreas, 10.1111/j.1502‐3885.2011.00213.x. ISSN 0300‐9483. Two sediment sequences from Sunnmøre, northern W Norway, were pollen‐analytically studied to reconstruct the Lateglacial vegetation history and climate. The coastal Dimnamyra was deglaciated around 15.3 ka BP, whereas Løkjingsmyra, further inland, became ice‐free around 14 ka BP. The pioneer vegetation dominated by snow‐bed communities was gradually replaced by grassland and sparse heath vegetation. A pronounced peak in Poaceae around 12.9 ka BP may reflect warmer and/or drier conditions. The Younger Dryas (YD) cooling phase shows increasing snow‐bed vegetation and the local establishment of Artemisia norvegica. A subsequent vegetation closure from grassland to heath signals the Holocene warming. Birch forests were established 500–600 years after the YD–Holocene transition. This development follows the pattern of the Sunnmøre region, which is clearly different from the Empetrum dominance in the Lateglacial interstadial further south in W Norway. The Lateglacial oscillations GI‐1d (Older Dryas) and GI‐1b (Gerzensee) are hardly traceable in the north, in contrast to southern W Norway. The southern vegetation was probably closer to an ecotone and more susceptible to climate changes.     

Fire and drought as drivers of early Holocene tree line changes in the Peruvian Andes

Three pollen and charcoal records from three lakes lying at 3400 m elevation in southern Peru provided a record of landscape change spanning the last ca.18 000 cal. a BP. The tree line lay close to the site between ca. 16 000 and 12 000 cal. a BP, with Polylepis woodlands growing near the lakes. Progressively drying conditions led to increased fire after 12 000 cal. a BP, coinciding with a decline in Polylepis cover and Andean forest relicts as puna grasslands expanded. A strong decrease in the rate of sediment deposition between ca. 12 000 and ca. 4400 cal. a BP was interpreted to indicate the presence of sedimentary hiatuses. With the return of wet conditions after 4400 cal. a BP, forests did not reassemble around the lakes. Instead, fire‐maintained grasslands dominated the landscape. Humans probably induced the intensified fire activity during the late Holocene and thereby deflected local successions. The modern fragmented landscape, with Polylepsis woodlands existing in fire‐resistant pockets above the general limit of the Andean tree line, resulted from the intensification of human land use practices during the last 4400 cal. a BP. Copyright © 2011 John Wiley & Sons, Ltd.     

Postglacial vegetation community change over an elevational gradient on the western Kenai Peninsula,Alaska: pollen records from Sunken Island and Choquette Lakes

The Kenai Peninsula of south‐central Alaska is a region of high topographic diversity with a complex glacial history. The sedimentary record of two small lakes [Sunken Island (SIL; 76 m a.s.l.) in the Kenai Lowlands; Choquette (CL; 527 m a.s.l.) in the Caribou Hills upland] exemplifies the postglacial development of the conifer–hardwood forest over an elevational range there. A herb–shrub tundra was established at both sites after deglaciation. By ~10.7 ka, poplar (Populus sp.) and alder (Alnus) dominated the lowland forest, while alder with minor poplar occurred at the upland site. Lake levels lower than today occurred during the early Holocene until ~8 ka. Subsequently at SIL, the near‐modern Kenai birch (Betula kenaica) – white spruce (Picea glauca) forest maintained prominence throughout the Holocene. However, at CL, alder dominated with dwarf birch and other subshrubs; small amounts of white spruce arrived ~5.2 ka. Black spruce (Picea mariana) grew around SIL by ~4 ka, but never gained prominence at CL. Fire, a prominent agent of disturbance in the Kenai Lowlands since ~8 ka, was essentially absent at the hardwood‐dominated upland site before ~6 ka, and rare thereafter. This suggests an important link between fire and spruce in Kenai forests.     

Chilean and Southeast Pacific paleoclimate variations during the last glacial cycle: directly correlated pollen and δO records from ODP Site 1234

Joint pollen and oxygen isotope data from Ocean Drilling Program Site 1234 in the southeast Pacific provide the first, continuous record of temperate South American vegetation and climate from the last 140 ka. Located at 36°S, 65 km offshore of Concepcion, Chile, Site 1234 monitors the climatic transition zone between northern semi-arid, summer dry-winter wet climate and southern year-round, rainy, cool temperate climate. Dominance of onshore winds suggests that pollen preserved here reflects transport to the ocean via rivers that drain the region and integrate conditions from the coastal mountains to the Andean foothills. Down-hole changes in diagnostic pollen assemblages from xeric lowland deciduous forest (characterized by grasses, herbs, ferns, and trees such as deciduous beech, Nothofagus obliqua), mesic Valdivian Evergreen Forest (including conifers such as the endangered Prumnopitys andina), and Subantarctic Evergreen Rainforest (comprised primarily of southern beech, N. dombeyi) reveal large rapid shifts that likely reflect latitudinal movements in atmospheric circulation and storm tracks associated with the southern westerly winds. During glacial intervals (MIS 2-4, and 6), rainforests and parkland dominated by Nothofagus moved northward into the region. At the MIS 6/5e transition, coeval with the rapid shift to lower isotopic values, rainforest vegetation was rapidly replaced by xeric plant communities associated with Mediterranean-type climate. An increased prominence of halophytic vegetation suggests that MIS 5e was more arid and possibly warmer than MIS 1. Although rainforest pollen rises again at the end of MIS 5e, lowland deciduous forest pollen persists through MIS 5d and 5c, into MIS 5b. Substantial millennial-scale variations occur in both interglacial and glacial regimes, attesting to the sensitivity of the southern westerly belt to climate change. Comparison of the cool, mesic N. dombeyi rainforest assemblage from Site 1234 with δ¹⁸O in the Byrd Ice core shows that on time scales longer than 10 ka, cool-moist conditions in central Chile were coherent with and occurred in phase with Antarctic cooling. This is also likely at millennial scales, although rainforest pollen lags Antarctic cooling with exponential response times of about 1000 years, which plausibly reflects the ecological response time to regional climate change.     

Millennial‐scale climate variability in northwest Patagonia over the last 15 000 yr

A pollen record from Lago Condorito (41°45'S, 73°07'W) shows prominent vegetation and climate changes at millennial time‐scales, superimposed on multimillennial trends in temperature and westerly activity in northwest Patagonia during the past 15 000 yr. The record shows that evergreen temperate rainforests have dominated the landscape over this interval, with floristic changes ranging from cold‐resistant North Patagonian forests with podocarp conifers to Valdivian forests with thermophilous, summer‐drought resistant species. The long‐term trend shows that cool‐temperate and humid conditions prevailed between 15 000 and 11 000 cal. yr BP, followed by an extreme warm and dry phase between 11 000 and 7600 cal. yr BP, and subsequent cooling events and increase in precipitation that peaked at ca. 5000 cal. yr BP, when Southern Hemisphere alpine glaciers achieved their first Neoglacial maximum. Modern conditions were established at ca. 1800 cal. yr BP, following a warm and dry phase between ca. 2900 and 1800 cal. yr BP. These results suggest that millennial‐scale climate variability during deglacial and post‐glacial times also affected the mid‐latitude region of the South Pacific, supporting the idea that changes in the tropical Pacific might be a key factor in the initiation and/or propagation of millennial‐scale climate variability at regional, hemispheric and global scales. Copyright © 2004 John Wiley & Sons, Ltd.     

Late Wisconsin periglacial environments of the southern margin of the Laurentide Ice Sheet reconstructed from pollen analyses

Full‐glacial pollen assemblages from four radiocarbon‐dated interstadial deposits in southwestern Ohio and southeastern Indiana imply the presence of herbaceous vegetation (tundra or muskeg with subarctic indicator Selaginella selaginoides) on the southern margin of the Miami lobe of the Laurentide Ice Sheet ca. 20 000 ¹⁴C yr BP. Scattered Picea (spruce) and possibly Pinus (pine) may have developed regionally ca. 19 000 ¹⁴C yr BP, and ca. 18 000 ¹⁴C yr BP, respectively. Spruce stumps in growth position support a local source of pollen. Prior to the ca. 14 000 ¹⁴C yr BP glacial advance, small amounts of Quercus (oak) and other deciduous pollen suggest development of regional boreal (conifer–hardwood) forests. Copyright © 2002 John Wiley & Sons, Ltd.     

Upper Quaternary vegetation dynamics and palaeoclimatology of the La Chonta bog area (Cordillera de Talamanca,Costa Rica)

Two pollen records from La Chonta bog (2310 m altitude) and one pollen record of a soil profile (2430 m altitude) at a short distance from the bog permit the reconstruction of the vegetational history and climatic sequence of probably the last ca. 80 000 yr of the Cordillera de Talamanca, Costa Rica. The three pollen records can be correlated on the basis of three radiocarbon-dated horizons (> 39 900 yr BP, 34 850 yr BP and 1390 yr BP) and palynostratigraphy. Pollen concentration data provide indirect time control between radiocarbon-dated horizons of the La Chonta 2 core. During the Early Glacial successive fluctuations of the upper forest line occurred between ca. 2300 m and 2800 m altitude and the bog is alternately situated close to the subalpine rainforest belt and in the uppermontane rainforest belt. During the Pleniglacial (ca. 50 000-13 000 yr BP) the forest line fell to ca. 2000 m altitude and paramo vegetation surrounded the La Chonta site. During the Late Glacial the forest line shifted rapidly in ca. 270 yr to 2700–2800 m altitude and the lake became surrounded by Quercus-dominated upper montane rainforest during a period of ca. 1655 yr. A distinct climate cooling (by 2–2.5°C) followed during ca. 580 yr and the upper forest line dropped to ca. 2400 m altitude. This temperature rebound is probably equivalent to the Younger Dryas event. In the depression extensive Alnus carr and swamp vegetation developed. Finally, during the Holocene, the forest line shifted to 3300–3500 m elevation and the lower montane rainforest reached close to ca. 2300 m altitude. In the later part of the Holocene subparamo scrub with Hypericum, Puya, Compositae, Escallonia and Ericaceae largely replaced Alnus carr and the regional and local vegetation closely resembles the presentday composition. During the Last Glacial to Holocene transition a shift of the upper forest line of ca. 1400 m is inferred, corresponding to a glacial temperature depression at 2300 m altitude of ca. 8°C.