Zoophycos and related trace fossils from the Chefar El Ahmar Formation,Upper Emsian-Frasnian Ia-Ib (Ougarta,SW Algeria)

The Upper Emsian to Frasnian Ia-Ib strata of the Marhouma area (or “km 30” outcrop), exposed in the Ougarta Range (SW Algeria) belong to the Chefar El Ahmar Formation. On the basis of distinct lithological and palaeontological features, this formation is subdivided into three members (Lower Marly Limestones Member, Middle Marly Limestones Member, and Upper Marly Limestones Member). The studied beds show low to moderate diversity of trace fossil assemblage which contains thirteen ichnotaxa: Chondrites intricatus, Chondrites isp., Chondrites cf. targionii, Circulichnis cf. montanus, Cochlichnus isp., Neonereites biserialis, Neonereites multiserialis, Nereites isp., Palaeophycus isp., Planolites isp., Thalassinoides isp., Zoophycos aff. cauda-galli, and Zoophycos isp. A. The two latter ichnotaxa are the most common trace fossils in the assemblage and occur at three different levels showing different bioturbation intensities. The first Zoophycos-bearing level (Zl 1) is characterised by an overall high bioturbation intensity reflecting a very high oxygenation rate and nutrient supply, allowing the development of large and dense Zoophycos specimens. The second Zoophycos-bearing level (Zl 2) has a considerable reduction of bioturbation intensity as compared to the previous level, with an abundance of Chondrites, which is probably due to radical palaeoecological changes that suggests dysoxic and stressful conditions. The third Zoophycos-bearing level (Zl 3) is characterised by an overall moderate bioturbation intensity. The distribution of trace fossils was influenced by lithology, sedimentation rate, energy level (storm events), bottom oxygenation, and nutrient supply. The lithofacies and trace fossils of the Chefar El Ahmar Formation both indicate a depositional environment fluctuating from the lower shoreface to lower offshore zone.     

The Faraoni event (latest Hauterivian) in ichnological record: The Río Argos section of southern Spain

The Early Cretaceous (latest Hauterivian) Faraoni Oceanic Anoxic Event (F-OAE) in the Río Argos section (Caravaca region, southern Spain), the candidate for the Global Boundary Stratotype Section and Point (GSSP) of the Hauterivian–Barremian transition, has been studied. Its ichnological bed-by-bed analysis allows for interpretation of oxygenation changes through the Faraoni Level interval and improvement upon previous characterization of oxygen conditions prior, during and after the F-OAE. The trace fossil assemblage belongs to the Zoophycos ichnofacies and it includes Chondrites intricatus, Chondrites targionii, Halimedides isp., Palaeophycus isp., ?Patagonichnus isp., Planolites isp., Rhizocorallium isp., Taenidium isp., Thalassinoides isp., Trichichnus linearis, Zavitokichnus fusiformis, and Zoophycos isp. Their diversity in particular beds fluctuates. Beds with four to six ichnotaxa reflect a multi-tiered macrobenthic trace maker assemblage living in good oxic and trophic conditions. In one bed below and one bed above the Faraoni Level (both marls without primary lamination), there are only two or three, mostly opportunistic ichnotaxa (Trichichnus, Chondrites, Planolites). They record dysoxic conditions. At the base of the Faraoni Level, one thicker and two thinner beds of marly mudstones (21.2 and 3.5 cm thick, respectively) are characterized by primary lamination. At the top and in the basal part of the thicker bed and in the thinner beds some trace fossils are present. These beds were deposited under anoxic conditions and later colonized by trace makers either from overlying beds deposited under oxic conditions or from the level of a greenish lamina in the lower part of the thicker bed, recording short episodes of dysoxic conditions. The thinner anoxic beds are separated by marls deposited under dysoxic conditions.     

东濮凹陷沙三段正常三角洲沉积中遗迹化石及意义

通过对钻井岩心的观察和遗迹化石的鉴定,在东濮凹陷古近纪沙河街组沙三段发现遗迹化石10属15种,包括:Skolithos vertivalis,Skolithos linearis,Skolithos isp.,Palaeophycus tubularis,Palaeophycus isp.,Planolites montanus,Planolites beverlegensis,Planolites isp.,Mermoides isp.,Taenidium isp.,Teichichnus isp.,Helminthoidichnites tenuis,Helminthopsis isp.,Beaconites isp.和Thalassinoides isp.。根据岩性特征、沉积构造和遗迹化石的组成、分布特征,研究区沙三段主要发育正常三角洲前缘亚相、前三角洲亚相,并划分了反映不同沉积环境和水体深度的2种遗迹组合,分别为Palaeophycus-Planolites遗迹组合和Planolites-Helminthoidichnites遗迹组合。     

Exceptionally favourable life conditions for macrobenthos during the Late Cenomanian OAE-2 event: Ichnological record from the Bonarelli Level in the Grajcarek Unit,Polish Carpathians

Pelagic and hemipelagic sediments of the Bonarelli Level (uppermost Cenomanian) in the Sztolnia section (Grajcarek Unit, Polish Carpathians) contain trace fossils of the Zoophycos ichnofacies, including (in descending order of abundance): Chondrites (smaller and larger forms), Planolites, Thalassinoides, Palaeophycus, Taenidium, Teichichnus, and Zoophycos. They occur in thick bioturbated layers, which are interbedded with rare, thin layers of unbioturbated black shales. The black shale layers mark the Bonarelli Level and are interpreted as a record of anoxia or dysoxia. Coeval sections in the Western Tethys contain similar trace fossils but they are less abundant and these sections are characterized by thicker unbioturbated black shale layers and thinner bioturbated layers. This confirms the exceptionally favourable life conditions in sediments of the Sztolnia section, which do not record strong global anoxia during the OAE-2 event. Such favourable conditions were probably caused by effective oxygenation of pore waters and deep burial of organic matter, which are a consequence of high rates of accumulation and the palaeogeographical location of the section on a flank of a submarine high, under strong circulation.     

Lower Devonian tempestites in western Yangtze,South China: insight from Zoophycos ichnofabrics

A Lower Devonian (Emsian) tempestite in the western Yangtze plate, China, was studied based on ichnological and stratigraphical features. The results indicated that the local tempestite can be grouped into clastic constituent types, which include bottom erosion structures, graded beds, swaley cross‐stratification (SCS), parallel lamination, bioturbation and the new composition Zoophycos ichnofabrics. Five storm sequences and three Zoophycos ichnofabrics (Zoophycos–Chondrites ichnofabric, Zoophycos–Chondrites–Thalassinoides ichnofabric and Zoophycos–Thalassinoides–Palaeophycus ichnofabric) were discerned. It is demonstrated that the Zoophycos‐producers from the complex types of Zoophycos in storm sequences were opportunistic organisms (r‐strategists). This study reveals storm‐generated physical and biogenic structures and textures and provides a good ichnological method to identify storm deposits. Copyright © 2013 John Wiley & Sons, Ltd.     

Thalassinoides in the Middle Miocene succession at Siwa Oasis,northwestern Egypt

At Siwa Oasis, the carbonate-rich Middle Miocene Marmarica Formation includes several firmground discontinuity surfaces, characterized by extensive burrow systems. These surfaces are confined to the boundary between the Oasis and Siwa Escarpment members in addition to several intraformational marine firmground discontinuities at several horizons within these two members. The marine trace fossils from all these discontinuities are represented mainly by the ichnogenus Thalassinoides. In all occurrences, Thalassinoides are characterized by a sharp erosional lower contact and grades upward into fossiliferous argillaceous limestone facies. Three Thalassinoides ichnospecies have been reported, including: Th. horizontalis Myrow, 1995, Th. suevicus (Rieth, 1932) and Th. paradoxicus (Woodward, 1830). They are mostly recorded as hypichnial/endichnial forms at the sole of fossiliferous argillaceous limestone facies. Rare exichnial occurrences are also recorded. Thalassinoides ichnospecies displayed considerable variations in their stratigraphic distribution, in addition to remarkable variations in their complexity and dimension. These variations are mostly reflecting environmental conditions prevailed during the trace-maker life. Paleoenvironmental interpretation of these burrows confirmed a well-oxygenated shallow marine environment with extremely slow or nearly nil rate of sedimentation.     

华南泥盆纪遗迹化石及遗迹相

通过对华南泥盆系6条剖面(四川北川甘溪、广西横县六景、广西桂林杨堤、贵州独山大河口—白虎坡、贵州贵阳乌当、四川广元后高坪)的遗迹沉积学系统研究,鉴定和描述遗迹化石15属24种,包括Arenicolites carbonaria,Arenicolites isp.,Chondrites cf.intricatus,Cho...     

Trace fossil assemblages in mixed siliciclastic-carbonate sediments of the Kaladongar Formation (Middle Jurassic), Patcham Island,Kachchh, Western India

The Middle Jurassic rocks of the Kaladongar Formation well exposed in the Kaladongar Hill range of the Patcham Island and Kuar Bet of the Northern Kachchh comprises of ∼450 m thick sequence of mixed siliciclasticcarbonate sediments intercalated with shales. These Mixed siliciclastic-carbonate sediments show wide variation in textural and mineralogical composition and represent genetically related six rock types: micritic sandstone, allochemic sandstone, sandy allochemic limestone, micrtic mudrock, sandy micrite and muddy micrite; which are highly bioturbated and show behaviourally diverse groups of trace fossils. Total 34 ichnogenera are identified, which includes, Arenicolites, Asterosoma, Beaconites, Bergaueria, Chondrites, Cochlichnus, Dactylophycus, Daedalus, Didymaulichnus, Diplocraterion, Gordia, Gyrochorte, Gyrolithes, Ichnocumulus, Laevicyclus, Lockeia, Margaritichnus, Monocraterion, Nereites, Ophiomorpha, Palaeophycus, Phoebichnus, Phycodes, Pilichnus, Planolites, Plug Shaped Form, Protovirgularia, Rhizocorallium, Scolicia, Skolithos, Taenidium, Teichichnus, Thalassinoides and Walcottia. These trace fossils are classified into six morphological groups namely, circular and elliptical structures; simple structures; branched structures; rosette structures; spreiten structures; and winding and meandering structures. These trace fossils are further group into eight assemblages which occurred together into mixed siliciclastic-carbonate sediments, include, Asterosoma assemblage, Gyrochorte assemblage, Rhizocorallium assemblage, Thalassinoides assemblage, Planolites-Palaeophycus assemblage, Phycodes assemblage, Ophiomorpha assemblage and Skolithos assemblage. The recurring pattern of these assemblages through the sequence displays the development of Skolithos and Cruziana ichnofacies and at places the mixed Skolithos-Cruziana ichnofacies which suggest a low wave and current energy conditions with intervening period of high wave and current energy conditions and an intermediate period of stressful environments, respectively. Sedimentological and ichnological data suggest that the deposition of the mixed siliciclastic-carbonate sediments of the Kaladongar Formation took place in the foreshore to offshore environment under fluctuating wave and current energy condition.     

Palaeoenvironmental significance of trace fossils from the palaeocene sediments of the Pondicherry area,South India

Sediments of Manaveli Formation (Palaeocene) are studied from the Thondaimanattam quarry section and Manaveli section of the Pondicherry area, which comprises of bioturbated arenaceous limestone-shale and shaly-sandstone sequences respectively. These sediments consist of high density and low diversity trace fossil genera which includes Lanicodichnus, Ophiomorpha, Planolites, Palaeophycus, Skolithos, Thalassinoides, and Polychaetes tubes. The abundance of horizontal structures of the Thondaimanattam quarry section indicate low to moderate wave and current energy in fully marine subtidal environments while the presences of vertical structures of the Manaveli section indicate moderate to high wave and current energy in the marginal marine environment of deposition for the Palaeocene sediments of the Pondicherry area.     

Upper Cretaceous bottom current deposits,north-east Greenland

Reported ancient bottom current deposits in deep marine settings are scarce and most of them remain contentious. This study describes sedimentological, ichnological and petrographical characteristics of a drill core that covers ca 10 Myr of Upper Cretaceous stratigraphy at Hold with Hope, north-east Greenland. The core is divided into four facies associations, which are interpreted to reflect deposition from bottom currents, turbidity flows and hemipelagic settling in slope and/or near slope environments. The evidence for bottom current influence is three-fold. Firstly, pervasive indications of winnowing such as marine bioclast-rich lags and outsized clasts on ‘mud on mud’ contacts are suggestive of low-sediment concentration flows capable of transporting up to pebble-sized clasts. Common Mn–Fe–Mg rich carbonate matrix cements and various types of hiatal chemogenic lag deposits showing glauconite, apatite and carbonate clasts also point to condensation, prolonged exposure at the sediment–water interface and recurrent phases of sea-floor erosion. Secondly, such deposits can show indicators for tidal processes such as double mud-drapes, tangential bottom sets in dune-scale cross-bedding and cyclic rhythmites. Thirdly, inverse to normal grading at various scales is common in fully marine, commonly seafloor-derived sediments. Ichnological data indicate considerable taxonomic variability in the bottom current deposits, but recurrent fabrics are characteristically dominated by morphologically simple burrows such as Thalassinoides and Planolites, with secondary Phycosiphon, Nereites, Zoophycos and/or Chondrites. In general, opportunistic taxa are common whereas mature composite ichnofabrics are rare. The omission surfaces are locally burrowed with stiffground to firmground trace fossil suites. The results contribute to establishing sedimentological, ichnological and mineralogical criteria for recognition of bottom current deposits as well as to the understanding of the Late Cretaceous palaeoenvironmental evolution of the Arctic region.     

Paleoenvironmental reconstruction of quaternary sedimentation in eastern Sri Lanka: An ichnological study

Quaternary sediment deposits are well exposed along the eastern coastal margin of Sri Lanka. Systematic trace fossil studies along with sediment characteristics have been carried out for the first time to interpret the paleoenvironment that prevailed during Quaternary sedimentation in eastern Sri Lanka. The trace fossils Thalassinoides, Skolithos and Polykladichnus have been recognized about 5m above the present mean sea level. Features of the major trace fossil, Thalassinoides, indicate that it was formed by Thalassinidean shrimp Calianassa. Isolated, vertical to steeply inclined, cylindrical to sub cylindrical tubes with thin internal wall lining and inner wall striations of Skolithos and Polykladichnus show that they were formed by polychaetes. The typical characteristic of Polykladichnus isY-shaped branching with slight enlargement at junctions. Morphology and association of burrows as well as mineralogy, grain size and roundness of sediments indicate that the environment prevailed was marine soft grounds of the intertidal zone. Presence of preserved fine-scale features of burrows indicate that the bioturbated sandy tidal flat was suddenly exposed to the tropical environment due to sea level regression in Quaternary periods. Due to the microbial activities of wall linings, burrows were preserved and it was supported by other microbial activities that produced secondary iron bearing minerals that suffered subsequent oxidation. The bio-origination of red sediments of eastern coastal area of the present study can be used to interpret the debatable origin of the red beds found in northwestern and southeastern coastal areas of the island.     

贵州贵阳地区下三叠统凝缩段的遗迹化石特征

贵阳地区下三叠统是一个完整的碳酸盐岩层序（相当于三级旋回层序）．该层序中的凝缩段由深色薄层（单层厚0．5cm～1cm）的泥晶灰岩和泥质条带灰岩组成．凝缩段中产丰富的遗迹化右．该遗迹群落为：Cochlichnus,Cosmorhaphe,Chondrites,Gordia,Huaxich－nus，Oldhamia,Phycosiphon.Protopaleodictyon,Pinnatpedus,Mammillichnus,Neonerites和生物扰动构造．遗迹化石组合与Nereites遗迹相相似。此外，文本还讨论了此遗迹群落的形成原因。     

Tidal signature recorded in burrow fill

The arrangement of sediment couplets preserved in Thalassinoides shafts suggests that tides regulated the passive filling of these trace fossils and, thus, represent tubular tidalites. The thickness variation in individual layers and couplets implies a mixed diurnal, semi‐diurnal tidal signature where packages of either thick‐layered or thin‐layered couplets alternate. Calcarenitic sediment accumulated when tidal current velocity was too high to allow deposition of mud, whereas a marly mud layer is interpreted to have formed during more tranquil times of a tidal cycle (in particular, low‐tide slack water). The tidal record within the burrows covers a few weeks and the corresponding spring–neap cycles. The fill of the Thalassinoides shafts is the only known record to decipher the tidal signature from otherwise totally bioturbated sediments. These deposits accumulated in a lower‐shoreface to upper‐offshore setting during the late Miocene on a shallow shelf extending from the Atlantic Ocean to the west into northern Patagonia. The fill of all investigated burrows started around spring tide and, thus, the behaviour of the burrow producers – probably crustaceans – is speculated to have been affected by tides or the high water level because all studied burrows became abandoned around the same period of a tidal cycle.     

Ichnofossils from the Miocene Middle Bhuban Formation, Mizoram,Northeast India and their Paleoenvironmental Significance

An ichnofossil assemblage is preserved in sediments of the Middle Bhuban Formation,Surma Group,of Miocene age from the Durtlang and Chanmari sections,Aizawl district,Mizoram,northeastern India.The studied sections comprise a thick,bioturbated and thinly bedded sandstone–silty shale facies.The preserved ichnofossils show high density and low diversity with the assemblage represented by feeding and resting traces,i.e.,Psilonichnus upsilon Frey et al.,1984,Ophiomorpha isp.,Teichichnus spiralis Mikulá.1990,Skolithos isp.,Palaeophycus isp.,and other horizontal burrows.Sedimentary structures associated with these traces are mainly cross bedding,flaser bedding and slump structures.Considering the distribution of the Psilonichnus ichnogenus across an integrated ichnological–sedimentological framework,the stratigraphic interval of the Middle Bhuban Formation,in which they occur,are interpreted to have been deposited under a shallow,marginal-marine channel complex dominated by tidal channels developed in quiet,brackish-water portions of a delta plain.     

First report of trace fossils from Palaeogene succession (Namunagarh grit) of Andaman and Nicobar islands

The Namunagarh grit of Eocene age in South Andaman island is a submarine fan deposit where sedimentation occurred in deep marine environment mainly by turbidity currents. The mudstone and fine-grained sandstone of the turbidite sequence yielded three ichnogenera viz Thalassinoides, Teichichnus and Lorenzinia. Typically shallow marine Thalassinoides burrows co-occur with deep water burrows represented by Teichichnus and Lorenzinia, indicating a dominant control of substrate, availability of food and rate of sedimentation against bathymetry on distribution of trace fossils. Trace fossils are particularly abundant in the mudstones pointing to a strong lithofacies control. A well-oxygenated muddy bottom with a low rate of sedimentation is envisaged. An order in the succession of traces could be deciphered. Simply constructed burrows of Thalassinoides appeared initially followed by complex structures of Teichichnus and ornate Lorenzinia.     

Depositional environments and ichnology of Upper Cretaceous deep-marine deposits in the Sistan Suture Zone,Birjand, Eastern Iran

The palaeoenvironmental significance of trace fossil assemblages in the flysch deposits of the Upper Cretaceous of the Sistan ocean – the Sefidabeh basin in the Sistan Suture Zone SSZ in Eastern Iran – has been assessed for the first time. The Sefidabeh basin of turbidite origin consists of 10 sedimentary facies, which can be grouped into 3 facies associations (FA) representing submarine channel-related facies associations (FA1), lobe-related facies associations (FA2), distal fan-basin floor facies associations of a deep-water turbidite system (FA3). Thirty three ichnogenera, with many ichnospecies, have been identified in this deep sea succession: Alcyonidiopsis, Arthrophycus, Asterostoma, Belorhaphe, Bergaueria, Cardioichnus, Chondrites, Cosmorhaphe, Desmograpton, Gyrophyllites, Halopoa, Helminthopsis, Helminthorhaphe, Laevicyclus, Lophoctenium, Mammilichnis, Megagrapton, Multina, Nereites, Ophiomorpha, Palaeophycus, Planolites, Phycodes, Phycosiphon, Paleodictyon, Rutichnus, Scolicia, ?Strobilorhaphe, Taenidium, Teichichnus, Thalassinoides, Zoophycos and Urohelminthoida. Their distribution is clearly linked with lithofacies and depositional palaeoenvironments. Changes in trace fossil assemblages and ichnocoenoses follow different environments of the turbidity system of the submarine channel to fan system of the Sefidabeh basin and are associated with variations in environmental controlling factors. Environmental controlling factors including hydrodynamic regime, oxygen level, organic content and sedimentation rates. Ten ichnocoenoses were recognized in the facies associations of the deep-sea fan system of this study. Taking into consideration the diversity, bioturbation level, and colonization order of bioturbated beds and the obvious deepening of the deep-sea depositional system from inner to outer parts of the succession, ichnocoenoses can express a bathymetric trend from shallower to deeper parts, and from higher-to-lower hydrodynamic condition of deep-sea fan systems of the Sefidabeh basin. This study reveals important sedimentological and ichnological features of turbiditic systems in deep sea settings of Iran and permits the development of predictive models for the palaeoenvironmental significance of trace fossil assemblages that can be readily translated to analogous depositional systems in the surface/subsurface.     

Geochemical and isotopic characterization of trace fossil infillings: New insights on tracemaker activity after the K/Pg impact event

Geochemical and isotopic analyses of the Cretaceous–Paleogene (K/Pg) boundary deposits were conducted at the Caravaca section (External Subbetic, southeast of Spain) in order to evaluate the recovery of the macrobenthic tracemaker community and the bioturbational disturbance. Samples from the infilling material of several lower Danian dark-colored trace fossils (Chondrites, Planolites, Thalassinoides and Zoophycos) located in the uppermost 8-cm of the light upper Maastrichtian strata, as well as samples from the host sedimentary rock of these trace fossils, were analyzed and compared with data from the lower Danian deposits. The values of element ratios indicative of extraterrestrial contamination (Cr/Al, Co/Al and Ni/Al) are higher in the infilling trace fossil material than in the upper Maastrichtian and lower Danian deposits, which suggests a contribution of the ejecta layer. Regarding the isotope composition, the δ¹³C values are lower in the infilling material than in the Maastrichtian host sedimentary rocks surrounding the traces, while the δ¹⁸O are higher in the infilling material. The geochemical and isotopic compositions of the infilling material evidence the unconsolidated character of the sediment, including the red boundary layer. Softground conditions confirm a relatively rapid recovery by the macrobenthic tracemaker community, starting a few millimeters above the K/Pg boundary layer. The mixture of the infilling material of the trace fossils moreover reveals a significant macrobenthic tracemaker activity affecting K–Pg boundary transition sediments that may have significantly altered original signatures.     

An example of a highly bioturbated,storm‐influenced shoreface deposit: Upper Jurassic Ula Formation,Norwegian North Sea

Integrated ichnological and sedimentological analyses of core samples from the Upper Jurassic Ula Formation in the Norwegian Central Graben were undertaken to quantify the influence of storm waves on sedimentation. Two main facies associations (offshore and shoreface) that form a progradational coarsening upward succession are recognizable within the cores. The offshore deposits are characterized by massive to finely laminated mudstones and fine‐grained sandstones, within a moderately to highly bioturbated complex. The trace fossil assemblage is dominated by deposit‐feeding structures (for example, Planolites, Phycosiphon and Rosselia) and constitutes an expression of the proximal Zoophycos to distal Cruziana ichnofacies. The absence of grazing behaviours and dominance of deposit‐feeding ichnofossils is a reflection of the increased wave energies present (i.e. storm‐generated currents) within an offshore setting. The shoreface succession is represented by highly bioturbated fine‐grained to medium‐grained sandstones, with intervals of planar and trough cross‐bedding, thin pebble lags and bivalve‐rich shell layers. The ichnofossil assemblage, forming part of the Skolithos ichnofacies, is dominated by higher energy Ophiomorpha nodosa ichnofossils and lower energy Ophiomorpha irregulaire and Siphonichnus ichnofossils. The presence of sporadic wave‐generated sedimentary structures and variability in ichnofossil diversity and abundance attests to the influence of storm‐generated currents during deposition. As a whole, the Ula Formation strongly reflects the influence of storm deposits on sediment deposition; consequently, storm‐influenced shoreface most accurately describes these depositional environments.     

The ichnology of carbonate drifts

Carbonate drifts have so far not been as intensely investigated as their siliciclastic equivalents, especially from an ichnological perspective. The aim of this work is therefore to provide an overview of the different bioturbation styles in carbonate drifts for ichnologists and sedimentologists working in such deposits. Different types of carbonate drifts from the Maldives were studied to address this objective. The cores recovered during International Ocean Discovery Program Expedition 359 were examined to provide the sedimentological and ichnological data for a detailed analysis of the ichnology of carbonate drifts. The ichnological characteristics of the Maldives drifts are compared to other carbonate drifts in order to discuss similarities and differences, and thus provide an overview of the general characteristics of carbonate drift ichnology. These drifts are located in the Santaren Channel which lines Great Bahama Bank, along the Marion Plateau in Australia, in the Limassol and Larnaca basins in Cyprus and in the Danish Basin in Denmark. The common characteristics of bioturbation in carbonate drifts are: (i) the complete bioturbation of the sediment with bioturbation indexes between four and six; (ii) the occurrence of distinctive trace fossils limited to facies contacts or condensed intervals; (iii) a typical ichnoassemblage consisting of Thalassinoides, Scolicia, Planolites, Zoophycos, Chondrites, Phycosiphon and Palaeophycus; (iv) the contiguous occurrence of ichnogenera from different tiers, with only Zoophycos and Chondrites as deep tiers; and (v) distinct infills of the traces including particulate organic matter, pyrite, silica and celestine. In addition, the main ichnofacies of carbonate drifts is the Zoophycos ichnofacies. Ichnofabrics grade from coarse‐grained and completely bioturbated to ichnofabrics with present to rare trace fossils and preserved sedimentary structures. The type and intensity of the bioturbation is controlled by the amount of organic matter and the oxygenation at the sea floor that is determined by the action of bottom currents and the sea‐level fluctuations affecting the carbonate factory in carbonate platforms bordering the basins where the carbonate drifts form. The study of the bioturbation in core and outcrop provides palaeoenvironmental information about carbonate‐drift deposits that complement the classical sedimentological data.     

Deep marine trace fossil assemblages from the Lower Carboniferous of Menorca,Balearic Islands,western Mediterranean

Sediments of Lower Carboniferous age in eastern Menorca, Balearic Islands contain a diverse and exceptionally well preserved ichnofauna, including Neonereites biserialis, Nereites isp., Arthrophycus isp., Dictyodora liebeana, two ichnospecies of Chondrites, several ichnospecies of Lophoctenium, two ichnospecies of Phycosiphon, Syncoprulus pharmaceus, annulated burrows and a vertical burrow. The host lithologies are conglomerates, sandstones, siltstones and mudstones; most are the product of debris flows, and of high concentration [Ta(bc) intervals] to relatively dilute (Tcd/Tde intervals) turbidity currents. The rocks are interpreted as deposits of an inner- to mid-fan palaeoenvironment. Channelized deposits, sequences of overbank deposition and interchannel deposits interspersed with the deposits of unconfined debris flows and high concentration turbidity currents are present. The ichnofauna is most frequently, and best, preserved within the Td interval of turbidities, which are interpreted as interchannel deposits, produced by low concentration turbidity flows. The distribution of the ichnofauna is partly controlled by the lithologies in which they are preserved. The rarity and poor preservation of trace fossils in the coarser grained facies contrasts with the detailed preservation of very delicate traces in the finer grained lithologies. However, the ichnofauna is also partitioned between different subenvironments of the mid-fan to produce a series of palaeoichnocoenoses. Thin intervals of interchannel deposits, separated by deposits of high concentration turbidity currents, repeatedly contain only Phycosiphon incertum and small (?juvenile)Dictyodora liebeana. These traces are interpreted as the products of opportunistic colonization of near-channel environments during episodes of quiescent deposition. Thicker intervals of interchannel deposits contain diverse assemblages of trace fossils characteristic of more stable environments, in which widespread colonization occurred. Overbank deposits at Cabo de Favaritz are medium- to thick-bedded, fine-grained beds. In these, the ichnofauna occurs in a simple, two-tier profile. The upper tier is dominated by Nereites isp.; this is underlain by a partially bioturbated layer characterized by large Dictyodora liebeana and Arthrophycus isp.