Intertidal clay-drape couplets (Gironde estuary, France)

Clay-drape couplets on subaqueous dunes have been regarded as a diagnostic feature of the subtidal environment since Visser's seminal paper (1980). The new observation of clay-drape couplets in the intertidal zone on a present day tidal bar of the Gironde estuary shows that they are not restricted to the subtidal zone.
In the intertidal zone, low-tide slack-water clay drapes are deposited in the bottomsets of the dominant current dunes when the muddy water retained in the troughs is absorbed into the sand during the emergence of the intertidal bar. They drape emergence run-off ripples generated by the drainage currents in the bottomsets. High-tide slack-water clay drapes are deposited over the entire dune surface and are preserved on the lee side of the dunes and in the bottomsets. They drape the subordinate current ripples. Low-tide and high-tide slack-water clay drapes enclose one thin rippled sand layer (the subordinate current bundle) and are isolated from other adjacent clay-drape couplets by the dominant current bundle.
The clay-drape couplets deposited in the intertidal zone can be distinguished from their subtidal counterparts on the basis of two morphological differences:
1. In the intertidal zone, the low-tide clay drape is only present in the bottomsets of the dunes, whereas in the subtidal zone equivalent clay drapes are also present on the lower part of the lee side of the dunes.
2. In the intertidal zone, low-tide clay drapes are deposited in the bottomsets of the dunes over emergence run-off ripples oriented in the direction of the drainage currents (i.e. in a direction normal to the tidal currents). Conversely, in the subtidal zone, the equivalent clay drapes are typically deposited over ripples oriented in the tidal-current direction (ebb or flood). There is a difference of polarity of 90° between the intertidal and subtidal small-scale bedforms draped by the low-tide slack-water drapes.     

A palaeoenvironmental interpretation of the Early Proterozoic Malmani dolomite from Zwartkops,South Africa

The Malmani Subgroup northwest of Johannesburg consists of dolomite and chert with only minor clastic sediments.A precise upper intertidal to marginal supratidal analogy and the associated relationship of varied structures suggest that much of the succession represents a tidal flat to intertidal complex formed within differing semiprotected to protected conditions. The dolomites from these environments are recrystallized, reflecting a meteoric influence, and the cherts which are commonly developed within them are also related to prevailing lower pH's. This dolomitization is considered to have been enhanced by the influx of meteoric waters which however resulted in the dolomites having undersaturated iron-manganese ratios. Rare colour-banded dolomites containing columnar stromatolites are thought to represent more steeply shelving intertidal conditions than are normally encountered in the epeiric sea. These dolomites contain quartz crystals rather than chert, suggestive of a lower concentration of silica in the original alkaline solutions. The exposure is part of a very widespread carbonate unit, dated at ca. 2.250 m.y.Subtidal conditions in which large elongate stromatolitic domes developed can be related to a marine transgression across a basal clastic beach sand; and secondly to a progradational tidal flat seawards of which a talus breccia developed on a steepened slope leading down to the subtidal regime. These dolomites formed by interaction with marine waters saturated with respect to iron and manganese, while the absence of chert reflects persisting alkaline conditions.A dark chert-free dolomitic facies with high iron and manganese contents of saturated ratios is considered to have developed in an alkaline lagoonal environment behind a subaqueous bar that is now represented by an overlying thick oolitic unit.The succession contains numerous chert breccias with which shales are associated. The breccias represent subaerial exposure phenomena related to regressions which were followed by periods of short-lived terrigenous influx.     

Eocene tidal sedimentation in the Anambra basin,Southern Nigeria

The Eocene Nanka Formation of the Anambra basin in southern Nigeria consists, in its type area, of four sand subunits each 50–90 m thick, separated by three gypsiferous glauconitic shales each about 2.5 m thick. The sand subunits are unconsolidated, uncemented quartzarenites, planar and trough cross-stratified, flaser and lenticular bedded, and burrowed (Ophiomorpha and Skolithos). Texturally the sands are medium to coarse, moderately sorted, positively skewed and leptokurtic. Two parallel belts, each with a distinctive paleocurrent pattern, are identified: one lying along the present eastern and northern margin of the formation, characterised by a shore-normal paleocurrent pattern; and the other lying on the west, distinguished by a shore-parallel orientation.An integrated analysis of stratigraphic, petrographic, textural, and structural data of the formation enabled reconstruction of an environmental model for the sand body. The model depicts sedimentation in a tidally influenced marine shoreline environment in which an intertidal and a subtidal zone can be delineated. Each zone is characterised by an assemblage of several sedimentary features. Interlaminations of shale and thin sand lenses, gypsiferous and glauconitic shale beds, flaser and lenticular bedding, herringbone structures, and shore-normal paleocurrent pattern are among the features that delineate the intertidal facies. The abundantly cross-stratified, coarse elastic sand-bars with rapid fining-upward sequences, burrowed pebbly horizons overlain by fine sand, and shoreparallel paleocurrent are the distinguishing features of the subtidal facies. The sandflat facies of the intertidal zone and the sandbar facies of the subtidal zone are, however, inseparable in most cases.     

扬子克拉通北缘中元古界神农架群乱石沟组叠层石类型及其沉积学意义*

扬子克拉通北缘神农架地区中元古界神农架群叠层石十分发育。该群下部乱石沟组中叠层石类型丰富,根据形态可分为柱状、穹状、层状叠层石以及锥状、层柱状、包心菜状、墙状叠层石和叠层石丘等。研究区叠层石以中小型为主,少数为大型和巨型;多数无壁,部分具单层壁;对称性和继承性由好到一般,部分较差。研究表明,叠层石的形态和规模,与水动力条件、水体深度及陆源组分含量关系密切,其中层状叠层石主要发育于潮间带上部,柱状叠层石产于潮间带下部和浅潮下带,穹状叠层石出现在潮间带和潮下带,锥状叠层石多见于潮间带下部和潮下带,层柱状、锥柱状叠层石则集中于潮间带,叠层石丘位于潮间带和潮下带。依据各类型叠层石宏观和微观特征、垂向组合类型、沉积构造及沉积岩相相互关系,并结合碳、氧同位素测试结果,综合研究认为: 虽然叠层石的形成受生物和环境等多重因素影响,但生物因素主要影响叠层石的微观组构,水动力条件则控制叠层石的宏观形态类型,水体深度影响叠层石的规模;另外,生物繁盛、陆源物质供给少的环境更有利于叠层石发育。     

冀西北铁岩类型与沉积环境

冀西北铁岩的结构和构造类似于碳酸盐岩,因此,借鉴于现代碳酸盐相模式与沉积环境,有可能恢复古代铁岩的沉积环境。本文对铁岩分类及其沉积环境进行了探讨。     

Community structure and intertidal zonation of the macrobenthos on a macrotidal, ultra-dissipative sandy beach: Summer-winter comparsion

Community structure and intertidal zonation of the macrobenthos on a macrotidal, ultra-dissipative beach were studied. On the beach of De Panne, Belgium, six transects perpendicular to the waterline (each with five stations) were sampled in September 1995 (summer) and March 1996 (winter). The 30 stations were distributed across the continuum from mean high water spring to mean low water spring in order to sample the macrobenthos at different levels of elevation. The 39 species found had total densities up to 5,500 ind m⁻² in summer and 1,400 ind m⁻² in winter. The highest densities were found in the spionid polychaetesScolelepis squamata andSpio filicornis, the nephtyid polychaeteNephtys cirrosa, the cirolanid isopodEurydice pulchra, and the haustorid amphipodsBathyporeia spp. Based on species composition, specific densities, and biomass, two species associations were defined: a relatively species-poor, high intertidal species association, dominated byS. squamata and with an average density of 1,413 ind m⁻² and biomass of 808 mg AFDW m⁻² (summer); and a relatively species-rich, low intertidal species association, dominated byN. cirrosa, and with an average density of 104 ind m⁻² and biomass of 162 mg AFDW m⁻² in summer. For both seasons, the high intertidal species association was restricted in its intertidal distribution between the mean tidal and the mean high-water spring level, whereas the low intertidal species association was found from the mean tidal level to the subtidal. The latter showed good affinities with the subtidalN. cirrosa species association occurring just offshore of De Panne beach, confirming the existence of a relationship between the low intertidal and subtidal macrobenthic species associations. Summer-winter comparison revealed a strong decrease in densities and biomass in the high intertidal zone during winter. Habitat continuity of the low intertidal zone with the subtidal allows subtidal organisms to repopulate the low intertidal zone.     

Upper Jurassic coal-bearing shoreline deposits,Hochstetter Forland,east Greenland

Upper Jurassic coal-bearing yellow sandstones occur at several localities on Hochstetter Forland, northern east Greenland. The sandstones have been divided into eight sedimentary facies, each facies characterized by an assemblage of sedimentary structures, a specific lithology and, occasionally by the content of fossils, notably oysters. The interpretations of the established facies range from high subtidal oyster banks over intertidal beach sand to lagoonal and coastal swamp deposits. The sedimentary facies occur in a vertical sequence where two slightly different facies associations could be discerned. It is suggested that both facies associations have been deposited in a barrier—lagoon coastal area characterized by minor shoreline oscillations. The palaeogeographic setting of the investigated sequence in the fault-controlled basins along the east Greenland continental margin is discussed.     

Subtidal occurrences of ladder-back ripples: their significance in palaeo-environmental reconstruction

Ladder-back ripples are common on modern intertidal flats and have been thought to be diagnostic of deposition in this environment. Recognition of superimposed, asymmetric ladder-back ripples in a subtidal back-barrier estuary and superimposed, symmetric ladder-back ripples in an embayed, shallow marine environment indicates that these features are not diagnostic of deposition in the intertidal zone only, but can occur in other depositional environments, produced by different mechanisms and on different scales. The presence of ladder-back ripples in the subtidal zones challenges aspects of a model used to determine the palaeotidal range of ancient tidal sequences.     

Microbial deposits in the aftermath of the end‐Permian mass extinction: A diverging case from the Mineral Mountains (Utah,USA)

The Lower Triassic Mineral Mountains area (Utah, USA) preserves diversified Smithian and Spathian reefs and bioaccumulations that contain fenestral‐microbialites and various benthic and pelagic organisms. Ecological and environmental changes during the Early Triassic are commonly assumed to be associated with numerous perturbations (productivity changes, acidifica‐tion, redox changes, hypercapnia, eustatism and temperature changes) post‐dating the Permian–Triassic mass extinction. New data acquired in the Mineral Mountains sediments provide evidence to decipher the relationships between depositional environments and the growth and distribution of microbial structures. These data also help to understand better the controlling factors acting upon sedimentation and community turnovers through the Smithian–early Spathian. The studied section records a large‐scale depositional sequence during the Dienerian(?)–Spathian interval. During the transgression, depositional environments evolved from a coastal bay with continental deposits to intertidal fenestral–microbial limestones, shallow subtidal marine sponge–microbial reefs to deep subtidal mud‐dominated limestones. Storm‐induced deposits, microbialite–sponge reefs and shallow subtidal deposits indicate the regression. Three microbialite associations occur in ascending order: (i) a red beds microbialite association deposited in low‐energy hypersaline supratidal conditions where microbialites consist of microbial mats and poorly preserved microbially induced sedimentary structure; (ii) a Smithian microbialite association formed in moderate to high‐energy, tidal conditions where microbialites include stromatolites and associated carbonate grains (oncoids, ooids and peloids); and (iii) a Spathian microbialite association developed in low‐energy offshore conditions that is preserved as multiple decimetre thick isolated domes and coalescent domes. Data indicate that the morphologies of the three microbialite associations are controlled primarily by accommodation, hydrodynamics, bathymetry and grain supply. This study suggests that microbial constructions are controlled by changes between trapping and binding versus precipitation processes in variable hydrodynamic conditions. Due to the presence of numerous metazoans associated with microbialites throughout the Smithian increase in accommodation and Spathian decrease in accommodation, the commonly assumed anachronistic character of the Early Triassic microbialites and the traditional view of prolonged deleterious conditions during the Early Triassic time interval is questioned.     

渤海湾西岸海岸带现代地质作用及影响因素分析

通过对渤海湾西岸粉砂淤泥质海岸带的现代地质作用调查与分析,阐述了渤海湾海岸带在现代地质作用下侵蚀和淤积并存的现状:目前潮间带上部在不断淤高,而潮下带则存在侵蚀,正处于淤蚀过渡期。遥感资料及沉积物粒度的进一步分析,认为渤海湾西岸现代地质作用的影响因素及海岸带淤积泥砂主要来自岸滩下部及浅海区的侵蚀作用。提出在海岸开发中注意防淤的同时,高度重视侵蚀作用可能对海岸带造成的危害。     

Predation Risk Experienced by Mummichog, Fundulus heteroclitus, in Intertidal and Subtidal Salt Marsh Habitats

It is often presumed that salt marshes provide a predation refuge for small fishes, but predation risks have rarely been compared in intertidal and subtidal habitats, making the importance of salt marshes as a predation refuge speculative. We measured relative survival of tethered mummichog (Fundulus heteroclitus) in four habitats in a salt marsh?Ctidal creek system: unvegetated and vegetated intertidal areas and the subtidal creek at high and low tide. At high tide, mummichog in the intertidal zone had significantly higher survival than in the subtidal creek in June through August. Survival rates in unvegetated and vegetated intertidal habitats were not significantly different, suggesting that higher intertidal survival was due to less abundant predators compared with the creek, rather than predators being less effective in vegetation. The lower predation risk experienced by mummichog in the intertidal marsh suggests that access to intertidal habitats will be important for production of small estuarine fishes.     

Sedimentary geology and stromatolites of the Middle Proterozoic Belt Supergroup, Glacier National Park, Montana

The Belt Supergroup is a thick, dominantly fine-grained sequence of Middle Proterozoic strata occurring in western Montana, northern Idaho, and parts of Washington state, Alberta, and British Columbia. The sequence in Glacier National Park is located along the northeastern part of present exposures of the Belt Supergroup; it is 2.9 km thick, extremely well exposed, and for the most part structurally simple. Although it was subjected to lowermost greenschist-facies metamorphism, primary sedimentary structures are exceptionally well preserved.Subtidal, intertidal, alluvial and possibly deltaic depositional environments appear to be represented in the Belt sequence in Glacier National Park. The lowermost unit, the Altyn Limestone, is not entirely exposed in the park. A partial section, 150 m thick, consists of impure dolostones deposited largely in shallow subtidal and intertidal settings. This carbonate unit is overlain by terrigenous strata of the Appekunny and Grinnell Argillites. The Appekunny Argillite is 700 m thick, consists largely of green-colored, fine-grained terrigenous material and appears to have been deposited predominantly in offshore and/or deltaic settings. The overlying Grinnell Argillite is 605 m thick and consists of red-colored terrigenous material deposited largely on an alluvial plain. The overlying Siyeh Limestone is 780 m thick and consists largely of impure dolostones and dolomitic limestones deposited in shallow subtidal and intertidal settings. Overlying the Siyeh Limestone is the 385 m thick Snowslip Formation, which consists of slightly dolomitic, predominantly fine-grained terrigenous strata deposited largely in intertidal settings. The overlying Shepard Formation is not exposed in its entirety in the central part of Glacier National Park. A 270 m thick section, which excludes the uppermost part of the formation, consists of impure dolostones and argillites, and appears to have been deposited in subtidal and intertidal settings.Stromatolites are abundant, diverse and well preserved in Glacier National Park, with mound-shaped forms and columnar forms of the group Baicalia occurring in the Altyn Limestone and Siyeh Limestone, and mound-shaped stromatolite-like structures occurring in the Snowslip and Shepard Formations. Particularly prominent is a 24–32 m thick stromatolite unit in the upper Siyeh Limestone, which contains Baicalia and Conophyton and appears to represent a prograding stromatolite reef, with Baicalia originating in a moderate-energy reef-front setting, and Conophyton originating in a lower energy back-reef setting. Individual units in these cycles can be correlated for 90 km. Many of the Conophyton in these cycles are inclined, probably as a result of gentle wave action, and the direction of inclination is relatively constant for 90 km, with the axes trending SW-SSW and plunging 30–60° SW.     

Depositional environments of Upper Miocene (Messinian) evaporite deposits of the Sicilian Basin*

In Sicily, Messinian evaporitic sedimentary deposits are developed under a wide variety of hypersaline conditions and in environments ranging from continental margin (subaerial), to basin-margin supratidal, to intertidal, to subtidal and out into the hypersaline basin proper. The actual water depth at the time of deposition is indeterminate; however, relative terms such as ‘wave base’ and ‘photic zone’ are utilized. The inter-fingering relationships of specific evaporitic facies having clear and recognizable physical characteristics are presented. These include sub-aerial deposits of nodular calcium sulphate formed displacively within clastic sediments; gypsiferous rudites, arenites and arenitic marls, all of which are reworked sediments and are mixed in varying degrees with other clastic materials (subaerial, supratidal, and intertidal to deep basinal deposits). Laminated calcium sulphate alternating with very thin carbonate interlaminae and having two different aspects; one being even and continuous and the other of a wavy, irregular appearance (subtidal, intertidal, and supratidal deposits). Nodular calcium sulphate beds, usually associated with wavy, irregular laminated beds (supratidal, sabkha deposits); very coarsely crystalline gypsum beds (selenite), associated with more even, laminated beds (subaqueous, intertidal to subtidal deposits); wavy anastomozing gypsum beds, composed of very fine, often broken crystals (subaqueous, current-swept deposits); halite having hopper and chevron structures (supratidal to intertidal); and halite, potash salts, etc. having continuous laminated structure (subaqueous, possibly basinal). Evidence for diagenetic changes is observed in the calcium sulphate deposits which apparently formed by tectonic stress and also by migrating hypersaline waters. These observations suggest that the common, massive form of alabastrine gypsum (or anhydrite, in the subsurface) may not always be ascribed to original depositional features, to syndiagenesis or to early diagenesis but may be the result of late diagenesis.     

A case of normal regression with sea level transgression: Example from the Ganurgarh shale,Vindhyan basin,Maihar area,M.P., India

The Ganurgarh shale, a formation belonging to the Bhander Group of Vindhyan basin is investigated using field based detailed lithofacies and petrofacies analyses in order to interpret the depositional environment in a sequence stratigraphic context. Five major lithofacies have been recognized consisting of calcareous sandstones, laminated mudstones, rippled siltstones, red-grey shales and sandy limestones characterized by small to large-scale cross-bedding, ripple cross-lamination of wave and current origin, parallel lamination, low-angle horizontal bedding, flaser and lenticular bedding, mud-cracks, salt pseudomorphs, convolute bedding and load structures. The constituent lithofacies are recurring and grouped into three lithofacies associations where, the association A is composed of fining upwards and B with coarsening upwards cycles at the lower and middle levels of the succession respectively, are dominantly arenaceous whereas, the association C occurring at upper levels is fining upwards (FU) and becomes calcareous with meager representation of clastics. Petrographically, the section offers three main petrofacies viz., (a) sandstone- (b) siltstone- (c) sandy limestone-petrofacies. Lithofacies characters complimented with petrography show that deposition occurred within the shoreface (subtidal) to foreshore intertidal domain involving tidal flats with sub-environments ranging from intertidal to supratidal. However, lithofacies associations within the Ganurgarh shale of Maihar area represent a case of normal regression during sea level transgression. In the beginning, probably because of excessive sediment supply the sea level had a falling trend during an overall transgressive phase ultimately culminating into limestone sedimentation.     

Tidal flat associations from a Lower Proterozoic carbonate sequence in South Africa

An association of structures is documented from a Lower Proterozoic carbonate sequence in South Africa: this is dominated by finely laminated material which may display flat (smooth), small domical (blister), crinkled (tufted) and pustular (mamillate) surface. The lamination may display overfolded structures and contain flat encapsulated features. Lens-like flat-pebble breccias are common, while edgewise breccias were seen less frequently. Small-scale features suggestive of vertical algal moulds were also recorded. This facies is considered to be analogous to contemporary tidal flat sedimentation developed in specific settings at Shark Bay in Western Australia, and at Abu Dhabi where it is the dominant growth form. This analogy is most notably clear in relation to the protected embayments at Shark Bay, and as a result, an inner intertidal to marginal supratidal environment is suggested for the specific horizons under consideration from the Transvaal Dolomite. Other structures are associated with this facies. These include larger domes, coarser bedding, oolites, ripple-marks and, in one case, columnar stromatolites. Using the basic facies as an environmental datum, a model is developed in which these latter structures extend into the outer intertidal zone. They are also considered to be associated with varying states of turbulence in the intertidal regime.     

Topographic control on distribution of modern microbially induced sedimentary structures (MISS): A case study from Texas coast

Modern back-barrier tidal flats of Galveston Island, Follets Island, and Matagorda Peninsula of the Texas coast are dominated by mud- to fine sand-sized siliciclastic sediments and prolific microbial mats. These microbial mats modify sediment behavior and result in a variety of microbially induced sedimentary structures (MISS). Common structures include: knobby surfaces, reticulated surfaces, gas domes, mat-cracks, sieve-like surfaces, erosional pockets, wrinkles, and mat chips. In general, mat thicknesses increase from ～ 1 mm in the upper supratidal to ～ 3 cm (maximum) in the lower supratidal and then decrease to ～ 2 mm in the lower intertidal areas. This same wedge-shaped pattern is displayed by detailed measurements of mat thicknesses from the rims into the deeper centers of depressions (pools) on the supratidal flats. Measurements of 175 mat-cracks show that height of the curled edges of the mat-crack polygons increases with increase in mat thickness. Similarly, measurements of 150 gas domes reveal that the size of the gas domes also increases with increasing thickness of the associated mats. Because mat thickness varies with elevation on the tidal flat, curl height of the mat-cracks and size of the gas domes are also related to elevation.Six zones were identified based on the occurrence of MISS within the supratidal (zone-I) to upper subtidal (zone-VI) areas. At the highest elevation, knobby surfaces characterize zone-I whereas zone-II is defined by reticulated surfaces. Along with reticulated surfaces, gas domes and mat-cracks characterize zone-III and zone-IV, respectively. Association of sieve-like surfaces with mat-cracks typifies zone-V whereas mat deformation structures and sieve-like surfaces define zone-VI. Boundaries between the MISS-zones in general are parallel and related to tidal zones. The distribution patterns of the MISS-zones are strongly controlled by local topography of the sediment surface because the degree of inundation is the primary controlling factor for the mat growth and resultant MISS. Therefore, distribution of the microbially induced sedimentary structures in siliciclastics, along with the dimension of the mat-cracks and gas domes, can be potentially helpful in interpretation of topography of paleodepositional surfaces.     

冀中坳陷奥陶纪岩相古地理*

碳酸盐岩是冀中坳陷下古生界重要的古潜山油气储集层。研究表明,冀中坳陷奥陶系属于陆表海型台地,进一步划分为潮上带、潮间带、局限潮下和开阔潮下4个沉积亚相,以及泥云坪、膏云坪、云坪和云质潟湖等11种沉积微相,并构成了4个三级层序(SQ1~SQ4),其中SQ4仅发育海侵域。奥陶纪首次海侵期沉积的冶里期(SQ1-TST)在区内由北向南发育了开阔潮下—潮间—潮上亚相沉积。亮甲山期(SQ1-HST)主要发育潮间亚相沉积,局部为潮上亚相。下马家沟期(SQ2-TST)和上马家沟组沉积晚期(SQ3-TST)分别为第2次和第3次海侵,区内均以局限潮下和开阔潮下亚相为主,局部发育生屑滩和砾屑滩;上马家沟组沉积早期(SQ2-HST)和峰峰组沉积早期(SQ3-HST)以发育潮间和潮上亚相为主,是白云岩的稳定沉积期。峰峰组沉积晚期(SQ4-TST)记录了第4次海侵,以发育开阔潮下亚相为主,北部地区零星发育潮间云灰坪微相。受加里东构造运动影响,华北地台整体抬升,冀中坳陷奥陶系顶部受到剥蚀,直至石炭纪再次接受沉积。     

The paleoenvironmental distribution of Phanerozoic wrinkle structures

Wrinkle structures are microbially induced sedimentary structures that are known to have formed in ancient marine siliciclastic environments. This study reviews the sedimentary context and paleoenvironmental distribution of these sedimentary structures throughout three distinct intervals of the Phanerozoic: the Cambrian, the post-Cambrian Paleozoic, and the Lower Triassic. During these times, wrinkle structures are found primarily within two sedimentary environments, storm-dominated subtidal environments and the intertidal zone. Subtidal occurrences of wrinkle structures during the post-Ordovician Phanerozoic only appear to occur during intervals of environmental stress, while earlier examples from the Cambrian and lowermost Ordovician likely formed prior to significant increases in extent and depth of bioturbation that hallmark most of the Phanerozoic. Intertidal examples occur regardless of severe environmental stress, and may have formed under low levels of bioturbation due to inherent vagaries of a marginal marine setting. These environmental preferences appear to be conservative throughout the Phanerozoic, and may very well extend throughout Earth's history.     

The necessity for intertidal foraging by estuarine populations of subadult dungeness crab,Cancer magister: Evidence from a bioenergetics model

Complex intertidal habitats characteristic of northeastern Pacific coastal estuaries provide critical nursery environments for young-of-the-year Dungeness crab,Cancer magister, yet their role in supporting subsequent year classes remains unclear. SubadultC. magister (40–130 mm; 1+ and >1+ year classes), which reach densities as high as 4,300 crabs ha^?1 in subtidal channels during low tides, migrate during flood tides from subtidal refuges into intertidal habitats to forage. As with other brachyuran species that undertake extensive tidally-driven migrations, intertidal foraging may contribute significantly to the energy budget of subadultC. magister. In order to explore the energetic incentive for intertidal migrations by subadult crabs, we developed an ontogenetically-based bioenergetics model for crabs within Willapa Bay, Washington. The model showed that energetic demand varied spatially across the bay, with the highest average energetic demand of a population of subadult crabs (2.13×10⁶ kJ ha^?1) occurring in a habitat stratum termed lower side channel (LSC) and characterized by relatively little subtidal area and extensive intertidal flats. Comparison of model results with subtidal prey production revealed that the latter could not satisfy subadultC. magister energetic demands, especially in LSC where modeled crab predation depleted subtidal prey biomass within 17 simulation days. We estimate that 1 ha of subtidal crabs from LSC would minimally require an additional 1.6 ha of intertidal area to satisfy energetic demands without depleting prey biomass. Our model results support the assertion thatC. magister make regular migrations to forage on productive intertidal flats, and suggest that intertidal foraging may contribute significantly to the diet of subadult crabs in coastal estuaries.     

Characterization of middle Eocene tide-influenced delta: A study from core samples of Hazad Member,Ankleshwar Formation,South Cambay Basin,India

The Hazad Member (Middle Eocene) of the Ankleswar Formation in Cambay Basin, India, is traditionally reported as deltaic system. Present work documents three major facies associations, namely, (i) sandstone-rich upper delta plain (FA-1) deposits, (ii) sandstone-mudstone heterolithic lower delta plain–delta front (FA-2) deposits, and (iii) shale-dominated prodelta (FA-3) deposits, in an overall coarsening-up to fining-up succession. Tidalites are well preserved in FA-2 and are represented by laterally accreted tidal bundles, tidal beddings and vertically accreted tidal rhythmites, described from drill core samples in this communication. Laterally accreted tidal bundles with reactivation surfaces in sand-dominated heterolithic units indicate time-velocity asymmetry in subtidal condition. Tidal beddings and tidal rhythmites in mud-dominated heterolithic units, associated with asymmetric/symmetric ripple forms and desiccation cracks, indicate periodic subaerial emergence in intertidal flat depositional setting. Systematic analysis of the architecture of the tidalites in different parts of the basin signifies rapid shift in sedimentation from subtidal to intertidal flat within the lower delta plain. Transitions from prodeltaic to tidally (subtidal-intertidal) affected delta front to lower delta plain and fluvial-dominated upper delta plain depositional systems attest to high frequency transgressive-regressive cycles in response to changing accommodation, as a result of sea level fluctuations and basinal tectonisms in the Cambay Basin.