The Faraoni event (latest Hauterivian) in ichnological record: The Río Argos section of southern Spain

The Early Cretaceous (latest Hauterivian) Faraoni Oceanic Anoxic Event (F-OAE) in the Río Argos section (Caravaca region, southern Spain), the candidate for the Global Boundary Stratotype Section and Point (GSSP) of the Hauterivian–Barremian transition, has been studied. Its ichnological bed-by-bed analysis allows for interpretation of oxygenation changes through the Faraoni Level interval and improvement upon previous characterization of oxygen conditions prior, during and after the F-OAE. The trace fossil assemblage belongs to the Zoophycos ichnofacies and it includes Chondrites intricatus, Chondrites targionii, Halimedides isp., Palaeophycus isp., ?Patagonichnus isp., Planolites isp., Rhizocorallium isp., Taenidium isp., Thalassinoides isp., Trichichnus linearis, Zavitokichnus fusiformis, and Zoophycos isp. Their diversity in particular beds fluctuates. Beds with four to six ichnotaxa reflect a multi-tiered macrobenthic trace maker assemblage living in good oxic and trophic conditions. In one bed below and one bed above the Faraoni Level (both marls without primary lamination), there are only two or three, mostly opportunistic ichnotaxa (Trichichnus, Chondrites, Planolites). They record dysoxic conditions. At the base of the Faraoni Level, one thicker and two thinner beds of marly mudstones (21.2 and 3.5 cm thick, respectively) are characterized by primary lamination. At the top and in the basal part of the thicker bed and in the thinner beds some trace fossils are present. These beds were deposited under anoxic conditions and later colonized by trace makers either from overlying beds deposited under oxic conditions or from the level of a greenish lamina in the lower part of the thicker bed, recording short episodes of dysoxic conditions. The thinner anoxic beds are separated by marls deposited under dysoxic conditions.     

Flysch Trace Fossils from the Hercynian and IndosinianOrogenic Belts of North western China andTheir Palaeoenvironmental Significance

Twenty-three ichnotaxa have been found in the Silurian and Carboniferous turbidites of the Tianshan orogenic belt and the Triassic turbidites of the East Kunlun-West Qinling orogenic belt of northwestern China. They are Acanthorhaphe isp., ?Arthrophycus isp., Aulichnites parkerensis, Chondrites isp., C. intricatus, C. targionii, Cochlichnus anguineus, coprolite, Dendrotichnium haentzscheli, Helminthopsis isp., Helminthopsis abeli, H. hieroglyphica, H. cf. irregularis, Imbrichnus isp., Kunlunichnus qinghaiensis, Laevicyclus rotaeformis, Lophoctenium tianshanensis, Megagrapton isp., Micatuba verso?, Muensteria isp., Neonereites, Palaeophycus, and Zoophycos caudagalli, two of which, namely, Kunlunichnus qinghaiensis and Lophoctenium tianshanensis, are new ichnospecies. The described trace fossils can be grouped into five ichno-assemblages: the Aulichnites-Imbrichnus ichno-assemblage representing turbidity current deposits of a restricted anoxic deep-sea basin, the Lophoctenium ichno-assemblage showing the deep-sea or ocean environments relevant to a plate subduction, the Zoophycos-Helminthopsis ichno-assemblage representing the upper-middle turbidity fan deposits of a pelagic and hemipelagic environment, the Megagrapton-Chondrites ichno-assemblage representing the middle-lower turbidity fan deposits of an oceanic archipelago and the Kunlunichnus ichno-assemblage indicating bathyal turbidity current deposits.     

Paleoecological significance of the trace fossils of Dhosa Oolite Member (Jumara Formation), Jhura dome,Mainland Kachchh,Western India

Dhosa Oolite Member of the Jumara Formation comprises alternating bands of oolitic limestones and shales, exposed in Jhura dome of Mainland Kachchh, Western India. This sequence is highly bioturbated and exhibits a moderate diversity and behaviourally complex assemblage of ichnospecies. The rhythmically bedded sequence shows three different levels of preservation of traces. Epichnial tiering consists of moderately bioturbated oolitic limestone exhibiting horizontal or low-angle protrusive/retrusive biogenic laminae, commonly dominated by feeding structures like Rhizocorallium jenense, R. irregulare, Zoophycos brianteus and Zoophycos isp. The endichnial structures within the oolitic limestone can be separated into two different preservational trace fossil suites. The endichnial shallow suites consist chiefly of deposit feeders like Chondrites intricatus, C. targionii, Planolites beverleyensis, Taenidium cameronensis, Thalassinoides isp., Z. brianteus, Z. cf circinnatus and Zoophycos isp. and few suspension feeder forms like Palaeophycus tubularis; while endichnial deep suites consist of Chondrites intricatus, Skolithos linearis and Zoophycos isp. Hypichnial structures consists abundant, cylindrical, branched, horizontal, large-sized three dimensional feeding burrows of Thalassinoides isp. and somewhat irregular, obtuse angle ramification burrows of Phycodes isp., which are attached to the lower surface of the casting medium. The trace fossil association indicates Cruziana ichnofacies and abundance of Zoophycus species below the fair weather wave base level is largely a preservational artifact. The preservational processes of the trace fossils indicate soft substrate and diversity and their abundance reflects the other palaeoecological parameters of the open shallow marine environments.     

Cambrian trace fossils of the Cruziana ichnofacies from the Bikaner-Nagaur Basin,north western Indian Craton

The Marwar Supergroup of the Bikaner-Nagaur Basin is composed of sediments deposited from the late Neoproterozoic (Ediacaran) to Upper Cambrian. The Nagaur Sandstone Formation of the Nagaur Group (uppermost division of the Marwar Supergroup) preserves trace fossils significant for establishing Early Cambrian biostratigraphic zones and depositional facies. Fifteen ichnospecies (and eight ichnogenera) identified in the Nagaur Sandstone Formation include “Treptichnus” pedum, Cruziana cf. tenella, Cruziana isp., Diplichnites ispp. A, B, and C, Gyrophyllites isp., Lockeia isp., Merostomichnites isp., Monomorphichnus gregarius isp. nov., Monomorphichnus isp., Planolites isp., Psammichnites isp., Rusophycus bikanerus isp. nov., Rusophycus cf. carbonarius, Rusophycus isp. and radial trace fossils.These trace fossils belong to ethological categories pascichnia, repichnia, cubichnia, and fodinichnia and represent arthropod and worm-like burrowing biota. The assemblage and a regional comparison with contemporaneous trace fossils in the eastern Gondwanan realm suggest that the sequence in the study area belongs to the Cruziana tenella Ichnozone and to Stage 2 (upper part of Terreneuvian), however the Middle Cambrian is not excluded. The trace fossil assemblage belongs to the archetypal Cruziana ichnofacies. Cross bedded sandstone, mud cracks and rainprints in the ichniferous strata of the Nagaur Sandstone Formation indicate deposition in an intertidal sand flat with channels that was exposed episodically.     

The ichnology of carbonate drifts

Carbonate drifts have so far not been as intensely investigated as their siliciclastic equivalents, especially from an ichnological perspective. The aim of this work is therefore to provide an overview of the different bioturbation styles in carbonate drifts for ichnologists and sedimentologists working in such deposits. Different types of carbonate drifts from the Maldives were studied to address this objective. The cores recovered during International Ocean Discovery Program Expedition 359 were examined to provide the sedimentological and ichnological data for a detailed analysis of the ichnology of carbonate drifts. The ichnological characteristics of the Maldives drifts are compared to other carbonate drifts in order to discuss similarities and differences, and thus provide an overview of the general characteristics of carbonate drift ichnology. These drifts are located in the Santaren Channel which lines Great Bahama Bank, along the Marion Plateau in Australia, in the Limassol and Larnaca basins in Cyprus and in the Danish Basin in Denmark. The common characteristics of bioturbation in carbonate drifts are: (i) the complete bioturbation of the sediment with bioturbation indexes between four and six; (ii) the occurrence of distinctive trace fossils limited to facies contacts or condensed intervals; (iii) a typical ichnoassemblage consisting of Thalassinoides, Scolicia, Planolites, Zoophycos, Chondrites, Phycosiphon and Palaeophycus; (iv) the contiguous occurrence of ichnogenera from different tiers, with only Zoophycos and Chondrites as deep tiers; and (v) distinct infills of the traces including particulate organic matter, pyrite, silica and celestine. In addition, the main ichnofacies of carbonate drifts is the Zoophycos ichnofacies. Ichnofabrics grade from coarse‐grained and completely bioturbated to ichnofabrics with present to rare trace fossils and preserved sedimentary structures. The type and intensity of the bioturbation is controlled by the amount of organic matter and the oxygenation at the sea floor that is determined by the action of bottom currents and the sea‐level fluctuations affecting the carbonate factory in carbonate platforms bordering the basins where the carbonate drifts form. The study of the bioturbation in core and outcrop provides palaeoenvironmental information about carbonate‐drift deposits that complement the classical sedimentological data.     

Deep marine trace fossil assemblages from the Lower Carboniferous of Menorca,Balearic Islands,western Mediterranean

Sediments of Lower Carboniferous age in eastern Menorca, Balearic Islands contain a diverse and exceptionally well preserved ichnofauna, including Neonereites biserialis, Nereites isp., Arthrophycus isp., Dictyodora liebeana, two ichnospecies of Chondrites, several ichnospecies of Lophoctenium, two ichnospecies of Phycosiphon, Syncoprulus pharmaceus, annulated burrows and a vertical burrow. The host lithologies are conglomerates, sandstones, siltstones and mudstones; most are the product of debris flows, and of high concentration [Ta(bc) intervals] to relatively dilute (Tcd/Tde intervals) turbidity currents. The rocks are interpreted as deposits of an inner- to mid-fan palaeoenvironment. Channelized deposits, sequences of overbank deposition and interchannel deposits interspersed with the deposits of unconfined debris flows and high concentration turbidity currents are present. The ichnofauna is most frequently, and best, preserved within the Td interval of turbidities, which are interpreted as interchannel deposits, produced by low concentration turbidity flows. The distribution of the ichnofauna is partly controlled by the lithologies in which they are preserved. The rarity and poor preservation of trace fossils in the coarser grained facies contrasts with the detailed preservation of very delicate traces in the finer grained lithologies. However, the ichnofauna is also partitioned between different subenvironments of the mid-fan to produce a series of palaeoichnocoenoses. Thin intervals of interchannel deposits, separated by deposits of high concentration turbidity currents, repeatedly contain only Phycosiphon incertum and small (?juvenile)Dictyodora liebeana. These traces are interpreted as the products of opportunistic colonization of near-channel environments during episodes of quiescent deposition. Thicker intervals of interchannel deposits contain diverse assemblages of trace fossils characteristic of more stable environments, in which widespread colonization occurred. Overbank deposits at Cabo de Favaritz are medium- to thick-bedded, fine-grained beds. In these, the ichnofauna occurs in a simple, two-tier profile. The upper tier is dominated by Nereites isp.; this is underlain by a partially bioturbated layer characterized by large Dictyodora liebeana and Arthrophycus isp.     

新疆准噶尔盆地西北缘石炭纪浊积岩系中遗迹化石的发现及其意义

准噶尔盆地西北缘石炭纪地层主要由碎屑岩组成,沉积厚度5000—7000m,在下石炭统和中石炭统中首次发现了包括浊流在内的重力流沉积和平流岩,大量的深水相遗迹化石。说明盆地西北缘是深水沉积,而非前人普遍认为的属浅水沉积环境。     

Ichnology of prodeltaic hyperpycnite–turbidite channel complexes and lobes from the Upper Cretaceous Prairie Canyon Member of the Mancos Shale,Book Cliffs,Utah, USA

The Upper Cretaceous Prairie Canyon Member of the Mancos Shale, Book Cliffs, Utah, contain outstanding examples of prodeltaic turbidity and hyperpycnal flow deposits. Sandstone‐rich, heterolithic and mudstone‐rich channel fills occur near the north‐west entrance to Tusher Canyon, Gunnison Butte and Bootlegger Wash. Mudstone‐rich and heterolithic‐rich hyperpycnal channel deposits are mostly unbioturbated, locally displaying a few specimens of Phycosiphon incertum, Protovirgularia dichotoma, Rosselia socialis, Schaubcylindrichnus coronus and Palaeophycus tubularis. Sandstone‐rich channel deposits consist of wave‐reworked turbidites and hyperpycnites, containing Helminthoidichnites tenuis, Lockeia siliquaria, Phycodes isp., Phycosiphon incertum, Protovirgularia dichotoma, Rosselia socialis, Skolithos linearis and Fugichnia. Scolicia isp. and Chondrites isp. occur locally. Strata along the south‐west entrance of Tusher Canyon record deposition in a prodelta turbidite lobe, but far from its axis. With the exception of a few specimens of Ophiomorpha isp., bioturbation is restricted to the top of the succession, where Curvolithus simplex, Gyrochorte comosa, Lockeia siliquaria, Palaeophycus tubularis and Ptychoplasma excelsum occur. Strata at Hatch Mesa record deposition in a hyperpycnal lobe, near to its axis. Sandstone beds include Curvolithus simplex, Gyrochorte comosa, Ophiomorpha nodosa, Palaeophycus tubularis, Phycosiphon incertum, Protovirgularia dichotoma, Ptychoplasma excelsum, Schaubcylindrichnus freyi, Skolithos linearis, large specimens of Rosselia socialis and indeterminate crustacean burrows. Chondrites isp. is present in the mudstone. High rates of both episodic and sustained sedimentation, degree of substrate consolidation, freshwater discharge and water turbidity are the most important stress factors in both channels and lobes. Taxonomic composition, uneven distribution of bioturbation through the successions, and overall low ichnodiversity help to distinguish these prodeltaic deposits from bathymetrically equivalent offshore strata in the same basin. Hyperpycnal flow deposits are formed in a wide variety of environmental settings, therefore displaying high ichnological variability. Such variability is summarized by characterising ichnofaunas from four different depositional settings: (i) lakes; (ii) shelf deltas; (iii) shelf‐edge deltas; and (iv) deep‐marine systems.     

Ichnofabric analysis of the Tithonian shallow marine sediments (Bhadasar Formation) Jaisalmer Basin,India

The shallow marine sedimentary sequence of the Jaisalmer Basin exhibits one of the important and well-developed Tithonian sedimentary outcrops for western India. The ichnology and ichnofabric of the lower part of Bhadasar Formation (i.e., Kolar Dongar Member) belonging to Tithonian age are presented and discussed. The Kolar Dongar Member represents a shallow marine succession that contains 16 ichnotaxa: Ancorichnus ancorichnus, Conichnus conicus, Gyrochorte comosa, cf. Jamesonichnites heinbergi, Imponoglyphus kevadiensis, Laevicyclus mongraensis, Monocraterion tentaculatum, Ophiomorpha nodosa, Palaeophycus tubularis, P. bolbiterminus, Phycodes palmatus, Planolites beverleyensis, Rhizocorallium isp., Rosselia rotatus, R. socialis, and Teichichnus rectus. The ichnofabric analysis divulges five distinct ichnofabrics, each typifying distinct depositional environment within shallow marine conditions. The ichnofabric Ophiomorpha 1 with syn-sedimentary faulting exemplifies high energy conditions typical of lower shoreface environment, whereas the Ophiomorpha 2 ichnofabric typifies upper shoreface environment. The Ancorichnus ichnofabric reflects lower offshore condition of deposition. The high ichnodiversity Ancorichnus–Rosselia ichnofabric is indicative of inner shelf conditions, while low ichno-diversity Teichichnus ichnofabric indicates prevalence of low energy brackish bay environment. Thus, Tithonian Kolar Dongar Member indicates depositional environment ranging from shoreface to offshore to inner shelf and finally to brackish bay environment.     

Oxfordian plug-shaped trace fossils from northwestern Algeria

Plug-shaped trace fossils B. perata, B. hemispherica, Bergaueria isp., Calycraterion samsonowiczi, Conostichus ornatus and Conostichus isp. are found in the middle-upper Oxfordian fine-grained sandstone of the Argiles de Saïda Formation (northwestern Algeria). Except for the trace Calycraterion which is interpreted as an annelid trace, these traces correspond to distinct behaviours of sea anemones, and their occurrence within tempestite deposits is influenced by a stress factors in a storm-dominated environment. The abundance of these plug-shaped trace fossils suggests that the upper Jurassic siliciclastic shelves with fine-grained soft clastic substrate were colonised by actinarian sessile benthos.     

Neoichnology and implications for stratigraphy of reworked Upper Oligocene oysters,Antigua, West Indies

The Late Oligocene oyster Hyotissa antiguensis (Brown) is locally common in the Antigua Formation of Hughes Point, eastern Antigua, Lesser Antilles; it was not commonly bored at that time. Its valves and shells are robust, and reworked into the shallow water near-shore environment in Antigua; it could potentially be incorporated into younger rocks. Its neoichnology includes clues that would facilitate identification of these oysters as reworked fossils. The suite of modern borings found in these specimens includes common Caulostrepsis taeniola Clarke, Gastrochaenolites isp. cf. G. turbinatus Kelly and Bromley and Entobia isp., and rare Oichnus simplex Bromley and Rogerella? isp. The latter three taxa are limited to oyster shell substrates. Of the common ichnotaxa, Caulostrepsis and Gastrochaenolites are particularly prominent in limestone clasts and limestone cemented to oyster shells, which would be an indicator of reworking if found in a post-Oligocene lithified deposit. Caulostrepsis and Gastrochaenolites are relatively less common in oyster shells and valves, and in many specimens are seen to terminate against the shell. Entobia is the only common boring limited to the shell substrate. The fidelity of preservation of modern borings is also superior in limestone clasts. This suite of borings is comparable with those found in the Neogene of the Antillean region.     

Pliocene trace fossils from oyster substrates in the Nijar Basin,Betic Cordillera,southern Spain

The Almería-Níjar Basin is a Neogene, intermontane depression marginal to the Mediterranean in southern Spain in the vicinity of El Argamasón, Andalucia. The Pliocene Cuevas Formation rests unconformably on the Upper Messinian rock succession in the Carboneras Fault Zone. The Cuevas Formation is a coarse-grained, bioclastic-rich, calcarenite to calcirudite shoreface deposit. Oysters, namely Saccostrea cucullata (Born), are locally common and preserve a moderate diversity of borings: Caulostrepsis taeniola Clarke; Entobia isp.; Gastrochaenolites isp. aff. G. lapidicus Kelly and Bromley; Maeandropolydora isp. cf. M. sulcans Voigt; Oichnus paraboloides Bromley; and Talpina isp. aff. T. hirsuta Voigt. All represent domiciles except for the predatory O. paraboloides trace. This suite of ichnotaxa is assigned to the Entobia ichnofacies sensu Bromley and Asgaard; they are comparable, particularly, with the Boulder Assemblage of the Pliocene of Rhodes, Greece. Physical disturbance is an important parameter in favouring this pattern of infestation, whether the bored clasts are boulders or oyster valves.     

Ichnofauna from the Harbans Bed of the Badhaura Formation (Sterlitmakian), Rajasthan,India

In the first ever systematic study of trace fossils from the Badhaura Formation, the authors described a nesting burrow, which they ascribed to a stomatopod. The purpose of this paper is two-fold: primarily, to document ichnofauna from (post-glacial marine late Palaeozoic rocks of peninsular India) the Badhaura Formation (Sterlitmakian) representing marine rocks deposited following the Late Palaeozoic glaciation and secondly to contribute to the data on post-glacial ichnofauna from constituent continents of the Gondwanaland. Trace fossils described here are from the Harbans Bed, the topmost lithounit of the Badhaura Formation. The ichnofauna includes Arenicolites tenuis, Beaconites isp., Curvolithus isp., Cylindrichnus concentricus, Didymaulichnus lyelli, Ophiomorpha isp., Palaeophycus tubularis, Planolites beverleyensis, P. montanus, Rosselia chonoides, R. socialis, Skolithos linearis, Taenidium cameronensis, Thalassinoides paradoxicus, Thalassinoides isp. and a flask-shaped brood chamber assigned to a stomatopod crustacean. This mixed assemblage is assigned to distal Skolithos ichnofacies and is suggestive of a period of relatively quiet, shallow water conditions of deposition. The ichnofauna, when viewed in context of peri-gondwanic ichnofaunas, mainly consisting of simple tracks and trails, from late Palaeozoic post-glacial deposits of other Gondwanan continents, is interesting due to dominance of domichnia. Profusion of brood chambers along with Thalassinoides in the Badhaura Formation validates the concept of pre-Mesozoic Thalassinoides being non-decapod in origin and suggestive of adaptive convergence.     

东濮凹陷沙三段正常三角洲沉积中遗迹化石及意义

通过对钻井岩心的观察和遗迹化石的鉴定,在东濮凹陷古近纪沙河街组沙三段发现遗迹化石10属15种,包括:Skolithos vertivalis,Skolithos linearis,Skolithos isp.,Palaeophycus tubularis,Palaeophycus isp.,Planolites montanus,Planolites beverlegensis,Planolites isp.,Mermoides isp.,Taenidium isp.,Teichichnus isp.,Helminthoidichnites tenuis,Helminthopsis isp.,Beaconites isp.和Thalassinoides isp.。根据岩性特征、沉积构造和遗迹化石的组成、分布特征,研究区沙三段主要发育正常三角洲前缘亚相、前三角洲亚相,并划分了反映不同沉积环境和水体深度的2种遗迹组合,分别为Palaeophycus-Planolites遗迹组合和Planolites-Helminthoidichnites遗迹组合。     

豫西登封地区寒武系张夏组中部生物扰动构造及其地质意义

掘穴生物作为生态系统工程的建造者,其对沉积物进行的改造直接影响了沉积物的物理化学性质,进而影响了底栖生态系统。豫西登封地区寒武系苗岭统张夏组中部薄层微晶灰岩中发育大量的生物扰动构造,利用地球生物学、古生物学和生物扰动指数等方法,探讨了生物扰动构造的形成环境及其对沉积物的影响。研究表明,研究区张夏组中部生物扰动构造特征多为扰动边界模糊不清,生物扰动指数（BI）为2 ~ 5,说明该沉积期食物充足,氧气含量高。此外,在生物扰动中识别出3种遗迹化石：Thalassinoides bacae,Planolites isp.和Rhizocorallium isp.,说明生物扰动构造为多种生物造迹形成的。依据寒武系苗岭统张夏组中部的遗迹化石以及沉积特征,可知其整体处于偶尔受到风暴影响的滩间局限台地环境。     

Paleoweathering and Paleoclimatic Reconstructions using Neoproterozoic Paleosol in the Eastern Margin of the Pranhita-Godavari Basin,India

Paleogene sediments of the inner fold belt, Naga hills, have very well preserved ichnofossils. 16 ichnospecies have been documented among 13 ichnogenera such as Arenicolites isp., Chondrites targionii, Cylindrichnus isp., Diplocraterion parallelum, Gyrochorte isp., Ophiomorpha annulata, O. irregulaire, O. nodosa, O. rudis, Palaeophycus tubularis, Planolites beverleyensis, Scolicia palaeobullia, Skolithos linearis, Trypinites weisei, Thalassinoides horizontalis and Zoophycos isp. The ichnofossil assemblages comprise mostly domichnia and fodinichnia benthos of the Skolithos and Cruziana ichnofacies. A shallow marine nearshore to offshore marine environment with fluctuating energy condition has been envisaged.     

Lower Devonian tempestites in western Yangtze,South China: insight from Zoophycos ichnofabrics

A Lower Devonian (Emsian) tempestite in the western Yangtze plate, China, was studied based on ichnological and stratigraphical features. The results indicated that the local tempestite can be grouped into clastic constituent types, which include bottom erosion structures, graded beds, swaley cross‐stratification (SCS), parallel lamination, bioturbation and the new composition Zoophycos ichnofabrics. Five storm sequences and three Zoophycos ichnofabrics (Zoophycos–Chondrites ichnofabric, Zoophycos–Chondrites–Thalassinoides ichnofabric and Zoophycos–Thalassinoides–Palaeophycus ichnofabric) were discerned. It is demonstrated that the Zoophycos‐producers from the complex types of Zoophycos in storm sequences were opportunistic organisms (r‐strategists). This study reveals storm‐generated physical and biogenic structures and textures and provides a good ichnological method to identify storm deposits. Copyright © 2013 John Wiley & Sons, Ltd.     

Exceptionally favourable life conditions for macrobenthos during the Late Cenomanian OAE-2 event: Ichnological record from the Bonarelli Level in the Grajcarek Unit,Polish Carpathians

Pelagic and hemipelagic sediments of the Bonarelli Level (uppermost Cenomanian) in the Sztolnia section (Grajcarek Unit, Polish Carpathians) contain trace fossils of the Zoophycos ichnofacies, including (in descending order of abundance): Chondrites (smaller and larger forms), Planolites, Thalassinoides, Palaeophycus, Taenidium, Teichichnus, and Zoophycos. They occur in thick bioturbated layers, which are interbedded with rare, thin layers of unbioturbated black shales. The black shale layers mark the Bonarelli Level and are interpreted as a record of anoxia or dysoxia. Coeval sections in the Western Tethys contain similar trace fossils but they are less abundant and these sections are characterized by thicker unbioturbated black shale layers and thinner bioturbated layers. This confirms the exceptionally favourable life conditions in sediments of the Sztolnia section, which do not record strong global anoxia during the OAE-2 event. Such favourable conditions were probably caused by effective oxygenation of pore waters and deep burial of organic matter, which are a consequence of high rates of accumulation and the palaeogeographical location of the section on a flank of a submarine high, under strong circulation.     

Early Cambrian Ichnofossils from the Mussoorie Syncline and revision of Trace Fossil Biozonation of the Lesser Himalaya,India

A new locality bearing ichnofossils of the Cruziana Assemblage Zone-Ⅲ from the Mussoorie syncline,Lesser Himalaya,is located in rocks of Member-B of the Dhaulagiri Formation,Tal Group,exposed along the Maldewta-Chhimoli fresh road cut section.The site yielded ichnofossils Bergaueria perata,Cochlichnus anguineus,？Diplocraterion isp.,Dimorphichnus obliquus,diplichnitiform Cruziana bonariensis,Diplichnites gouldi,Glockeria isp.,Helminthopsis isp.,Monomorphichnus lineatus,Phycodes palmatum,Palaeophycus striatus,Planolites beverleyensis,Planolites montanus,Treptichnus cf.T.pedum,scratch marks and an undetermined worm impression.An Early Cambrian age （Cambrian Series 2） is assigned to the ichnofossil-bearing strata based on the stratigraphic position between the Drepanuroides and Palaeoolenus trilobite zones.A revised Cambrian ichnofossil zonation is presented for the Tal Group of the Mussoorie syncline.Together with their occurrence on rippled surfaces,and the lateral displacement of some trackways （due to current action）,a sub-aqueous shallow-marine depositional setting is proposed for the rocks of Member-B.     

新疆柯坪地区志留系柯坪塔格组的遗迹化石

通过对柯坪地区剖面志留系的研究,在柯坪塔格组中已识别出遗迹化石15属20种,它们是：Asterichnus lawrencensis,Cochlichnus anguineus, Cruziana ichnosp., Cruziana qinlingensis, Cruziana rouaulti, Diplichnites bipartibilis,Gordia molassica, Helminthopsis abeli, Helminthopsis isp., Lockeia siliquaria ,Monocraterion cf. teataculatum, Palaeophycus tubularis, Planolites isp.,Planolites montanus, Planolites punctatus, Rusophycus ramellensis,?Scolicia ichnosp., Sagittichnus lincki,Spirophycus bicornis,Taenidium cameronensis。根据这些遗迹化石的组成、产状、分布规律及其围岩的沉积特征,可以划分为2种不同沉积环境条件下的遗迹组合,即(1)Helminthopsis-Gordia遗迹组合, 主要组成分子有Asterichnus, Diplichnites,Gorida, Helminthopsis, Planolites, Rusophycus,?Scolicia和Spirophycus,等,被解释形成于低能的滨外远岸浅海陆棚环境;(2)Cruziana-Rusophycus遗迹组合,常见组成分子有Cochlichnus,Cruziana, Lockeia, Palaeophycus,Rusophycus, Monocraterion,Sagittichnus,和Taenidium等,解释它们形成于滨外近岸浅海陆棚环境。