Modern pollen assemblages from surface lake sediments and their environmental implications on the southwestern Tibetan Plateau

The relationships amongst modern pollen assemblages, vegetation, climate and human activity are the basis for reconstructing palaeoenvironmental changes using pollen records. It is important to determine these relationships at regional scales due to the development of vegetation under different climatic conditions and human activities. In this paper, we report on an analysis of modern pollen assemblages of 31 surface lake samples from 31 lakes (one sample per lake) on the southwestern Tibetan Plateau where the knowledge of modern pollen and their relationships with vegetation, climate and human activities is insufficient. The region includes five vegetation zones: sub‐alpine shrub steppe, alpine steppe, alpine meadow and steppe ecotone, mountain desert and alpine desert. The lakes span a wide range of mean annual precipitation (50–500 mm) and mean annual temperature (?8 to 6 °C). Modern pollen assemblages from our samples mainly consist of herb taxa (Artemisia, Cyperaceae, Poaceae, Chenopodiaceae, etc.) and some tree taxa (Pinus, Fagaceae, Alnus, etc.). The results indicate that modern pollen assemblages are able to reflect the main vegetation distribution. Redundancy analysis for the main pollen types and environmental variables shows that precipitation is the leading factor that influences the pollen distribution in the study area with the first axis capturing 13.7% of the variance in the pollen data set. The Artemisia/Chenopodiaceae ratio is valid for separating the desert component (<2) from the steppe and other vegetation zones (>2) but is unable to distinguish moisture variations. The Artemisia/Cyperaceae ratio is able to identify meadows (<1) and steppes (>1) and can be used as a moisture index on the southwestern Tibetan Plateau. Our results show that an appropriate range is needed for a modern pollen data set in order to perform pollen‐based quantitative climate reconstructions in one region. It is essential to perform modern studies before using pollen ratios to reconstruct palaeovegetation and palaeoclimate at a regional scale.     

青藏高原与黄土高原毗邻地区晚更新世以来孢粉植物群及沉积环境

本文对青藏高原与黄土高原毗邻地区近20个剖面(点)的黄土沉积进行了孢粉分析表明,自晚更新世以来,研究区黄土剖面中古土壤层的孢粉种类、数量以及木本植物花粉成分均比黄土中多得多,反映了该时期研究区的气候条件比现今要温和湿润,植被较繁茂,以后逐渐旱化,但并不连续。此阶段研究区主要植物群的演替规律是:针叶林→森林草原→草原。     

A method to determine warm and cool steppe biomes from pollen data; application to the Mediterranean and Kazakhstan regions

An objective method for the assignment of pollen spectra to appropriate biomes has been published recently. The aim of this paper is to improve the distinction between warm and cool steppes, thus refining vegetation and climate reconstruction, particularly during the Last Glacial Maximum. A set of modern pollen spectra from the Mediterranean and Kazakhstan regions, dominated today by open vegetation types, has been analysed statistically in order to relate pollen taxa abundances to warm and cool grass/shrub plant functional types (PFTs). A statistical test using modern pollen data shows that the method is able to distinguish between cool and warm steppe biomes with a high degree of confidence. The method has been applied to two fossil pollen records. The results of this exercise showed that cool steppe dominated in central Greece between 18 000 and 13 000 yr BP, while in western Iran the vegetation was at the boundary between cool and warm steppes. These vegetation types were replaced by warm mixed forest in Greece and warm steppe in Iran after that time span. © 1998 John Wiley & Sons, Ltd.     

中国黄土高原洛川剖面S5以来的孢粉学记录

中国黄土-古土壤序列记录了距今约2.6 Ma以来的环境变迁。孢粉作为恢复植被演替的敏感指标,被广泛用于古植被重建,但是针对高原中部地区长时间尺度植被演替历史的孢粉学研究还较缺乏。通过对黄土高原洛川剖面S5以来的黄土-古土壤序列开展较高分辨率的孢粉学工作,揭示出该区约0.6 Ma以来的植被演化历史。孢粉谱分析表明,洛川地区0.6 Ma以来以温带草原植被为主,不支持塬区历史时期存在大范围落叶阔叶林的观点。现代黄土高原缺少森林植被主要是由自然原因引起,因此塬区的现代生态恢复应以退耕还草为主。     

黄土高原小流域植被变化和侵蚀产沙的孢粉示踪研究初探

如同137Cs和210Pbex核尘埃,土壤和坝库沉积物中的孢粉也源于大气沉降,是否能用于侵蚀产沙的示踪研究?在陕北吴旗周湾水库采集了一些表层土壤和现代淤地坝、古代聚湫坝沉积泥沙的样品,尝试土地利用/覆被变化和侵蚀产沙的孢粉示踪研究。初步研究表明:草地表土孢粉丰富,平均浓度为26077粒/g;有33个科属的花粉,孢粉组合反映了当地以蒿属为主的半干旱温带草原植被;坡耕地表土的孢粉组合和草地一致,但孢粉平均浓度远低于草地,仅5844粒/g。坝库沉积剖面中,和137Cs核素一样,孢粉也富集于每一洪水沉积旋迴的上部层位,可用于沉积旋迴的区分。现代沉积旋迴的孢粉平均浓度,远低于草地和坡耕地表土,这表明草地和坡耕地土壤不是沉积泥沙的主要来源。古代沉积物的孢粉平均浓度高于现代沉积,表明流域内300年前的植被好于现代,或300年前沟谷地的相对产沙量低于现代。古代孢粉组合中十字花科花粉比例高达36·85%,明显高于现代的0·58%含量,这表明草地退化严重,十字花科植物比例急剧降低。孢粉示踪技术在植被环境恢复,沉积旋迴区分,和泥沙来源确定等方面具有一定的潜力,但应建立针对土地利用/覆被变化和侵蚀产沙研究的孢粉示踪技术方法。     

An 11 000-year record of vegetation and environment from Lago di Martignano,Latium, Italy

A pollen diagram from Lago di Martignano, a maar lake in central Italy, provides an 11000-year record of vegetation and environment change. The earliest pollen spectra are dominated by Artemisia and Gramineae, representing late glacial steppe vegetation typical of the Mediterranean region. Broad-leaved forests were established by ca. 11 000 yr BP. Although Quercus initially dominated their canopy, a wide range of other mesophyllous trees were also present. Pollen values for sclerophyllous tree and shrub taxa characteristic of Mediterranean woodlands and scrub are initially low (<10%). After ca. 7000 yr BP, however, they begin to increase and rise to a peak of >40% of total land pollen at ca. 6700 yr BP, with Olea europaea the single most abundant taxon. Human influence upon the vegetation only becomes significant somewhat after this peak, with progressive clearance of woodland and expansion of herbaceous communities. Castanea sativa and luglans regia pollen is recorded consistently from the beginning of the rise in pollen values for taxa characteristic of Mediterranean scrub communities. Pollen values for arable crops increase progressively after ca. 5500 yr BP, following the peak pollen values for taxa characteristic of Mediterranean scrub vegetation. Late glacial and Holocene climate changes have been complex in this region, with the present character of the climate developing only during the last millennium. Rates of change of pollen spectra peak during this period.     

青藏高原现代表土中冷杉和云杉花粉的空间分布

文章通过对青藏高原北纬25°～45°,东经75°～106°范围内海拔1000～5700m的地区所采集的598个表土样品的孢粉分析,研究了现代表土中冷杉和云杉花粉的分布与植被、气候和海拔之间的关系。表土样品中冷杉和云杉花粉高含量范围对应着青藏高原南部和东部山地暗针叶林分布区,花粉等值线图较准确地反映了冷杉和云杉的丰富程度及其与植被、环境的关系;冷杉和云杉花粉百分含量随海拔和气候梯度变化表现出单峰分布模式,冷杉和云杉花粉百分含量大于20%的最适宜海拔高度是2500～4000m,最高含量中心限制在海拔3200m左右、年均温2～3℃范围内;较高的花粉百分含量带出现在年均温0～8℃、年均降水量400～850mm的区域。结果显示了青藏高原现代冷杉和云杉花粉的空间分布与植被、气候、海拔的空间变化之间很好的相关性,为利用花粉资料进行青藏高原地区的古气候重建提供了有益的资料     

Terrestrial molluscan records of Weichselian Lower to Middle Pleniglacial climatic changes from the Nussloch loess series (Rhine Valley, Germany): the impact of local factors

Two malacological sequences sampled in loess sections P1 and P3 of Nussloch (Rhine Valley, Germany) provide the most complete and precise molluscan record of western Europe for the Weichselian Lower and Middle Pleniglacial from about 70 to 34 cal. kyr BP. Qualitative and statistical analyses were performed on 134 mollusc samples. In the most complete Lower Pleniglacial record (P1), malacofauna changes reflect three short phases of vegetation development and climatic improvement related to soils and probably interstadials. A steppe to herb/shrub tundra shift characterizes the Lower-Middle Pleniglacial transition and is followed in both malacological records by the same general environmental trend (decline in vegetation and humidity increase) ending with a new increase in temperature and vegetation cover at the top of P3. In the Middle Pleniglacial, the impact of each shorter climatic change on the malacofauna is less recognizable due to a higher sediment compaction and also to being differently recorded in both sequences as the local topography affects soil water resources, soil and vegetation development and malacofauna adaptation. A comparison shows that the western European biostratigraphical framework can thus be improved by coupling molluscan records from loess sections to pollen sequences.     

川北若尔盖高原玛曲-红原一带表土中孢粉组合的特征

通过对四川若尔盖高原玛曲—红原一带20个表土孢粉样的分析,研究了该区域表土孢粉组合与现代植被的关系。结果显示,玛曲—红原一带表土的孢粉组合基本反映了现代植被的整体特征,与现代植被在植被类型、植被带的空间排列、主要建群种属和优势种属等方面基本一致。依据孢粉的百分含量,可以划分为亚高山灌丛植被区,亚高山灌丛、亚高山草甸植被区和高寒草甸、草原植被区3个植被带。表土花粉的代表性因种属的不同而各有差异,松属和菊科花粉具超代表性,云杉属花粉的代表性较好,禾本科和莎草科具低代表性,而桦木属则不具有超代表性,可能与桦木属花粉不易搬运有关。3个植被区中的表土孢粉的种类、比例与现代植被均存在一定的差别,这可能与植被退化有关。     

晚更新世以来洛川黑木沟黄土地层中植物硅酸体研究及古植被演替

文中扼要介绍了植物硅酸体在古环境分析中的应用基础,详细研究了洛川黑木沟黄土剖面晚更新世以来植物硅酸体的组合特点、古植被及其演替规律,探讨了植物硅酸体分析在研究高分辨率气候事件及古气候空间格局中的意义。     

柴达木盆地东部36～18kaB.P.期间的孢粉记录及其气候环境

对柴达木盆地察尔汗古贝壳堤剖面的沉积物进行孢粉分析,结果显示这个地区植被与气候在晚更新世中晚期经历以下几个阶段的变化:36.2～31.2kaB.P.(未校正14C年代,下同)期间,以禾本科、藜科、蒿属、莎草科为主,发育草原-草甸植被,气候温和湿润,盘星藻出现较多,反映淡水湖泊,水深在10m左右;31.2～27.6kaB.P.期间,松属、云杉属、桦属等为主的木本植物的含量增加,周围山地森林发育,表明气候温暖,降水量增多。但由于蒸发量大,有效湿度下降,荒漠成分柽柳属等增加,盘星藻在30kaB.P.以后消失,反映湖泊盐度增大;27.6～23.3kaB.P.期间,植被中荒漠成分显著增加,周围山地森林萎缩,气候趋向相对寒冷干旱,湖面积缩小;23.3～18.0kaB.P.,孢粉浓度很低,蒺藜科、藜科等荒漠成分明显增加,植被稀疏,已趋向荒漠化草原,反映气候寒冷干旱。从整个剖面来看,主要的陆生植物孢粉类型为禾本科、柽柳属、蒺藜科、麻黄属、松属、云杉属、柏科、胡桃属和桦属等,藜科和蒿属含量很少,这与柴达木盆地东部地区的表土分析结果完全不同,也与其他草原以及荒漠草原的表土花粉结果相异。这说明晚更新世中晚期柴达木盆地东部地区的植被和现在无法进行比较,气候环境与现在显著不同。     

末次间冰期以来黄土高原南部植被演替的植物硅酸体记录

对于本次间冰期以来黄土高原是否存在大面积森林植被的问题一直存在争议,本文通过对渭南、宝鸡和兰州剖面的植物硅酸体记录研究表明,末次间冰期以来,在黄土高原南部,主要发育草原和森林草原的植被类型,只有在气候最适宜期年降水量超过800mm以上,才可能有短期的森林植被发育。在黄土高原源区发育森林植被所需要的降水量要比基岩山地高250mm以上。     

The importance of plant macrofossils in the reconstruction of Lateglacial vegetation and climate: examples from Scotland,western Norway,and Minnesota,USA

Lateglacial and early Holocene (ca 14–9000 ¹⁴C yr BP; 15–10,000 cal yr BP) pollen records are used to make vegetation and climate reconstructions that are the basis for inferring mechanisms of past climate change and for validating palaeoclimate model simulations. Therefore, it is important that reconstructions from pollen data are realistic and reliable. Two examples of the need for independent validation of pollen interpretations are considered here. First, Lateglacial-interstadial Betula pollen records in northern Scotland and western Norway have been interpreted frequently as reflecting the presence of tree-birch that has strongly influenced the resulting climate reconstructions. However, no associated tree-birch macrofossils have been found so far, and the local dwarf-shrub or open vegetation reconstructed from macrofossil evidence indicates climates too cold for tree-birch establishment. The low local pollen production resulted in the misleadingly high percentage representation of long-distance tree-birch pollen. Second, in the Minnesotan Lateglacial Picea zone, low pollen percentages from thermophilous deciduous trees could derive either from local occurrences of the tree taxa in the Picea/Larix forest or from long-distance dispersal from areas further south. The regionally consistent occurrence of low pollen percentages, even in sites with local tundra vegetation, and the lack of any corresponding macrofossil records support the hypothesis that the trees were not locally present. Macrofossils in the Picea zone represent tundra vegetation or Picea/Larix forest associated with typically boreal taxa, suggesting it was too cold for most thermophilous deciduous trees to grow. Any long-distance tree pollen is not masked by the low pollen production of tundra and Picea and Larix and therefore it is registered relatively strongly in the percentage pollen spectra.Many Lateglacial pollen assemblages have no recognisable modern analogues and contain high representations of well-dispersed ‘indicator’ taxa such as Betula or Artemisia. The spectra could have been derived from vegetation types that do not occur today, perhaps responding to the different climate that resulted from the different balance of climate forcing functions then. However, the available contemporaneous plant-macrofossil assemblages can be readily interpreted in terms of modern vegetation communities, suggesting that the pollen assemblages could have been influenced by mixing of locally produced pollen with long-distance pollen from remote vegetation types that are then over-represented in situations with low local pollen production. In such situations, it is important to validate the climate reconstructions made from the pollen data with a macrofossil record.     

东亚中高纬Heinrich 1事件的表现特征:呼伦湖孢粉记录

Heinrich 1事件是发生于末次冰消期的极端气候突变事件之一,对全球大气环流和陆地生态格局产生了深刻影响.基于对东亚夏季风边缘区最北端呼伦湖 HL08 孔5. 75 m 以上沉积岩芯的 AMS 14C定年技术和 415~275 cm段140个样品的孢粉分析,重建了东亚中高纬地区呼伦湖21500~13000 cal. a B. P.高分辨率植被变化历史,在此基础上揭示了Heinrich 1事件期间呼伦湖区植被响应过程,明确了Heinrich 1事件在东亚中高纬地区的表现特征.结果显示:呼伦湖区Heinrich 1事件发生于16500~15400 cal. a B. P.,以剧烈降温和显著干旱化为表现特征;事件发生期间湖区周围山地发育亚高山草甸,森林植被稀疏;湖盆区域以藜科为主的荒漠草原显著扩张,区域植被盖度降低、生态环境明显恶化;同时,不同植被类型对Heinrich 1事件的响应存在明显差别,亚高山草甸和蒿属为主的典型草原较藜科为主的荒漠草原和桦属为主的落叶阔叶林响应更为快速、敏感.     

Steppe expansion in Patagonia?

Westward expansion of the Patagonian steppe and retrocession of Andean forests due to increasing aridity over the past one or two millennia has been a persistent theme in the ecological and paleoecological literature for at least half a century. New evidence from pollen profiles, tree-ring analysis, vegetation structure, and photographic and documentary historical sources does not show the expansion of the steppe. Instead, over the past century trees have invaded the steppe as a consequence mainly of human-induced changes in the fire regime, and trees have regenerated in forest areas that were heavily burnt at the onset of European colonization.     

Pollen taphonomy in a canyon stream

Surface soil samples from the forested Chuska Mountains to the arid steppe of the Chinle Valley, Northeastern Arizona, show close correlation between modern pollen rain and vegetation. In contrast, modern alluvium is dominated by Pinus pollen throughout the canyon; it reflects neither the surrounding floodplain nor plateau vegetation. Pollen in surface soils is deposited by wind; pollen grains in alluvium are deposited by a stream as sedimentary particles. Clay-size particles correlate significantly with Pinus, Quercus, and Populus pollen. These pollen types settle, as clay does, in slack water. Chenopodiaceae-Amaranthus, Artemisia, other Tubuliflorae, and indeterminate pollen types correlate with sand-size particles, and are deposited by more turbulent water. Fluctuating pollen frequencies in alluvial deposits are related to sedimentology and do not reflect the local or regional vegetation where the sediments were deposited. Alluvial pollen is unreliable for reconstruction of paleoenvironments.     

Vegetation patterns, recent pollen deposition and distribution of non-pollen palynomorphs in a polygon mire near Chokurdakh (NE Yakutia, NE Siberia)

Fine-scaled distribution patterns of vegetation and palynomorphs in a polygon mire in NE Siberia were studied in 31 × 31 plots of 1 m² (vegetation) and along a transect at intervals of 1 m (palynomorphs). Pollen and non-pollen-palynomorph contents clearly reflect differences between polygon ridges and centres. For most single pollen types, however, no unambiguous relation with actual plant abundances and distribution exists. High pollen values are restricted to the immediate vicinity of the pollen sources, which is attributed to the low height of the vegetation. For several plant taxa, high pollen values were found to be restricted to particular plant specimens in the population. High values of pollen ascribed to taxa currently absent at the relevant plot show that the sampled mosses contain a pollen signal of several years. Betula exilis produces two morphologically clearly distinct pollen types. The distribution of selected non-pollen palynomorphs correlates to microrelief, moisture regime and/or trophic conditions. The study shows that single palaeo-sections from arctic peatlands should not be over-interpreted with respect to their palaeoecological value. Local vegetation types and site conditions, however, can be accurately reconstructed from the total assemblages and from combined occurrences of specific palynomorph types.     

Late Quaternary vegetation in the Southwestern Columbia Basin,Washington

A 33,000-yr pollen record from Carp Lake provides information on the vegetation history of the forest/steppe border in the southwestern Columbia Basin. The site is located in the Pinus ponderosa Zone but through much of late Quaternary time the area was probably treeless. Pollen assemblages in sediments dating from 33,000 to 23,500 yr B.P. suggest a period of temperate climate and steppe coinciding with the end of the Olympia Interglaciation. The Fraser Glaciation (ca. 25,000–10,000 yr B.P.) was a period of periglacial steppe or tundra vegetation and conditions too dry and cold to support forests at low altitudes. Aridity is also inferred from the low level of the lake between 21,000 and 8500 yr B.P., and especially after about 13,500 yr B.P. About 10,000 yr B.P. Chenopodiineae and other temperate taxa spread locally, providing palynological evidence for a shift from cold, dry to warm, dry conditions. Pine woodland developed at the site with the onset of humid conditions at 8500 yr B.P.; further cooling is suggested at 4000 yr B.P., when Pseudotsuga and Abies were established locally.     

Ecological significance of common pollen ratios: A review

Pollen ratios have been commonly used to indicate landscape change and climate variation. However, the reliability of these indicators needs to be verified by studies on modern pollen process. Here, we synthesized the major pollen ratios used in previous studies and found that pollen ratios are valuable indicators for the change of vegetation types and climate, e.g., precipitation and moisture. Artemisia/Chenopodiaceae (A/C) ratio could increase from desert to steppe and positively correlate with mean annual precipitation (MAP). Artemisia/Cyperaceae (A/Cy) ratio could be used to identify cool meadow and warm steppe, and it is positively correlated with temperature of July (T_July) and negatively correlated with MAP. Arboreal pollen/nonarboreal pollen (AP/NAP) ratio can be used as a semi-quantitative indicator for landscape and regional precipitation changes. In spite of the significant climatic and environmental implications of the pollen ratios, they were also questioned in some studies under various circumstances and thus caution is needed when using them to indicate climate in different vegetation zones.