Macrobenthic zonation patterns along a morphodynamical continuum of macrotidal,low tide bar/rip and ultra-dissipative sandy beaches

The species composition, densities, biomass and zonation patterns of the macrobenthos of sandy beaches are greatly influenced by the morphodynamics and morphology of the beaches. Macrobenthic zonation patterns along a small-scale morphodynamic gradient, comprising eight Belgian beach sites, were investigated. By taking into account the dimensionless fall velocity (Ω) and the relative tidal range, the beach sites were ordered along the gradient from the ultra-dissipative beach type (UD) to the low tide bar/rip beach type (LTBR). The resulting beach state index varied between 1.8 and 4.2 and the beach profiles were related with the beaches' morphodynamic state.In total 35 macrobenthic species, mainly polychaetes and crustaceans, were encountered, varying between 19 and 23 species per beach site. The species composition was quite similar among beach sites, with Scolelepis squamata being abundant at all eight sites. Furthermore, the macrobenthic distribution patterns were mainly related to elevation at all beach sites. Some remarkable difference in metrics, largely related to the beach morphodynamics and the consequent hydrodynamics, were found. At the hydrodynamically benign and consequently macrobenthos-rich UD beaches, the highest macrobenthic densities and biomass occurred on the upper beach, while at the hydrodynamically harsh and thus macrobenthos-poor LTBR beaches, the maximum densities and biomass occurred lower on the beach. Species, typically occurring on the upper UD beaches, such as Eurydice pulchra, S. squamata, and Bathyporeia sarsi, were restricted to the sub-optimal middle and lower beach zone at LTBR beaches. Only Bathyporeia pilosa was found on the upper beach of both UD and LTBR beaches. The more robust polychaete Ophelia rathkei and the interstitial polychaete Hesionides arenaria were exclusively found in the hydrodynamically harsh conditions of the middle LTBR beach zone.     

The effect of the dominant polychaete Scolelepis squamata on nematode colonisation in sandy beach sediments: An experimental approach

The effect of an abundant sandy beach polychaete, Scolelepis squamata, on the colonisation of defaunated sediments by marine nematodes indicates that sandy beach fauna can be partially controlled by biological interactions within and across size groups. Experimental cores, equipped with windows allowing infaunal colonisation, were filled with defaunated sandy beach sediment containing two different treatments with and without S. squamata. These cores were inserted into microcosms filled with sediment with indigenous meiofauna collected from the field. The treatments were incubated in the laboratory at ambient temperature and salinity for 2, 7, 14 and 21 days, in order to follow the colonisation process of the defaunated sediments by the indigenous nematode fauna over time. Nematodes initially colonised both treatments, with abundances of up to 10% of the densities in the control; after 2 weeks, nematode densities in the cores without S. squamata surpassed the control densities. Nematode assemblages in both treatments were not species rich, and also differed in composition from the natural assemblages. The most successful colonising species, Enoplolaimus litoralis, was rare in the surrounding sediment, suggesting that colonisation was determined by species-specific characteristics such as body size, motility and feeding strategy. Initially the presence of macrofauna did not affect the nematode community composition, but after 2 weeks of the experiment, the presence of the polychaete seemed to facilitate the earlier establishment of non-opportunistic species.     

Macrobenthic community structure of soft-bottom sediments at the Belgian Continental Shelf

Within the frame of different research projects, a large number of sites at the Belgian Continental Shelf (BCS) have been sampled for the macrobenthos between 1994 and 2000. These samples cover a diverse range of habitats: from the sandy beaches to the open sea, from the gullies between the sandbanks to the tops of the sandbanks, and from clay to coarse sandy sediments. To investigate the large-scale spatial distribution of the macrobenthos of the Belgian Continental Shelf, the data of all these research projects—728 samples—were combined and analysed. By means of several multivariate techniques, 10 sample groups with similar macrobenthic assemblage structure were distinguished. Each sample group is found in a particular physico-chemical environment and has a specific species composition. Four sample groups differ drastically, both in habitat and species composition, and are considered to represent four macrobenthic communities: (1) the muddy fine sand Abra alba–Mysella bidentata community is characterized by high densities and diversity; (2) the Nephtys cirrosa community occurs in well-sorted sandy sediments and is characterized by low densities and diversity; (3) very low densities and diversity typify the Ophelia limacina–Glycera lapidum community, which is found in coarse sandy sediments and (4) the Eurydice pulchra–Scolelepis squamata community is typical for the upper intertidal zone of sandy beaches. These macrobenthic communities are not isolated from each other, but are linked through six transitional species assemblages. The transition between the A. alba–M. bidentata community and the N. cirrosa community is characterized by a reduction in the mud content and is dominated by Magelona johnstoni. The transition between the N. cirrosa and the O. limacina–G. lapidum community is distinctive by decreasing densities and coincides with a gradual transition between medium and coarse sandy sediments. From the N. cirrosa to the E. pulchra–S. squamata community, transitional species assemblages related to the transition from the subtidal to the intertidal environment were found. Each community or transitional species assemblages was found over a specific range along the onshore–offshore gradient, four types can be discerned: (1) almost restricted to the near-shore area, but possible wider distribution; (2) distributed over the full onshore–offshore gradient; (3) restricted to the near-shore area and (4) restricted to the sandy beach environment. The diversity pattern on the BCS follows this division, with species rich and poor assemblages in the near-shore area to only species poor assemblages more offshore. The distribution and diversity patterns are linked to the habitat type, distinguished by median grain size and mud content.     

Community structure of macrobenthos in two tropical sandy beaches with different morphodynamic features, Rio de Janeiro, Brazil

The macrobenthos of two exposed tropical sandy beaches in Rio de Janeiro (Brazil) were compared in relation to density, species richness, and vertical zonation. Biological and sediment samplings were carried out in the austral winter of 2002 and the austral summer of 2003. The sampling design consisted of 10 transects perpendicular to the water line, evenly divided into strata. A sampling unit was taken in each stratum with a 0.04 m² quadrat sampler. Beaches were also compared according to physical features, such as slope, wave period, wave height, and grain size. According to Dean's Ω morphodynamic index the Pontal is a dissipative beach while the Costa Azul is a reflective one. The mean grain size ranged from median to coarse sand in Costa Azul, whereas in Pontal it ranged from median to very fine sand. Eleven species were collected in the two beaches. Crustaceans were the dominant in the Costa Azul Beach, while the polychaete Scolelepis squamata dominated the Pontal beach. A negative correlation was found between the density of the macrobenthos and mean grain size, and beach slope. On the other hand, the Dean's parameter correlated positively with faunal density. Based on the results of ANOSIM, in both beaches, two groups of stations were identified, defining an upper and a lower beach zone along the vertical distribution of the macrobenthos.     

Stability of an artificially nourished beach,Balaena Bay,Wellington Harbour,New Zealand

Balaena Bay, Wellington Harbour, New Zealand, has a small pocket beach that was covered originally by pebbles and cobbles. In February and October 1982, the beach was nourished with sandy granular gravel, the stability of which was monitored until February 1984. Although isolated from oceanic swell, the new beach readily responded to locally generated wind waves which induced both northwards and southwards longshore drift. The net effect was erosion of the southern beach, aggradation over the central beach, and minor fluctuations at the northern end. Yet despite this mobility nearly all the nourishment sediment was retained in the littoral zone. Beach volumes, calculated for each survey, varied little and sediment distribution patterns revealed negligable transport of nourishment sediment to adjacent beaches and offshore areas. Stability is further confirmed by compositional data which record no preferential loss of the sandstone, argillite, and quartz components. The only compositional changes were the incorporation into the new beach of small (< 10%) quantities of sediment derived from the old beach surface and from biogenic productivity.     

威海典型海滩沉积地貌及质量评价

选择威海双岛湾附近海滩、国际海水浴场海滩、金海滩、青叽岛以西海滩、青叽岛以东海滩、天鹅湖海滩、桑沟湾海滩、楮岛海滩、南海新区海滩和乳山银滩十处代表性岸滩进行了海滩地貌和底质调查,结合沙滩表层样品分析测试数据,对研究区海滩地貌、沉积物粒度分布特征及海滩质量进行了综合评价。结果表明:威海海滩沉积物主要包括砾砂、粗砂、中砂、细砂四种类型,其中中砂分布最广,约占所有点位的35.6%,细砂、粗砂、砾砂分别占34.4%,26.7%和3.3%。岸滩坡度一般较缓,宽度中等以上,岸滩质量整体较好,部分岸滩受沉积物粒度影响,质量稍差。     

Seasonal persistence of a small southern California beach fill

Torrey Pines State Beach, a site with large seasonal fluctuations in sand level, received a small shoreface beach fill (about 160,000 m³) in April 2001. The 600 m-long, flat-topped nourishment pad extended from a highway riprap revetment seaward about 60 m, terminating in a 2 m-tall vertical scarp. A 2.7 km alongshore span, centered on the nourishment region, was monitored prior to the nourishment and biweekly to monthly for the following 2 years. For the first 7 months after the nourishment, through fall 2001, significant wave heights were small, and the elevated beach fill remained in place, with little change near and above Mean Sea Level (MSL). In contrast, the shoreline accreted on nearby control beaches following a seasonal pattern common in southern California, reducing the elevation difference between the nourished and adjacent beaches. During the first winter storm (3 m significant wave height), the shoreline retreated rapidly over the entire 2.7 km survey reach, forming an alongshore-oriented sandbar in 3 to 4 m water depth [Seymour, R.J., Guza, R.T., O'Reilly, W., Elgar, S., 2004. Rapid erosion of a Southern California beach fill. Coastal Engineering 52 (2), 151–158.]. We show that the winter sandbar, most pronounced offshore of the nourishment, moved back onto the beach face during summer 2002 (following the usual seasonal pattern) and formed a wider beach above MSL at the site of the original nourishment than on the control beaches. Thus, the April 2001 shoreline nourishment was detectable until late fall 2002, persisting locally over a full seasonal cycle. In an extended 7-year time series, total sand volumes (summed between the back beach and 8 m water depth, over the entire 2.7 km reach) exhibit multi-year fluctuations of unknown origin that are twice as large as the nourishment volume.     

The biomass of macro- and interstitial fauna on clean and wrack-covered beaches in Western Australia

The benthic faunal spectrum including bacteria, protozoans, meiofauna, wrack epifauna and macrofauna, was quantitatively surveyed on two modally reflective, moderate energy, Western Australian beaches. The more exposed beach had coarser sand, no intertidal macrofauna and a poor interstitial fauna. The less exposed beach had a large deposit of wrack totalling 161 kg m^?1 dry mass concentrated on the lower shore. The amphipod Allorchestes compressa was abundant in the fresh wrack comprising most of the macrofauna. There were also fairly abundant small epifauna on the wrack. Dry biomass of macrofauna, epifauna, meiofauna, protozoans and bacteria was 0, 0, 15, 4 and 180 g m^?1 on the more exposed beach and 160, 3, 112, 9 and 901 g m^?1 on the less exposed beach with wrack. On the latter beach there was an inverse correlation between meiofaunal densities and the densities of protozoans and bacteria, suggesting grazing by the former on the latter. On both beaches meiofauna was concentrated in the mid- to upper beach, protozoans near the surface and bacteria in the mid- to lower beach. It is estimated that bacteria are responsible for most of the secondary production on both beaches.     

Exposed sandy beaches as semi-closed ecosystems

On the basis of various lines of evidence, mostly coming from exposed sandy beaches in Southern Africa, it is postulated that under certain conditions high energy sandy beaches and their adjacent surf zones may function as viable ecosystems. Where surf zones are reasonably broad and shallow, cellular circulation patterns predominate and these tend to retain nutrients generated by the macrofauna and interstitial fauna of the beach. These nutrients may then cause blooms of surf zone phytoplankton which in turn serve as food for macrofauna filter feeders. With the perimeter of the circulation cells of the surf zone forming its marine boundary, the beach and surf zone may together be considered an ecosystem with surf phytoplankton the primary producers, beach macrofauna the consumers and interstitial fauna the decomposers.     

台风季节朱家尖岛砂、砾质岬湾海滩的不同沉积地貌动态变化

岬湾砂、砾质海岸是海岛、海岸带重要的旅游资源, 具有较高的社会经济和生态价值, 长期以来备受关注。本文以浙江朱家尖岛东岸的5个代表性岬湾海滩为例, 基于2019年台风季节早期、中期及晚期测量获得的海滩地形和沉积物数据, 结合海滩近岸的水动力数据, 分析了砂质海滩和砾石海滩这两种不同类型岬湾海滩的沉积地貌动态变化。结果表明, 朱家尖岛东岸5个海滩在台风季节出现了不同的体积变化, 大沙里、东沙、千沙、乌石塘和小乌石塘海滩的单宽体积变化分别为11.93m³·m^-1、-54.41m³·m^-1、-19.75m³·m^-1、2.19m³·m^-1和-1.96m³·m^-1。砾石滩较砂质海滩更为稳定, 无人类活动干扰的砂质海滩在台风季节侵蚀更少、变化更小。台风季节大沙里、东沙、千沙、乌石塘和小乌石塘海滩表层沉积物的平均粒径分别为2.47Փ、2.24Փ、2.64Փ、-5.96Փ和-6.03Փ, 粒径粗化和离岸输运是5个海滩表层沉积物在台风季节的主要表现, 砂质海滩的沉积物粒度特征变化比砾石海滩要大。沉积物粒径、台风强度及台风期间的主要波向与海滩走向之间的关系、海岸工程这3种因素都可能对海滩在台风季节的沉积地貌动态变化产生影响。本文研究结果可为台风季节的海滩管理提供参考。     

Meiobenthic diversity in space and time: The case of harpacticoid copepods in two Mediterranean microtidal sandy beaches

Meiobenthic data from two microtidal sandy beaches of the eastern Mediterranean (Crete, Greece) were used to investigate patterns of both alpha and beta diversity in space and time. Copepod assemblages and environmental variables related to sediment characteristics, morphodynamics and food were studied over a year at four distinct habitats at each beach; the retention, resurgence and saturation zones of Salvat's intertidal scheme (midlittoral zone), and the surf zone of the sublittoral. Αlpha diversity analysis indicated similar species richness at both beaches when the whole 13-month data set was considered but was higher at the sheltered site when each sampling period was examined separately. Both beaches supported higher diversity in the sublittoral zone. Species richness increased seawards at the midlittoral zone of the sheltered site whereas, no pattern was evident at the exposed site, where the intense hydrodynamic conditions homogenized the sediments. Beta diversity increased markedly towards the sublittoral, indicating greater differences in alpha diversity between the sublittoral and the midlittoral zone. Species turnover was more variable at the exposed beach and at the most landward stations, where environmental conditions change often between extremes. A proportion of the variation in alpha diversity was explained by food availability at both beaches and additionally by grain size at the sheltered site. However, no environmental variable explained beta diversity patterns. Although the results of our study support the hypothesis of Multicausal Environmental Severity proposed for sandy beach macrofauna, we believe the classic Intermediate Disturbance Hypothesis is a more appropriate framework for the meiofauna communities of the studied sites.     

Structure and composition of the consolidated mud tube of Maldane sarsi (Polychaeta: Maldanidae)

Maldanid polychaetes can be important components of marine benthic communities, playing significant roles in particle subduction or sediment irrigation. Many maldanids are known to inhabit tubes consisting of sediments consolidated by mucus; the structure and composition of these tubes, and their potential impact on benthic environments, are poorly known. We examined the three-dimensional organization of Maldane sarsi tubes, using CT scanning together with analyses of sediment grain size and concentrations of Fe, Mn, organic carbon and bacteria in tube material. M. sarsi tubes consist of stacks of individual consolidated mud disks, surrounding a dense, continuous, inner tube. The tubes of M. sarsi contained fewer fine particles than surrounding sediments, and greater concentrations of Fe, Mn, organic carbon and bacteria, especially in the inner zone. These distributions suggest that tube irrigation affects Fe and Mn oxidation and precipitation in a narrow zone surrounding M. sarsi, and that mucous secretion and potential feeding activities (the hoeing of surface sediments) lead to increases in organic carbon and bacteria in the inner, and deepest parts of the tube. The finding of relict tubes, buried at up to 15 cm depth, indicates a relatively high longevity for these structures and suggests a potential importance in biogeochemical cycling.     

人工海滩研究进展

海滩是激浪作用下泥沙在激浪带的堆积,是海岸带最活跃的地貌单元。近年来,由于世纪性的海平面上升,侵蚀型海滩范围扩大,侵蚀加重,海滩宽度变窄、坡度变陡和滩面物质粗化等问题日益严重,引起了广泛关注,亦成为海岸工程研究的热点课题之一。几十年的工程实践表明,海滩喂养与人工海滩是当前防护海滩侵蚀最有效的措施,并已为欧、美、日等国广泛应用。利用海岸地貌学原理方法,借鉴模拟环境条件类似的周边自然海滩的形成和演变规律,合理的设计海滩物质的颗粒大小和组成,海滩的坡度、高度、宽度以及填砂范围等关键参数,能够有效地使人工海滩更快的达到平衡状态并维持其稳定。在海滩的娱乐休闲功能在国际范围内日益受到重视的今天,这一方法在海岸工程应用上具有重要的现实意义。     

深圳龙岐湾海滩资源调查与开发建议

文章基于2022年春、秋季开展的龙岐湾沙滩资源调查数据,分析较场尾沙滩、大塘角沙滩、黄泥湾沙滩和桔钓沙沙滩的剖面特征、沉积物粒径和裂流发生情况。研究结果表明：4处沙滩剖面的坡度秋缓春陡,剖面受风浪和潮汐直接作用时坡度变化大,受遮蔽时坡度变化小,整体呈冬春淤积、夏秋侵蚀;沙滩沉积物以中砂、粗砂和砾石为主,粒径变化不大;沙滩均无明显裂流发生,为低风险等级。建议开发沙滩旅游应考虑季节性的补沙方案,开展围垦养殖和海岸工程应注意削弱ESE向和SE向的波浪作用。     

An interpretation of wave refraction and its influence on foreshore sediment distribution

To analyze the grain size and depositional environment of the foreshore sediments, a study was undertaken on wave refraction along the wide sandy beaches of central Tamil Nadu coast. The nearshore waves approach the coast at 45° during the northeast(NE) monsoon, at 135° during the southwest(SW) monsoon and at 90° during the non-monsoon or fair-weather period with a predominant wave period of 8 and 10 s. A computer based wave refraction pattern is constructed to evaluate the trajectories of shoreward propagating waves along the coast in different seasons. The convergent wave rays during NE monsoon, leads to high energy wave condition which conveys a continuous erosion at foreshore region while divergent and inept condition of rays during the SW and non-monsoon, leads to moderate and less energy waves that clearly demarcates the rebuilt beach sediments through littoral sediment transport. The role of wave refraction in foreshore deposits was understood by grain size and depositional environment analysis. The presence of fine grains with the mixed population, during the NE monsoon reveals that the high energy wave condition and sediments were derived from beach and river environment. Conversely, the presence of medium grains with uniform population, during SW and non-monsoon attested less turbulence and sediments were derived from prolong propagation of onshore-offshore wave process.These upshots are apparently correlated with the in situ beach condition. On the whole, from this study it is understood that beaches underwent erosion during the NE monsoon and restored its original condition during the SW and non-monsoon seasons that exposed the stability of the beach and nearshore condition.     

The sandy beach macrofauna of Gulf of Gabès (Tunisia)

This study describes the macrofauna of the three beaches situated in central Gulf of Gabès (Tunisia): Ouderef, Gabès and Zarrat. The Gulf of Gabès has the largest tides in the Mediterranean and the beaches showed a wide intertidal zone. The beaches were sampled once during the spring low tides of June 2005. A transect was extended at each beach, from above the drift line to below the swash line at five sampled levels; at each level six 0.05 m² replicates were taken to a depth of 30 cm and sieved through a 1-mm mesh, and the organisms collected and preserved. The three beaches showed a different physical environment. Sediment type was medium sand at the steeper Ouderef beach, fine sands at Gàbes beach, and very fine sands at the flatter Zarrat beach. The total number of species collected was 31: 12 crustaceans, 10 polychaetes, four molluscs and five insects. The supralittoral and mediolittoral zones were very different. The supralittoral zone was dominated by Talitrus saltator and insects. The most abundant mediolittoral species were the amphipod crustacean Bathyporeia guilliamsoniana at Ouderef beach (23069 ind.m⁻¹), the surf clam Donax trunculus at Gabès beach (60711 ind.m⁻¹) and the spionid polychaete Scolelepis mesnili at Zarrat beach (18345.6 ind.m⁻¹).     

Multigrain sedimentation/erosion model based on cross-shore equilibrium sediment distribution: Application to nourishment design

In the light of global warming and sea level rise there are many coastal beaches that suffer from erosion. Beach nourishment has become a common practice to maintain the sediment balance on a shore-face. In this paper, a three-dimensional numerical model for evaluating long-term impact of beach nourishment projects has been developed. The model addresses the longstanding complex issue of coastal morphology and sediment grain size distribution from an unconventional angle, which exploits the strong links between grain size distribution and the prevailing transport direction of each sediment constituent under ‘average’ wave and storm action. The present model predicts the redistribution of nourished sediment according to the subtle clues implied by equilibrium distribution curves and latest coastal wave transformation theories. After verification against recent field observations in Terschelling, The Netherlands, the model was used to predict long-term effects of different beach nourishment strategies. It was found that: (a) given the source sediment available in Terschelling the tactics of large volume and less frequent implementation are better than otherwise; and (b) from a pure engineering point of view, waterline nourishment outperforms offshore trough nourishment.The model offers an additional tool for coastal engineers to evaluate the feasibility, effectiveness and the optimization of dumping locations for beach nourishment projects. It is also a useful tool for stratigraphic modelling of shallow-marine sedimentation in conjunction with sea level changes.