Lower Bathonian Belemnites and Biostratigraphy of the Central and Southern Parts of the East European Platform: Part 2. Cylindroteuthididae and Belemnotheutididae

Belemnites from the lower Bathonian deposits of the Russian Plate are revised and discussed on the basis of study of the two reference sections—Pletnyovka and Sokur quarries. This is the second part of the investigation dealing with representatives of the family Cylindroteuthididae, and “rostrum-less” belemnites of the family Belemnotheutididae, while members of the family Megateuthididae were discussed in detail in the first part (Ippolitov, 2018). The cylindroteuthidids are found only in the Sokur Quarry and comprise four species all assigned to the genus Pachyteuthis, including a new species, Pachyteuthis mittai sp. nov. For Belemnotheutididae, the presence of a single species Acanthoteuthis foliorostris sp. nov. is recorded, which is the first confirmed record of the genus Acanthoteuthis in the pre-Callovian strata. The study of morphological changes within Cylindroteuthididae across the succession allowed two biohorizons to be recognized and the existing belemnite-based scheme for the lower Bathonian of the Volga Region and its correlation with sections in the Pechora River basin to be updated. As the result of the revision, a complex zonal and infrazonal belemnite-based stratigraphic scheme of the lower Bathonian of the Volga Region is introduced. The presence of an endemic genus deriving from Arctic-Boreal immigrants in the lower Bathonian of the Middle Russian Sea allows the belemnite fauna of this age to be interpreted as the first isolated episode of differentiation of the Boreal-Atlantic Province in belemnites. The study of the development of the cephalopod fauna in the early Bathonian of the Russian Plate indicates a biphase formation of the meridional strait connecting the Middle Russian Sea with the Arctic basins and also supports the hypothesis of a short-term opening of a sublatitudinal Pripyat Strait during the maximum highstand.     

Late Carboniferous lycopsids of the Karantrav locality (the Southern Urals)

Numerous plant remains as the imprints of lycopsid strobili and stems and the sporadic imprints of articulate stems were first found in the Upper Carboniferous deposits from the Western Slope of the Southern Urals (Karantrav Settlement area). The brief stratigraphic characteristic of the studied locality is given. The studied plant assemblage is represented by Lepidodendron ophiurus Brongniart, L. vaselgense Anikeeva et O. Orlova, sp. nov., Lepidodendron sp., Lepidostrobus tevelevii O. Orlova, Mamontov et Anikeeva, sp. nov., L. ronnaensis Bek et Oplustil, Knorria sp., Calamites sp., and Mesocalamites ramifer (Stur) Hirmer. In-situ microspores of the Lycospora type were discovered in the sporangia of some strobili of the genus Lepidostrobus. Four lycopsid species (two are new) were described.     

Smaller foraminifers from type sections of the Bolorian stage,the Lower Permian of Darvaz

Smaller foraminifers from Upper Yakhtashian and Bolorian deposits of the stratotype area (Pamir, Darvaz, Tajikistan) are investigated. Four assemblages are defined. The first assemblage is from Chalaroschwagerina vulgaris-Pamirina darvasica Zone. The second assemblage found in the transitional Yakhtashian-Bolorian beds includes Globivalvulina, Palaeotextulariidae, Hemigordiidae, and Glomospira, associated with the first Pachyphloia and Langella forms. Characteristic taxa of third assemblage from the Misellina (Brevaxina) dyhrenfurthi Zone are Geinitzinidae, Globivalvulina, Palaeotextulariidae, Glomospira, and rare Pachyphloia. The forth assemblage of Hemigordiidae, Pachyphloia, Palaeotextulariidae, Geinitzinidae, Pseudoagathammina is identified in the M. (Misellina) parvicostata Zone. The assemblages were compared with concurrent analogs from China, Japan, and Russia. New species and subspecies Glomospira paleograndis sp. nov., G. darvasica sp. nov., Agathammina darvasica sp. nov., Pachyphloia darvasica sp. nov., Nodosinelloides cubanicus elongatus subsp. nov., and Hemigordius saranensis darvasicus subsp. nov. are described.     

Campanian-Lower Paleogene gastropods from the Kharga Oasis,south Western Desert,Egypt

The Campanian-Lower Paleogene successions exposed at the Kharga Oasis contain well-preserved gastropods. Nineteen gastropod species are identified and described in detail. Two new species are established: Gisortia khargaensis from the lower Eocene El Rufuf Formation and Ornopsis? barisensis from the lower Paleocene Kurkur Formation. In addition to, Turritella (Torquesia) derbesi Coquand is recorded herein for the first time from Egypt. The studied gastropods belonging to four clades: Sorbeoconcha (Metacerithium abictiforme (Wanner), Mesalia hofana Mayer-Eymar, Mesalia sp., Turritella (Torquesia) derbesi Coquand, Turritella (Torquesia) forgemoli Coquand, Turritella kaserensis Hewaidy and Azab, Turritella sp.), Littorinimorpha (Cypraedia bullina Oppenheim, Gisortia khargaensis nov. sp., Natica farafrensis Wanner, Aporrhais schweinfurthi (Quaas), Drepanocheilus bicarinatus Abbass, Drepanocheilus? sp., Anchura? unicarinatus (Abbass), Epitonium sp.), Neogastropoda (Ornopsis? barisensis nov. sp., Rostellana daniensis (Quaas), Vouta sp.), and Heterobranchia (Avellana cretacea Quaas).     

Ammonites and Stratigraphy of the Terminal Part of the Middle Volgian Substage (Upper Jurassic; Epivirgatites nikitini Zone and Its Equivalents) of the Panboreal Realm: 2. <Emphasis Type="Italic">Titanites</Emphasis> and <Emphasis Type="Italic">Glaucolithites</Emphasis>

The genera Titanites and Glaucolithites, which were previously known only from the Portlandian of Northwestern Europe are now established in Epivirgatites nikitini Zone of European Russia. The Volgian species of Titanites are represented by both the endemic taxa Titanites (Paratitanites) manipulocostatus subgen. et sp. nov. and T. (Pseudogalbanites) triceps subgen. et sp. nov. and Titanites (Titanites) ex gr. titan Buckman of the Portland type. A new species Glaucolithites gardarikensis is described. It is likely that these ammonites migrated to the Central Russian Sea from northwestern Europe via the Norway-Greenland seaway and further to the Mezen-Pechora system of straits, which is supported by occurrences of Titanites in the north of Central Russia and Glaucolithites in East Greenland and on Spitsbergen. The direct and indirect correlations show that the middle part of the Nikitini Zone (Lahuseni Subzone) should at least partly be correlated with the Portlandian Kerbeus Zone.     

Marine Early Bajocian Deposits of the Lower Volga Region (Volgograd Region) and Their Belemnite-Based Stratigraphy

A representative early Bajocian belemnite assemblage containing four genera and 11 species (two identified in open nomenclature and five new: Eocylindroteuthis weisi sp. nov., E. mariottii sp. nov., E. yarkovi sp. nov., Homaloteuthis volgogradensis sp. nov., and Hastites orphana sp. nov.) is described from a section near the Dubovoi hamlet in the Greater Don Bend area (southern termination of the Don-Medveditsa dislocations). Some members of this assemblage were previously known from the Caucasus; however, it is possible to state the discovery of a Euroboreal belemnite fauna completely new for Russia, previously described only from Central Europe and almost unknown for Eastern Europe. The age of the studied assemblage collected from the lower part of the section, traditionally considered to be Upper Bajocian, corresponds to the Laeviuscula ammonite Chronozone and is the first reliable evidence of marine settings in the Volga Region for the early Bajocian. Two new biostratigraphic units, Beds with Eocylindroteuthis weisi and Beds with H. orphana, are introduced. The former, judging from the occurrences of its characteristic species in Western Europe, has a broad correlative potential. On the basis of datings obtained, the scheme of the formational subdivision of the Middle Jurassic of the Volga Region near Volgograd is revised, and the formerly abandoned Bakhtemir Formation is reinstated as a valid unit. Its total range is reestablished as corresponding to the upper part of the Discites (?)/Laeviuscula Chronozone (lower Bajocian)–Garantiana Chronozone (upper Bajocian).     

Representatives of genera <Emphasis Type="Italic">Malbosiceras</Emphasis> and <Emphasis Type="Italic">Pomeliceras</Emphasis> (Neocomitidae,Ammonoidea) from the Berriasian of the Crimean Mountains

The revised representatives of ammonite genera Malbosiceras and Pomeliceras from the Berriasian of the Crimean Mountains are classed with seven species, four of the first genus [M. malbosi (Pictet), M. chaperi (Pictet), M. broussei (Mazenot), M. pictetiforme Tavera] and three of the second one [P. aff. boisseti Nikolov, P. breveti (Pomel), P. (?) funduklense Lysenko et Arkadiev sp. nov.]. The identified species are described. The genus Mazenoticeras is considered as synonym of Malbosiceras. The above species prove that all the Berriasian zones (jacobi, occitanica and boissieri) are characteristic of corresponding deposits in the Crimean Mountains.     

The Bajocian–Bathonian plant fossil from the Hojedk formation,Lenjan section,Kerman, Iran

The Jurassic successions represent a wide distribution in North of the Kerman province. These successions include Ab-Haji, Badamu, and Hojedk formations. The Hojedk Formation contains the plant fossils. The Lenjan section is one of the suitable areas for paleontological studies on the Hojedk Formation. The study section is mostly composed of green sandstone and shale with several interbedded coal veins with different thicknesses. The thickness of the Hojedk Formation is about 200 m in the Lenjan section. In this study, seven genera and 13 species of macro plant fossils were identified and described, including Nilssonia undulata, Nilssonia bozorga, Nilssonia berriesi, Nilssonia sp., Klukia cf. exilis, Klukia exilis, Cladophlebis antarctica, Coniopteris lobata, Coniopteris murrayana, Elatocladus confertus, Podozamites sp., Equisetites sp., and Coniopteris sp. The Bajocian–Bathonian can be attributed to the Lenjan section based on the recognized flora.     

Permian nonmarine bivalve zonation of the East European platform

New finds and revision of available collections of nonmarine bivalves provided grounds for development of a zonal scale for terrestrial sequences of the Permian System based on species belonging to the genus Palaeomutela Amalitzky, 1891, which are characterized by regular changes in the structure of the shell hinge. The scale includes two parallel zonal successions that are based on the stratigraphic distribution and evolutionary trends of two morphological lineages of the genus. The zonal succession based on development of the P. umbonata group (dwellers of mobile waters and silty-psammitic substrates) includes 11 range zones: stegocephalum, ovatiformis, umbonata, quadriangularis, krotowi, wohrmani, numerosa, ulemensis, keyserlingi, curiosa, golubevi. The zonal succession based on development of the P. castor group (dwellers of calm waters and silty-pelitic substrates) includes eight range zones: larae, castor, olgae, doratioformis, marposadica, fischeri, obunca, amalitzkyi. The proposed zonal units are correlated with scales based on ostracod, fish, and tetrapod fossils. New species Palaeomutela golubevi sp. nov. and P. amalitzkyi sp. nov. are described with the extended diagnosis of the genus Palaeomutela.     

Gymnospermes s.l. du Jurassique supérieur du massif Manamana,Sud-Ouest de Madagascar: une nouvelle paléoflore gondwanienne

Fossil plants from Upper Jurassic sediments belonging to the Gymnosperms s.l. have been studied in the Manamana massif from the southwestern part of Madagascar. The levels are Upper Oxfordian in age on the basis of the palaeontological content, especially with ammonites. The majority of fossil plants are devoided of organic matter but in some cases collodion peels have been made to present epidermal characters. The cuticle of Brachyphyllum madagascariense nov. sp. has been observed with scanning electron microscope (SEM). This new flora is constituted by Pteridospermales and Coniferales. Fragmentary fronds of Pteridospermales have been collected of Pachypteris sp. A and Pachypteris sp. B. Some epidermal characters are described. Coniferales are represented by leafy stems. The diagnosis of Brachyphyllum madagascariense nov. sp. is given with morphological and epidermal characters. Others conifers have been determined: Brachyphyllum sp. A, Brachyphyllum sp. B, Elatocladus sp., Cupressinocladus sp., Cyparissidium sp.. Araucarites sp. remains are probably isolated fertile scales of a female cone. Complete cones of Conites sp. are also present.Comparisons of the systematic composition are made with other Gondwana floras of the same age from Antarctica, South America, Madagascar, India, South Africa, New Zealand and Australia. This flora shows xerophytic adaptations. With the Filicales described by Appert (1973a, b) from the same localities, it is now the most diverse flora from the Upper Jurassic of Madagascar, despite the lack of Cycadales, Bennettitales and Angiosperms.     

<Emphasis Type="Italic">Rhizopalmoxylon nypoides</Emphasis> – a new palm root from the Deccan Intertrappean beds of Sagar,Madhya Pradesh,India

A new species of fossil palm rhizome having root-mat under the organ genus Rhizopalamoxylon (Rhizopalmoxylon nypoides sp. nov.) is reported. The specimen shows the closest resemblance with the modern monotypic genus Nypa Wurmb of the Arecaceae. The specimen was collected from the late Maastrichtian–early Danian sediments of Deccan Intertrappean beds, Mothi, Sagar district, Madhya Pradesh, India. Nypa is a mangrove palm naturally found in estuaries and swamps of the tropical region and represents one of the oldest records of the genus from the Deccan Intertrappean beds of central India. The abundance of palms, including Nypa and previously recorded coastal and mangrove elements such as Acrostichum, Barringtonia, Cocos, Sonneratia and marine algae (Distichoplax and Peyssonellia) from the Deccan Intertrappean beds indicate marine influence and existence of tropical rainforest ecosystem in the vicinity of fossil locality in contrast to the deciduous forests occurring there at present.     

Early Triassic palynomorphs from Kuraloi Block,Belphar area,Ib-River coalfield,Mahanadi basin,Odisha

A rich assemblage of palynomorphs including miospores and megaspores has been recorded from the bore core no. IBKAN-2 drilled at a depth of 17.80m in Kuraloi Block-A, south-west part of the Ib-River Coalfield, Jharsuguda district, Odisha. The miofloral assemblage recovered shows prominence of cingulate, zonate, taeniate, non-striate and non saccate palynotaxa which indicate an early Triassic age and is comparable with the palynoassemblages of the same age known from the Damodar basin, India. The megaspore assemblage includes eight genera and fifteen species namely, Banksisporites karanpuraensis, B. indicus, B. utkalensis, Banksisporites sp., Barakarella pantii, Barakarella sp., Biharisporites spinosus, Biharisporites sp., Erlansonisporites triassicus, Erlansonisporites cf. erlansonii, Erlansonisporites sp. Jhariatriletes sp., Singhisporites sp., Ramispinatispora sp. and Talchirella trivedii. Records of megaspores are sparse in the early Triassic of Gondwana basins of India. The present study records the first occurrence of miospores and megaspores from the early Triassic of Mahanadi basin besides substantiating the earlier data.     

<Emphasis Type="Italic">Suraqalatia brasieri</Emphasis> n.gen., n.sp. (larger foraminifera) from the Maastrichtian of Sulaimani area in northern Iraq

Suraqalatia brasieri n.gen., n.sp. from the family Dicyclinidae Loeblich and Tappan 1964 occurs on the Maastrichtian carbonate platform of northern Iraq. The new genus is recognizable by its large very compressed conical test, up to 55–70 mm in diameter, to 0.3–1.6 mm in thickness, planspiral cooling having very small proloculus in the initial part and later circular chambers including numerous chamberlets with an agglutinated wall. Suraqalatia brasieri n.gen., n.sp. is associated with textulariids, miliolids and rotaliids as Loftusia elongata Cox, L. morgani Douvillé, Orbitoides medius d’Archiac, O. megaloformis Papp & Kupper, O. gruenbachensis Papp, O. apiculatus Schlumberger, Omphalocyclus macroporus (Lamarck), Siderolites calcitrapoides Lamarck, Sirtina orbitoidiformis Brönnimann & Wirz. The associated macrofauna comprises large and rich giant rudists (Preradiolites sp.), other bivalves (Gryphaea sp. and Glycymeris sp.), gastropods (Acteonella sp.), echinoderms and corals. The fauna indicates shallow marine carbonate platform conditions within the Maastrichtian green house. It is also worth mentioning that the new genus has only been recorded from the Maastrichtian age.     

Microbiostratigraphy of the Upper Paleocene to Middle Eocene Jahrum Formation in the Folded Zagros Zone,SW Iran

The Jahrum Formation (Upper Paleocene to Middle Eocene) is composed of carbonate and dolomitic carbonate rocks in the Zagros Basin. The Zagros is located at the boundary between the Arabian and Eurasian lithosphere plates and represent the orogenic response to a collision between Eurasia and advancing Arabia during the Cenozoic. The study area is located in the northern part of Kuh-E-Tudej, ~175 km southeast of Shiraz in the Folded Zagros Zone. The Jahrum Formation at Kuh-E-Tudej, with a thickness of 190 m, consists of medium to massive bedded limestone. The following foraminiferal index species are identified in the studied section: Fallotella alavensis, Kathina sp., Miscellanea sp., Lockhartia sp.,Orbitolites shirazeinsis, Nummulites sp., Opertorbitolites sp., Dictyoconus cf. egyptiensis, Orbitolites cf. complanatus, Dictyoconus sp., Coskinolina sp., Somalina stefaninii, Discocyclina sp., Praerhapydionina sp., Coskinolina cf. liburnica, Nummulites cf. globulus, Nummulites cf. aturicus, and Alveolina sp. The age of the studied sediments ranges from Upper Paleocene to Middle Eocene. The microbiostratigraphic studies revealed four biozones based on the foraminifers identified in the studied section.     

Some Calloporid (Bryozoa: Cheilostomata) species from the Cenozoic sediments of western Kachchh,Gujarat

Seven species of Calloporid (family Calloporidae Norman, 1903) Bryozoa are described from the Cenozoic sediments of the western Kachchh, Gujarat, India. Among these Planicellaria walsariensis, Dionella sp., Pyriporella charopadiensis, P. vadsariensis and Wilbertopora sp. are new and the other species viz; Crassimarginatella blandfordi and Reptoporina chhasraensis are reported for the second time from these sediments. The phylogenetic analysis suggests all the species of Calloporid bryozoans are monophyletic i.e. all of these species have common ancestor.     

A diverse rodent fauna from the middle Bartonian (Eocene) of Les Alleveys,Switzerland: snapshot of the early theridomyid radiation

A fossiliferous fissure filling at Les Alleveys, Mormont (Canton de Vaud, Switzerland), sampled in 1986 and representing part of the same fissure system as that excavated by Chavannes and Morlot in 1852, contains an Eocene mammal fauna of 27 species-group taxa, dominated by micromammals. Unlike the 1852 collection, thanks to screenwashing techniques, there is an abundance of rodents, especially theridomyids, which provide new information on the early radiation of the family. The homogeneity of the new assemblage suggests that the Les Alleveys fauna is entirely Bartonian in age and not as previously thought mixed with Priabonian elements. Bartonian faunas rich in a diversity of micromammals are relatively uncommon in Europe outside the Quercy region of France and this is the first such fauna from Mormont and from Switzerland. The rodents are here treated systematically and a new species of Elfomys, E. engesseri sp. nov. is described. A few teeth of an undescribed species tentatively referred to ‘Protadelomys’ provide an archaic element, whereas a single milk premolar of Patriotheridomys? suggests a considerable range for the genus prior to the middle Priabonian. The Les Alleveys theridomyids, together with a range of other early family members are analysed cladistically. The analysis confirms both the distinctness of the subfamily Remyinae and that the genus Estellomys is a primitive theridomyine; it shows Paradelomys to be a primitive member of the Columbomyinae, but finds no dental synapomorphies to support the Issiodoromyinae as represented by Elfomys and Pseudoltinomys.     

Record of Leaf and Pollen cf. <Emphasis Type="Italic">Sloanea</Emphasis> (Elaeocarpaceae) from the Middle Siwalik of Darjeeling sub-Himalaya,India and its Palaeobiogeographic Implications

Fossil leaf impressions and pollen grains comparable to modern Sloanea sp. of Elaeocarpaceae collected from the middle part of the Siwalik sediments (Geabdat Sandstone Formation; Pliocene) in Darjeeling foothills of eastern Himalaya are reported in the present communication. On the basis of macro morphological features, leaf remains are described as a new species Sloanea siwalika sp. nov. This is the first authentic record of the occurrence of leaf and pollen grains comparable to the genus Sloanea L. from the Cenozoic sediments of India and Asia as well. The recovery of this species and other earlier-described evergreen taxa from the same formation, suggests the existence of a tropical, warm and humid climatic conditions during the depositional period. The present study further suggests that after Pliocene the taxon might have shifted from Darjeeling Himalayan region to the adjoining southeast Asian land masses, due to possible climate change caused by post-Pliocene orogenic movement of the Himalaya.     

Stratigraphy of the Bathonian-Callovian boundary deposits in the Prosek section (Middle Volga Region). Article 1. Ammonites and infrazonal biostratigraphy

In European Russia, the most complete succession of Boreal sediments of the terminal Bathonian and lower Callovian is exposed near the Prosek Settlement. After its revision, the infrazonal division of the upper Bathonian and lower Callovian and position of the Bathonian-Callovian boundary are difined more carefully. The Calyx Zone and bodylevskyi Biohorizon are established in the upper Bathonian. The base of the lower Callovian is defined at the first occurrence level of Macrocephalites jacquoti. Based on four successive ammonite assemblages occurring in lower part of the Elatmae Zone, the breve, frearsi, quenstedti, and elatmae biohorizons are identified. The joint occurrence of Boreal, Subboreal, and Tethyan ammonites in the section facilitate its correlation with the other sections of the Panboreal paleobiogeographic superrealm.     

New data on microbiota of the Middle Volgian substage in the Loino Section (Kirov oblast)

The calcareous nannoplankton was established for the first time in Upper Volgian deposits of the Loino Section (Kirov oblast). Here, single coccolites of representatives of genus Watznaueria of wide stratigraphic and geographic distributions dominate. Representatives of genus Zuegrhabdotus are less common, whereas species of genus Polypodorhabdus are rare. The taxonomic composition of the calcareous nannoplankton assemblage does not allow us to distinguish the nannoplankton zone in the Loino Section, but it gives us an idea about its distribution at the Volgian Stage in the Russian Plate. As corresponding microfauna, foraminifers, assigned to the Lenticulina infravolgaensis–Saracenaria pravoslavlevi Zone, and ostracods were identified to substantiate the age of host deposits. The ostracod assemblage is assigned to the Macrodentina (Polydentina) subtriangularis Beds established on the Volga River left bank in deposits corresponding to the ammonite zones Panderi and Virgatus assigned to the middle Volgian substage. On the basis of ostracods, the warm-water well-aerating environment of the upper sublittoral (as deep as 50 m) was reconstructed.