New Azhdarchid Pterosaur from the Early Cretaceous of Western Liaoning

A nearly complete skeleton with a lower jaw of pterosaur from the Early Cretaceous of western Liaoning is described and assigned to a new genus, Eoazhdarcho gen. nov.. The new genus is characterized by a relatively small size,the ratio of the length to width of the middle series cervical vertebrae approximately 3.5 and the ratio of humeral length to femoral length approximately 0.96. The humerus of Eoazhdarcho shows great resemblances to that of previously described Azhdarchidae, so it is assigned to the family Azhdarchidae.     

A New Enantiornithine Bird from the Lower Cretaceous Jiufotang Formation in Jinzhou Area,Western Liaoning Province,China

A new Early Cretaceous enantiornithine bird from Liaoning Province of northeastern China, Shengjingornis yangi, gen. et sp. nov., is reported. This new bird possesses the following unique combination of features: a long rostrum, with some teeth in the front; short nasal; slender jugal; Y-shaped furcula, with expanded distal end of the hypocleidum; cake-like sternum, with a low and caudally distributed keel; strut-like and caudally concave coracoid. The derived features of the scapula and the wings suggest a powerful flapping flight capability.     

New Tapejarid Pterosaur from Western Liaoning, China

A new tapejarid pterosaur Huaxiapterus jii gen. et sp. nov. is erected based on a nearly complete skeleton with a skull. The skull morphology of Huaxiapterus jii gen. et sp. nov. appears transitional between the skulls of Sinopterus and Tapejara. The skull morphology （low crest） of Huaxiapterusjii indicates that Huaxiapterus is more closely related to Sinopterus than to Tapejara, which has a high crest. The relatively long and shallow skull of Sinopterus indicates that it is a primitive form. Huaxiapterus is more derived than Sinopterus but more primitive than Tapejara.     

辽西早白垩世一反鸟类新属种的发现

新反鸟化石（Gracilornis jiufotangensis gen. et sp. nov.）产于辽宁省朝阳地区早白垩世九佛堂组。根据该反鸟的骨骼细弱、肢骨关节发达、胸骨与躯干相比较小（胸骨宽与躯干长比率约为0.11;胸骨长与躯干长比率约为0.13）等不同于其他已知反鸟的特征,建立一新属种,归于华夏鸟科（Cathayornithidae）。     

New Material of Pterosaur Sinopterus （Reptilia： Pterosauria） from the Early Cretaceous Jiufotang Formation, Western Liaoning, China

1 Introduction The Jiufotang Formation is one of the most fossil- productive Early Cretaceous deposits of China. It overlies the famous Yixian Formation from which many important taxa were found (Ji et al., 2004; Chang et al., 2003; Zhou et al., 2003). In addition to other vertebrates, pterosaurs, the first flying reptiles in Earth history, are also abundant in the Jiufotang Formation. At present, nine species belonging to five genera have been reported from the Jiufotang Formation. The…     

New Eoenantiornithid Bird from the Early Cretaceous Jiufotang Formation of Western Liaoning, China

1 Introduction Enantiornithes were the most blooming branch among the early birds and had distinct diversities. They flourished in the Early Cretaceous (Kurochkim, 1985; Chiappe, 1991; Sanz and Bonaparte, 1992; Boles, 1995; Elzanowski, 1995), and become extinct at the end of the Late Cretaceous. Lots of the birds have been reported in western Liaoning, China (Sereno and Rao, 1992; Zhou, 1995; Hou, 1996, 1997; Sanz and Chiappe, 1996; Sanz et al., 1997; Hou and Chen, 1999; Hou et al., 1…     

An Early Cretaceous Avialian Bird, Shenzhouraptor sinensis from Western Liaoning, China

This paper describes an avialian bird from the Early Cretaceous Jiufotang Formation in Yixian county, western Liaoning, China, which is named as Shenzhouraptor sinensis by Ji et al. on July 15, 2002. Shenzhouraptor sinensis is characterized by no teeth in its mouth, the forelimbs longer than the hindlimbs, a long tail with more than 23 caudal vertebrae, U-shaped wishbone, and remiges longer than the total length of ulna and manus. It is certain that the new avialian bird is really capable of powerful flight, representing a missing link between theropod dinosaurs and birds.     

A New Specimen of Liaoceratops yanzigouensis (Dinosauria:Neoceratopsia) from the Early Cretaceous of Liaoning Province, China

A new specimen of basal neoceratopsian dinosaur Liaoceratops yanzigouensis is described. The specimen comes from the Lujiatun Bed of the Lower Cretaceous Yixian Formation in Beipiao City of western Liaoning Province, and is represented by a very well preserved three-dimensional partial skull and mandible. It is also the smallest among the three specimens of L. yanzigouensis, and several features in the new specimen, such as the short preorbital length and the round rostroventral orbital rim, can be ontogenetically-related. The superb exposure of the palatal complex may be caused by the removing of its brain by a small predator in the contemporary Jehol Biota.     

辽西义县组发现一新的翼手龙类化石

根据产自中国辽宁省西部早白垩世义县组上部、具有不完整头骨的翼手龙类骨架确立一新属新种：李氏始无齿翼龙Eopteranodon lii（gen．et sp．nov．）。从它窄长的头骨且其上、下颌尖长而没有牙齿，短的颈椎椎体上来看，显示了该翼龙与无齿翼龙类相似。始无齿翼龙的肱骨没有像其他无齿翼龙类那样具有弯曲的肱骨三角嵴，而且几乎所有的无齿翼龙类均个体较大。始无齿翼龙的个体较小，两翼展宽约为1．1m左右。腕骨没有愈合，肱骨三角嵴较平直以及第Ⅳ翼掌骨稍微长于尺骨显示了它的原始性，它代表一新的、较原始的、类似无齿翼龙的翼手龙类。     

辽宁西部早白垩世朝阳翼龙科 (Chaoyangopteridae)化石一新种

中国辽宁西部早白垩世翼龙类化石非常丰富,其上部的九佛堂组翼龙类组合以含有丰富的无齿的古神翼龙科、朝阳翼龙科等进步类型而与下部的义县组翼龙类组合相区别。本文描述了辽宁西部建昌盆地九佛堂组一新的大型翼龙类化石,其以上下颌无齿、吻端指数为3.7、前颌骨背支细长且封闭鼻眶前孔背缘、很大的鼻眶前孔后端超过上下颌关节位置、轭骨的上颌骨突基部宽大而应被归入朝阳翼龙科(Chaoyangopteridae)神州翼龙属(Shenzhoupterus)。其以较大的体形(翼展2.05 m)、平直的上下颌咬合面、轭骨的眶后骨突较泪骨突长且基部略宽、第4～7颈椎长度依次减小等特征,区别于朝阳神州翼龙(Shenzhoupterus chaoyangensis),而被命名为三亚神州翼龙(新种)(Shenzhoupterus sanyainus sp. nov.)。这是辽宁西部早白垩世翼展最大且不具牙齿的翼龙类属种,它的发现丰富了九佛堂组翼龙类组合内容,对认识朝阳翼龙科的骨骼形态与生态习性亦具有积极意义。     

A New Ornithocheirid from the Early Cretaceous of Liaoning Province, China

Based on a nearly complete skeleton with skull from the Early Cretaceous of Liaoning Province, a new ornithocheirid pterosaur: Boreopterus cuiae gen. et sp. nov. is erected. Boreopterus cuiae is different from other pterosaurs preserved with skulls known from the western Liaoning Province and its neighboring areas. Tiffs new oterosaur has more and larger teeth than those in other ornithocheirids. Its anterior nine pairs of teeth are larger than other teeth. The fourth pair of upper and lower teeth are slightly larger than the third pair. Overall, Boreopterus cuiae shows much small range of tooth size variation than Anhanguerapiscator and Coloborhynchus robustus. The new taxon shares with other ornithocherids in having a relatively large size of the third and fourth pairs of teeth.     

辽西早白垩世一反鸟类新发现

描述发现于中国辽宁朝阳早白垩世九佛堂组地层中的一件鸟类标本。该标本体型较小,具有许多反鸟类的近裔特征。通过对新标本后肢形态特征的分析,可判断其为一小型树栖反鸟类。新标本具有发育的前肢和进步的乌喙骨,及发育滞后的后肢和相对原始的胸骨形态。胸骨长宽比约为1,顶端呈圆弧形,两个后侧突末端不膨大,中央突末端稍扩大,龙骨突基本不发育等。新标本同时具有反鸟类的原始和进步特征,以及原始的胸骨特征。其综合特征不同于已知任意一种小型反鸟类,因此建立一个反鸟类新属种——普通小反鸟(Microenantiornis vulgaris gen.et sp.nov.)。     

A New Boreopterid Pterodactyloid Pterosaur from the Early Cretaceous Yixian Formation of Liaoning Province, Northeastern China

<正>A new boreopterid pterosaur:Zhenyuanopterus longirostris gen.et sp.nov.from the Yixian Formation of western Liaoning is erected,based on the complete skeleton with a skull and lower jaws preserved.It is characterized by:a large boreopterid pterosaur with a high number of teeth,where the anterior teeth are much larger than posterior ones;the length of the dorsal + sacral vertebrae is nearly half the length of the skull;ratio of the length of the humerus to metacarpal IV is approximately 91%and the,humerus,femur and third wing phalanx are all equal in length and the feet are specially small.It represents the largest boroepterid pterosaur discovered from western Liaoning and its surrounding areas so far     

A New Troodontid Dinosaur from the Lower Cretaceous Yixian Formation of Liaoning Province, China

A new troodontid dinosaur,Daliansaurus liaoningensis gen.et sp.nov.,is erected based on a nearly complete specimen from the Lower Cretaceous Yixian Formation of Beipiao City,Liaoning Province,China.This well preserved skeleton provides important new details of the anatomy for Liaoning troodontids,and gives new insight into their phylogenetic relationships and evolution.Daliansaurus is distinguished from other troodontids by an enlarged ungual on pedal digit IV,which is approximately the same size as the sickle-shaped second ungual,and is differentiated from other Liaoning troodontids by a number of characters of the skull,manus,pelvis,and hindlimb.A phylogenetic analysis recovers Daliansaurus within a subclade of Liaoning troodontids that also includes Sinovenator,Sinusonasus,and Mei.We erect a name for this group—Sinovenatorinae—and argue that it reflects a localized radiation of small-bodied troodontids in the Early Cretaceous of eastern Asia,similar to previously recognized radiations of Liaoning dromaeosaurids and avialans.As more Liaoning theropods are discovered,it is becoming apparent that small,feathered paravians were particularly diverse during the Early Cretaceous,and future work is needed to clarify how this diversity arose,which species coexisted,and how these numerous species partitioned niches.     

A new pterodactyloid pterosaur from the Early Cretaceous of the western part of Liaoning Province, northeastern China

Pangupterus liui gen. et sp. nov. from the Jiufotang Formation of Sihedang, Lingyuan City, Liaoning Province is erected based on a nearly complete lower jaw. It is characterized by having a total of 36 slender curved teeth with sharp tips, forming a distinct fish-grabbing mechanism; the teeth are well-spaced and are circular in section; the length ratio of the mandibular symphysis to the whole jaw is 20%; and the ratio of the tooth root width to tooth length is 12%. Toothed pterosaurs make up about 56.3% of the pterosaur assemblage from the Jiufotang Formation, which indicates that toothed forms played a key role in the ecosystem.     

Large-Sized Ankylosaur (Dinosauria) from the Lower Cretaceous Jiufotang Formation of Western Liaoning, China

Two incomplete bones referred to ankylosaur scapulocoracoid and humerus are described, which were discovered from the Early Cretaceous Jiufotang Formation of Dachengzi Town, Kazuo County, Liaoning Province. Ankylosaur dinosaurs have ever been found in the Lower Cretaceous Yixian, Fuxin and Sunjiawan Formations of western Liaoning. The occurrence of ankylosaurs within Jiufotang Formation improves our knowledge on the ankylosaur stratigraphic distribution. Based on the length of scapulocoracoid, the new dinosaur is estimated to exceed 6.0 meters long, representing the largest ankylosaur in western Liaoning.     

中国早白垩世热河生物群中大型霸王龙类化石的发现

一件包括部分头骨、下颌、肠骨等的不完整兽脚类恐龙骨架发现于辽宁喀左早白垩世九佛堂组中,估计身体全长可达9~10m。其前颌骨主体部分高、肠骨外侧面具显著的直立的嵴等特征表明该化石应归霸王龙类。它以外鼻孔大、上颌骨背缘前部略内凹、上颌骨孔向前达眶前窝前缘、腹缘与眶前窝腹缘存在较大距离等特征不同于晚白垩世的霸王龙科分子,故被命名为一新属种——喀左中国暴龙(Sinotyrannus kazuoensis gen. et sp. nov.),或许代表了最早的霸王龙科类型。该属种是已知个体最大的前晚白垩世霸王龙类,也是辽西及周边地区热河生物群中个体最大的兽脚类恐龙。它的发现不仅表明东亚是霸王龙类最主要的演化地区之一,而且为探讨霸王龙科的起源、热河生物群的组成与生态系统等提供了重要依据。     

辽宁西部一新的帆翼龙类化石

根据发现于辽西朝阳九佛堂组一具有部分头骨的不完整骨架,命名一新的帆翼龙——赵氏龙城翼龙（新属新种）。头骨和牙齿形态显示龙城翼龙与宽齿帆翼龙的关系要比与其他翼龙的关系密切,龙城翼龙是目前辽西九佛堂组中发现的唯一的帆翼龙类。帆翼龙科原仅一属种,发现于英国早白垩世地层中,辽西早白垩世新帆翼龙的发现,不但扩大了其地理分布,也证明辽西是晚侏罗世至早白垩世翼龙辐射、分异的重要地区。这对研究翼龙的起源和演化有重要意义。     

中国辽西早白垩世义县组始无齿翼龙新材料

根据新材料,对始无齿翼龙的特征进行了修订：中部颈椎体没有神经棘;肱骨三角嵴的长度与肱骨长度之比率约为0．2．5：翼掌骨的长度与第二翼指骨的长度几乎相等;第二翼指骨与第一翼指骨的长度之比率约为0．76。始无齿翼龙的新标本与正型稍微不同,可能由个体发育所引起。     

中国辽宁义县早白垩世燕鸟一新种

辽宁锦州义县早白垩世义县组发现了一种新型燕鸟(Yanornis)化石.该新型化石具有燕鸟的典型特征,如齿骨直且约占头骨全长的2/3、每侧约含20枚牙齿,颈椎细长,异凹型.愈合荐椎包括9枚脊椎,前肢约为后肢长的1.1倍.近端脚趾较其它趾节长和粗壮.然而,该新型化石部分特征又不同于燕鸟的模式种——马氏燕鸟(Yanornis martini),如其肱骨三角肌脊强烈发育,约为肱骨长度的一半.尺骨、桡骨及手部近等长,均长于肱骨.耻骨联合部较短,与尾综骨等长,不到耻骨长度的1/6.腓骨较长,约为股骨长度的4/5,约为胫跗骨长度的2/3.跗跖骨较短并完全愈合,不及胫跗骨长度一半.根据该新型化石的独特性及其与义县鸟、松岭鸟和马氏燕鸟的特征对比,笔者建立了燕鸟一新种——国章燕鸟(Yanornis guozhangisp.nov.).此外,国章燕鸟(新种)颈部还保存有3条鱼化石,这一特殊现象为研究燕鸟的生活习性和发育模式提供了重要信息.