A baby mosasauroid (Reptilia,Squamata) from the Turonian of Morocco – Tethysaurus ‘junior’ discovered?

New Middle Turonian mosasauroid remains were discovered in the same large-sized nodule that yielded a specimen referred to Tethysaurus nopcsai Bardet et al., 2003, from the Goulmima region (southern Morocco). They comprise isolated, fragmentary cranial elements (skull and mandible) and some vertebrae. Their very small size suggests a juvenile condition, an observation supported also by anatomical (spongious nature of numerous bone parts), micro-anatomical (loose inner spongiosa) and histological (numerous remains of calcified cartilage deep into the centrum; radial vascular canals) data. These bones belonged to a basal mosasauroid that cannot be distinguished from Tethysaurus nopcsai to which taxon we tentatively assign the material.     

Redescription of the elasmosaurid plesiosaurian Libonectes atlasense from the Upper Cretaceous of Morocco

The holotype of Libonectes atlasense is an almost complete skeleton from Upper Cretaceous (mid-Turonian) rocks of the Goulmima area in eastern Morocco. Initial assessment of this specimen in 2005 proposed generic referral based on stratigraphical contemporaneity with Libonectes morgani from the Cenomanian–Turonian of Texas, U.S.A. Nevertheless, relative differences in the profile of the premaxillary-maxillary tooth row, position of the external bony nasal opening, number of teeth and rostrad inclination of the mandibular symphysis, proportions of the axial neural arch, and number of cervical and pectoral vertebrae were used to distinguish between these species. As part of an on-going comparative appraisal of elasmosaurid plesiosaurian osteo-anatomy, we re-examined the type and formally referred material of both L. atlasense and L. morgani in order to establish species validity, as well as compile a comparative atlas for use in future works. Our inspections revealed that these reportedly distinct species-level fossils are in fact virtually indistinguishable in gross morphology. Indeed, the only substantial difference occurs in relative prominence of the midline keel along the mandibular symphysis, which might be explained by intraspecific variation. Furthermore, our observations permit an amendment to the published generic diagnosis of Libonectes with the confirmation of important states such as the likely presence of a pectoral bar, distocaudad expansion of the humerus, and an epipodial foramen. In addition, novel features include a prominent ‘prong-like’ ventral midline process on the coracoids, and the development of a median pelvic bar that encloses a central fenestration. The composite remains of L. morgani thus constitute one of the most complete elasmosaurid skeletal hypodigms documented worldwide, and evidence a trans-Atlantic distribution for this apparently dispersive species during the early–Late Cretaceous.     

Note on Upper Cretaceous ostracods from south-western Morocco

Cenomanian, Turonian and Coniacian ostracods from a borehole in the coastal zone of the Tarfayan Basin, south-western Morocco, display affinities with associations from Algeria, Tunisia and the Middle East as well as with the Iberian Peninsula. Relationships with central and southern West Africa are relatively slight, which could be a result of palaeoecological particularities.     

Zarafasaura oceanis, a new elasmosaurid (Reptilia: Sauropterygia) from the Maastrichtian Phosphates of Morocco and the palaeobiogeography of latest Cretaceous plesiosaurs

Though the Maastrichtian Phosphates of Morocco have yielded very rich marine vertebrate assemblages, plesiosaurs remain very scarce in these strata. The only hitherto recognized taxon was Plesiosaurus mauritanicus Arambourg, 1952, regarded here as a nomem dubium. Here we describe a new genus and species of elasmosaurid plesiosaur, Zarafasaura oceanis, which represents the first valid elasmosaurid plesiosaur described from the latest Cretaceous of Africa, and the second one from this continent. A phylogenetic analysis of plesiosauroids indicates that Zarafasaura oceanis has close affinities with elasmosaurids from the Late Cretaceous of North America and Japan. Among its distinctive suite of characters, the general shape and organisation of its squamosal and palate are unique among elasmosaurids. This new taxon completes our understanding of Late Cretaceous plesiosaur palaeobiodiversity and palaeobiogeography, and shows that Maastrichtian plesiosaurs were characterized by a quite high degree of endemism. They were also highly diversified and distributed worldwide, which supports the hypothesis of a catastrophic extinction of plesiosaurs at the K/T boundary.     

Vulnerability mapping of the Turonian limestone aquifer in the Phosphates Plateau (Morocco)

Turonian limestones are an important source of water indispensable for industrial, urban and agricultural development in the Phosphates Plateau. This is an elaboration of a vulnerability map of the aquifer, based on a multicriterion approach according to the DRASTIC charter that serves as a tool for assessment, management and protection of the groundwater quality.     

A new Plattenkalk Konservat Lagerstätte in the Upper Cretaceous of Gara Sbaa, south-eastern Morocco

Laminated, dolomitised marine limestones within a transgressive fluvio-lagoonal-carbonate platform sequence at Gara Sbaa in the Kem Kem region of south eastern Morocco yield well preserved marine fishes and crustaceans. A rarer terrestrial component includes delicate fern fronds, insects and a lizard. Sedimentological structures and stratigraphic context indicate initial shallow waters with in a carbonate lagoon with benthic microbial mats followed by deepening. High precision dating has yet to be accomplished, but a Late Cenomanian or Early Turonian age is indicated on the basis of faunal and sequence-stratigraphic considerations. The fish assemblage has affinities with mid-Cretaceous ichthyofaunas in South America and Lebanon, including taxa in common at generic level.     

A new protacanthopterygian fish from the Upper Cretaceous (Turonian) of the Pelotas Basin,southern Brazil

The Turonian age in the Pelotas Basin is represented by the Atlântida Formation, which consists mainly of marine dark shales and siltstones with intervening layers of carbonates. It has yielded a neritic fish assemblage that includes lamniform, pycnodontid, cladocyclid, clupeomorph, basal euteleostean, dercetid, enchodontid and holocentrid remains. In this assemblage there is a small fish, here described as Parawenzichthys minor gen. et sp. nov. It is characterized by having a low and elongate body, prominent snout, drop-shaped nasal, smooth dermal bones, toothless jaws, a mandible-suspensorium joint placed at the posterior portion of the orbit, very small pleural ribs, and at least 50 smooth vertebrae. Although superficially very similar to Santanasalmo elegans, known from the Albian of the Araripe Basin, it is separated by a combination of features of skull and, particularly, in the caudal endoskeleton. The first uroneural shows reduced anterior outgrowth and is fused to the compound centrum (first preural + ural centra), and the second uroneural is very long. There are six hypurals, the first of which is the largest. Taking into account the presence of derived conditions in the skull and caudal endoskeleton, the fish is assigned to Protacanthopterygii, sharing certain derived features with the argentinoid Wenzichthys congolensis from the non-marine Lower Cretaceous of Gabon (Africa).     

Ornithomimidae (Dinosauria: Theropoda) from the Bissekty Formation (Upper Cretaceous: Turonian) of Uzbekistan

The stratigraphically oldest remains of ornithomimid theropod dinosaurs are known from the Cenomanian Khodzhakul Formation and the Turonian Bissekty Formation of Uzbekistan. The ornithomimid from the Bissekty Formation is documented by more than 800 isolated bones that represent much of the skeleton. It shows at least three unambiguous synapomorphies of Ornithomimidae: length of anterior cervical centra three to five times greater than transverse width; low and rounded fibular crest of tibia; metatarsal III pinched between metatarsals II and IV; and proximal end of metatarsal III not visible in anterior view. Phylogenetic analysis, based on a dataset with 568 morphological characters and including all known ornithomimosaurian taxa, places the ornithomimid from the Bissekty Formation near the base of the ornithomimid radiation, between Archaeornithomimus asiaticus and Sinornithomimus dongi.     

Early Cretaceous (Valanginian – Hauterivian) belemnites from western Morocco: stratigraphy and palaeoecology

Recent field work in Lower Cretaceous successions of the Agadir-Essaouira region, western High Atlas, Morocco, have resulted in 489 belemnite guards collected bed-by-bed from the lower Valanginian – upper Hauterivian interval. These findings are described and discussed in their biostratigraphic, taxonomic, palaeoecologic and palaeobiogeographic context. The most common belemnite genera, Hibolithes, Vaunagites?, and Duvalia, are of Meditteranean affinities and closely related to Spanish, French, and Bulgarian faunas. Less common taxa include Pseudoduvalia and Berriasibelus. The genus Pseudobelus, however, common in southeastern France and southern Spain, is rare in the Moroccan sections. The palaeoecological interpretation of these observations result in the recognition of three belemnite assemblages, characterizing different marine settings: 1) A Hibolithes - Duvalia assemblage which is typical for inner to middle shelf settings. These are relatively shallow marine environments with a high input of siliciclastics. 2) A Duvalia - Hibolithes assemblage of the outer shelf setting. Duvalia spp. is the most common genus, Hibolithes spp. is less common, while Berriasibelus spp. and Pseudobelus spp. are rare. 3) A Duvalia - Pseudobelus assemblage of hemi-pelagic preference. Duvalia is most common, next is Pseudobelus.     

Processes of water–rock interaction in the Turonian aquifer of Oum Er-Rabia Basin, Morocco

Possible water–rock interaction processes, in the Moroccan basin of Oum Er-Rabia, were discussed by a geochemical study of groundwater from the Turonian limestone aquifer, the most important water resource in the region. Different types of water according to the classification of Piper were defined. Waters have shown an evolution from dominant CHO₃–Ca–Mg type through mixed to SO₄–Cl–Ca–Mg type. The use of geochemical diagrams and chemical speciation modeling method has shown that water–rock interaction is mainly controlled by carbonate and anhydrite dissolution, ion exchange and reverse ion exchange processes. Water–rock equilibrium conditions are favorable for the precipitation of calcite, dolomite, kaolinite and magnesian smectite.     

Mapping of groundwater quality in the Turonian aquifer of Oum Er-Rabia Basin, Morocco: a case study

This study takes the groundwater of the Moroccan limestone aquifer of Oum Er-Rabia as an example of statistical and cartographical approaches in water resources management. Statistical analyses based on frequency distribution and PCA methods revealed the homogeneity of waters with the existence of abnormal points and have helped to assess correlations between the studied variables. The mapping approach illustrated that waters are influenced by the lithology of the surrounding rocks and are of Ca–Mg–HCO₃, Ca–Mg–Cl–SO₄, and mixed types according to the Piper classification. The quality of water is of high to medium, north of the basin, but it is of medium to bad, NE and south, due to excessive contents of chloride, sulfate and nitrate. According to the US Salinity Laboratory classification, water used for irrigation in the eastern and the southern parts of the basin should take into consideration the drainage conditions, the nature of plants and the addition of gypsum doses.

Skeletal remains of Tyrannosauroidea (Dinosauria: Theropoda) from the Bissekty Formation (Upper Cretaceous: Turonian) of Uzbekistan

There exists a major gap in the fossil record of tyrannosauroid theropod dinosaurs spanning the early part of the Late Cretaceous. We report on skeletal remains referable to Tyrannosauroidea indet. from the Turonian Bissekty Formation at Dzharakuduk in the central Kyzylkum Desert of Uzbekistan. Phylogenetic analysis of the hypodigm places the Bissekty tyrannosauroid as a non-tyrannosaurid tyrannosauroid more basal than the Campanian tyrannosauroids Appalachiosaurus and Bistahieversor from North America.     

Hesperornithiformes (Aves: Ornithurae) from the Upper Cretaceous Pierre Shale,Southern Manitoba,Canada

A historical collection of hesperornithiform fossils from the Gammon Ferruginous, Pembina, and Millwood members of the Pierre Shale (Campanian, Upper Cretaceous) in southern Manitoba, Canada, was examined to revise their taxonomy. Only two species of Hesperornis have been recognized in previous studies on the Pierre Shale in Manitoba, but our study recognizes six species of two genera, including H. lumgairi sp. nov. H. regalis is the most common species but absent in the uppermost unit within the studied sequence. The result of this study supports the paleobiogeographic subdivision of the Campanian vertebrate fauna within the Western Interior Seaway, but not the faunal boundary that distinguishes the avian fauna of Manitoba from that of South Dakota and Kansas.     

A new mawsoniid coelacanth (Actinistia) from the Upper Cretaceous of Southern France

Axelrodichthys megadromos sp. nov. is a coelacanth described based on a single specimen collected in the lower Campanian site of Ventabren motorway, Southern France. The new species is referred to the mawsoniids because of the ornamentation of the skull roof and of the proportionally wide supraorbital series. The specimen belongs to the Mawsonia-Axelrodichthys complex based on features present on the lower jaw and on the basisphenoid. The new species is referred to the genus Axelrodichthys because of proportions of its parietonasal shield and because of the arrangement of the posterior parietals relative to the supraorbitals. Autapomorphic characters, in particular on the parasphenoid, justify the inclusion of the specimen in a new species. The occurrence of a mawsoniid in the Ibero-Amorican Island that formed part of the European Archipelago in the terminal Cretaceous is an evidence of a dispersal event from the southern land masses. The occurrence of A. megadromos in the Campanian-Maastrichtian represents the last occurrence of mawsoniids worldwide, after a gap in the fossil record of about 30 million years. This belated occurrence of Axelrodichthys extends the time range of this genus to approximately 40 myr and suggests that this genus, together with its sister genus Mawsonia, were organisms with a slow morphologic evolution.     

Transatlantic latest Cretaceous mosasaurs (Reptilia,Lacertilia) from the Maastrichtian type area and New Jersey

Cranial and postcranial elements of the mosasaurid reptiles Mosasaurus hoffmanni Mantell 1829 and Plioplatecarpus marshi Dollo 1882 from the Maastrichtian type area, and of Mosasaurus maximus Cope 1869 and Plioplatecarpus depressus (Cope 1869) from New Jersey are compared. Views held by previous authors are discussed. It is concluded that these European and North American taxa are conspecific.     

The Plio-Pleistocene evolution of the Southern Middle Atlas Fault Zone (SMAFZ) front of Morocco

The South Middle Atlas front constitutes a northeast-trending shear zone, located north of the Neogene Missour basin and east of the Taza Guercif basin. This paper analyses the Southern Middle Atlas Fault Zone (SMAFZ) deformation since the Pliocene. The set of structures observed suggests that reverse and thrust faulting along the central part of the SMAFZ are combined with left-lateral slip along N–S striking faults of its south-western termination and right-lateral faulting along E–NE striking faults of the east–northeast termination. Thrusts and oblique thrust-related anticlines of the two lateral ramps partly accommodate north-west directed motion of the African plate. The Thrusts probably resulted from rejuvenation of Jurassic normal faults; they were active during the Upper Miocene–Pliocene and the Pleistocene. The geometries of positive inversion structures and buttressing effects are clearly dependent on the geometry and sedimentology of the original basin-controlling fault system and on the presence of a décollement level. Field mapping is integrated with Landsat imagery and a digital elevation model to investigate the morphotectonic evolution of the south-eastern range front of the Middle Atlas. Geomorphological features provide significant information on the processes that govern lateral propagation of active anticlines. Both suggest that the deformation front may have been active since Pliocene.     

New species of Limoniidae (Diptera) from Myanmar amber,Upper Cretaceous

A new species of Dicranoptycha Osten Sacken, 1860, Dicranoptycha plicativa sp. nov., and a unassigned species are illustrated and described from Myanmar (Burmese) amber (lowermost Cenomanian, Upper Cretaceous). A complete list of fossil species of the genus Dicranoptycha is summarized. In addition, an updated key to the known fossil species of the Dicranoptycha is provided.     

Carbon isotope stratigraphy and depositional oxia through Cenomanian/Turonian boundary sequences (Upper Cretaceous) in New Zealand

Stratigraphic sections across the Cenomanian/Turonian boundary (C/T boundary) are identified in New Zealand and were deposited in southern high latitudes of the palaeo-Pacific. Lithological evidence for Cretaceous Oceanic Anoxic Event 2 (OAE2), which preceded and spanned the C/T boundary, is lacking in these sections. The correlative interval is identified, however, from a positive 2‰ carbon isotope excursion (CIE) and from clustered highest occurrences of Cenomanian-restricted dinoflagellate taxa together with the lowest occurrence of Turonian Heterosphaeridium difficile. A zone lacking benthic macrofossils encompasses the CIE. In some sections, this interval is also characterized by distinctive red mudstone beds; the thickest such red bed (6–18 m thick) may overlap or just overlie the main part of the CIE interval. Shelly macrobenthos, notably inoceramid bivalves, disappeared >500 kyr prior to the CIE. This suggests that environmental deterioration associated with OAE2 may have preceded the inferred volcanic trigger that has been identified from other regions. Strong intermediate water depth oxia during OAE2, which contrasts with oceanic anoxic conditions that occurred elsewhere on the globe, apparently prevailed during the later phase of OAE2 in the southernmost Pacific. New data from New Zealand indicate that causal mechanism(s) of OAE2 may be complex.