Kaikaifilu hervei gen. et sp. nov., a new large mosasaur (Squamata,Mosasauridae) from the upper Maastrichtian of Antarctica

We present a large, fragmentary skull and the humerus of a mosasaur (Squamata, Mosasauroidea) recovered from upper Maastrichtian beds of the López de Bertodano Formation in Marambio (=Seymour) Island, Antarctica. The material belongs to a large, adult individual with marked heterodonty as well as unusual humeral features. Different phylogenetic analyses returned the studied specimen within the Tylosaurinae, while the distinctive features of the skull and humerus allow distinguish it from the unique Antarctic known tylosaurine species, Taniwhasaurus antarcticus (Novas et al., 2002), as well as from other known Late Cretaceous mosasaurids from the Southern Hemisphere, thus, justifying the erection of a new taxon, Kaikaifilu hervei gen. et. sp. nov. The different dental types documented in the specimen studied have been previously recorded through isolated teeth from the same locality and were subsequently referred to several genera. This new find is relevant for assessing the previously known fragmentary records of Antarctic mosasaurids, suggesting that its local diversity could be more reduced than previously interpreted. The new material represents the youngest occurrence of tylosaurines in Antarctica.     

Shallow water agglutinated foraminiferal response to Late Cretaceous–Early Paleocene sea-level changes in the Dakhla Oasis,Western Desert,Egypt

The Late Cretaceous (Maastrichtian) to early Paleocene (Thanetian) shallow water (<100 m) agglutinated foraminifera from a section at Dakhla Oasis (Western Desert, Egypt) were analyzed for their assemblage, species and genera distribution, diversity, depositional environment, community structure and palaeobathymetry with respect to regional tectonics, climate and global eustasy. Data suggest an equitable benthic environment with low species dominance deposited in a brackish littoral and/or marsh setting. Sea level curves using characteristic benthic foraminiferal species, genera and assemblages corroborate quantitatively generated estimate and statistical analysis. Data suggests that in the absence of or of an impoverished benthic foraminiferal fauna, a high resolution agglutinated foraminiferal dataset can be as good a predictor of the benthic community structure and environment, as its calcareous counterpart, at least for shallow settings (<100 m). Present data also provides a good window in better understanding the distribution and interrelationship between the three dominant genera, Haplophragmoides, Trochammina and Ammobaculites. Faunal changes at boundaries (Cretaceous/Paleogene, Danian/Selandian and Selandian/Thanetian) are also evaluated.     

Late Cretaceous (Santonian) Atractosteus (Actinopterygii,Lepisosteidae) remains from Hungary (Iharkút,Bakony Mountains)

Lepisosteid fishes are well known from the Upper Cretaceous of Europe, but only by fragmentary remains from some Cenomanian and Campanian–Maastrichtian deposits. Here we report various cranial and postcranial remains of gars, discovered in the Upper Cretaceous (Santonian) Csehbánya Formation of Iharkút (Bakony Mountains, Hungary). These remains represent one of the most diverse assemblages of lepisosteid fish material from Upper Cretaceous continental deposits of Europe. Based on tooth morphology, scale-microstructure and the features of the supracleithrum we refer these remains to the genus Atractosteus. Besides some uncertain remains from the Cenomanian of France and Spain, the Santonian aged fossils from Iharkút represent the oldest undisputable occurrence of the family Lepisosteidae in the European continental Cretaceous. Using tooth crown morphology, the surface microstructure of the ganoid scales and the anatomy of the supracleithrum a review of the Late Cretaceous lepisosteid record suggests the occurrence of both Atractosteus and Lepisosteus in the European archipelago.     

A new species of Pluridens (Mosasauridae: Halisaurinae) from the upper Campanian of Southern Nigeria

The Upper Cretaceous of Africa has produced a diverse fauna of mosasaurs, including the highly specialized, long-jawed Pluridens. The type of Pluridens walkeri comes from the Maastrichtian Farin-Doutchi Formation of Niger, with a second, referred specimen coming from the Campanian section of the Campanian-Maastrichtian Nkporo Shale near Calabar, in southern Nigeria. Comparisons of this referred specimen with the holotype suggest that it represents a distinct and more primitive species. The Calabar jaw resembles P. walkeri in being long and narrow anteriorly with a shallow subdental shelf, and in having small, numerous, recurved teeth with medially positioned replacement pits. However, it lacks many of the derived features that characterize Pluridens walkeri, such as the extremely long and straight jaw, the extreme lateral protrusion and subcircular section of the dentary, strong transverse expansion of the dental thecae, and extreme reduction and increase in number of the teeth. The Calabar Pluridens is therefore referred to a new species, Pluridens calabaria. Following recent studies, Pluridens is considered to represent a highly derived member of the Halisaurinae. The marked differences between the Campanian and Maastrichtian species of the genus underscore the rapid pace of mosasaur evolution during the Cretaceous.     

Maastrichtian to Paleocene depositional environment of the Dakhla Formation, Western Desert, Egypt: sedimentology, mineralogy, and integrated micro- and macrofossil biostratigraphies

Integrated sedimentology, mineralogy, geochemistry, and microfossil and macrofossil biostratigraphies of the Maastrichtian–early Paleocene Dakhla Formation of the Western Desert, Egypt, provide improved age resolution, information on the cyclic nature of sediment deposition, and the reconstruction of depositional environments. Age control based on integrated biostratigraphies of planktic foraminifera, calcareous nannofossils and macrofossils yields the following ages for stratigraphic and lithologic sequences. The contact between the Duwi and Dakhla formations marks the Campanian/Maastrichtian boundary (zone CF8a/b boundary) and is dated at about 71 Ma. The age of the Dakhla Formation is estimated to span from 71 Ma at the base to about 63 Ma at the top (zones CF8a–Plc). The Cretaceous/Tertiary (K/T) boundary is within the upper unit of the Kharga Shale Member and marked by a hiatus that spans from 64.5 Ma in the lower Paleocene (base Plc) to at least 65.5 Ma (base CF2, base M. prinsii zones) in the upper Maastrichtian at Gebel Gifata, the type locality of the Dakhla Formation. As a result, the Bir Abu Minqar horizon, deposited between about 64.2 and 64.5 Ma (Plc(l) zone), directly overlies the K/T boundary hiatus. Major hiatuses also span the late Maastrichtian–early Paleocene in sections to the northwest (c. 61.2–65.5 Ma at North El Qasr, c. 61.2–69 Ma at Bir Abu Minqar and c. 61.2–65.5 Ma at Farafra), and reflect increased tectonic activity.During the Maastrichtian–early Paleocene a shallow sea covered the Western Desert of Egypt and the clastic sediment source was derived primarily from tectonic activity of the Gilf El Kebir spur to the southwest of Dakhla and the Bahariya arch. Uplift in the region resulted in major hiatuses in the late Maastrichtian–early Paleocene with increased erosion to the southwest. The area was located near the palaeoequator and experienced warm, wet, tropical to subtropical conditions characterized by low seasonality contrasts and predominantly chemical weathering (high kaolinite and smectite). A change towards perennially more humid conditions with enhanced runoff (increased kaolinite) occurred towards the end of the Maastrichtian and in the early Paleocene with shallow seas fringed by Nypa palm mangroves. Sediment deposition was predominantly cyclic, consisting of alternating sandstone/shale cycles with unfossiliferous shales deposited during sea-level highstands in inner neritic to lagoonal environments characterized by euryhaline, dysaerobic or low oxygen conditions. Fossiliferous calcareous sandstone layers were deposited in well-oxygenated shallow waters during sea-level lowstand periods.     

Stratigraphy,sedimentology and tectonic evolution of the Upper Cretaceous/Paleogene succession in north Eastern Desert,Egypt

The stratigraphy, sedimentology and syn-depositional tectonic events (SdTEs) of the Upper Cretaceous/Paleogene (K–P) succession at four localities in north Eastern Desert (NED) of Egypt have been studied. These localities are distributed from south-southwest to north-northeast at Gebel Millaha, at North Wadi Qena, at Wadi El Dakhal, and at Saint Paul Monastery. Lithostratigraphically, four rock units have been recorded: Sudr Formation (Campanian–Maastrichtian); Dakhla Formation (Danian–Selandian); Tarawan Formation (Selandian–Thanetian) and Esna Formation (Thanetian–Ypresian). These rock units are not completely represented all over the study area because some of them are absent at certain sites and others have variable thicknesses. Biostratigrapgically, 18 planktonic foraminiferal zones have been recorded. These are in stratigraphic order: Globotruncana ventricosa Zone (Campanian); Gansserina gansseri, Contusotruncana contusa, Recimguembelina fructicosa, Pseudohastigerina hariaensis, Pseudohastigerina palpebra and Plummerita hantkenenoides zones (Maastrichtian); Praemurica incostans, Praemurica uncinata, Morozovella angulata and Praemurica carinata/Igorina albeari zones (Danian); Igorina albeari, Globanomanlina pseudomenradii/Parasubbotina variospira, Acarinina subsphaerica, Acarinina soldadoensis/Globanomanlina pseudomenardii and Morozovella velascoensis zones (Selandian/Thantian); and Acarinina sibaiyaensis, Pseudohastigerina wilcoxensis/Morozovella velascoensis zones (earliest Ypresian). Sedimentologically, four sedimentary facies belts forming southwest gently-dipping slope to basin transect have been detected. They include tidal flats, outer shelf, slumped continental slope and open marine hemipelagic facies. This transect can be subdivided into a stable basin plain plus outer shelf in the extreme southwestern parts; and an unstable slope shelf platform in the northeastern parts. The unstable slope shelf platform is characterized by open marine hemipelagic, fine-grained limestones and fine siliciclastic shales (Sudr, Dakhla, Tarawan and Esna formations). The northeastern parts are marked by little contents of planktonic foraminifera and dolomitized, slumped carbonates, intercalated with basinal facies. Tectonically, four remarkable syn-depositional tectonic events (SdTEs) controlled the evolution of the studied succession. These events took place strongly within the Campanian–Ypresian time interval and were still active till Late Eocene. These events took place at: the Santonian/Campanian (S/C) boundary; the Campanian/Maastrichtian (C/M) boundary; the Cretaceous/Paleogene (K/P) boundary; and the Middle Paleocene–Early Eocene interval. These tectonic events are four pronounced phases in the tectonic history of the Syrian Arc System (SAS), the collision of the Afro-Arabian and Eurasian plates as well as the closure of the Tethys Sea.     

A mosasaur,cf. Plotosaurus,from the upper Maastrichtian Quiriquina Formation in Central Chile

Here we describe a mosasaurid from the upper Maastrichtian Quiriquina Formation near the town of Cocholgüe about 30 km north of Concepción, central Chile. The specimen comprises a concretion preserving the rostral part of a pair of mandibular rami. The bulbous base of the teeth allows referral to cf. Plotosaurus sp. This is southernmost record of this piscivorous and pelagic taxon. The southern dispersal of mosasaurs during the late Maastrichtian may not only have depended from warm south-bound Equatorial currents but mainly from the nutrient-rich oceanic regime along an upwelling zone of cold abyssal water that mixed with the warm surface water, resulting in an enormous primary production. Apparently mosasaur dispersal depended more on food than on suitable temperatures or currents.     

The first evidence of paleo-wildfire from the Campanian (Late Cretaceous) of North Africa

Although the fossil record of plant macro- and mesofossils, including fossil charcoal, is patchy geographically and temporally, such remains play an important role for the interpretation of paleoenvironmental and paleoclimatic developments in the continental realm. In Egypt, previous palynological studies on the Upper Cretaceous (Campanian) deposits suggested presence of lush subtropical forests, dominated by angiosperms and pteridophytes, which developed under warm and wet climatic conditions. In the present study, the occurrence of paleo-wildfires during the Late Cretaceous (Campanian) is presented for the first time, based on samples from a surface exposure in the vicinity of the Baris Oasis, south Western Desert, Egypt. Macroscopic charcoal was collected and subsequently analyzed under a stereomicroscope and scanning electron microscope (SEM). The charred wood remains were identified as belonging to gymnosperms, which were important components of the North African paleoflora during the Cretaceous. These charcoal remains represent the first verified occurrence of paleo-wildfires in Africa during the Campanian.     

Dinosaur eggshells from the Santonian Milk River Formation of Alberta,Canada

The North American fossil record of dinosaur eggshells for the Cretaceous is primarily restricted to formations of the middle (Albian–Cenomanian) and uppermost (Campanian–Maastrichtian) stages, with a large gap in the record for intermediate stages. Here we describe a dinosaur eggshell assemblage from a formation that represents an intermediate and poorly fossiliferous stage of the Upper Cretaceous, the Santonian Milk River Formation of southern Alberta, Canada. The Milk River eggshell assemblage contains five eggshell taxa: Continuoolithus, Porituberoolithus, Prismatoolithus, Spheroolithus, and Triprismatoolithus. These ootaxa are most similar to those reported from younger Campanian–Maastrichtian formations of the northern Western Interior than they are to ootaxa reported from older middle Cretaceous formations (i.e., predominantly Macroelongatoolithus). Characteristics of the Milk River ootaxa indicate that they are ascribable to at least one ornithopod and four small theropod species. The taxonomic affinity of the eggshell assemblage is consistent with the dinosaur fauna known based on isolated teeth and fragmentary skeletal remains from the formation, although most ornithischians and large theropods are not represented by eggshell. Relative to the Milk River Formation eggshell, similar oospecies occurring in younger Cretaceous deposits tend to be somewhat thicker, which may reflect an increase in body size of various dinosaur lineages during the Late Cretaceous.     

Late Cretaceous marine reptiles (Elasmosauridae and Mosasauridae) of the Chatham Islands, New Zealand

A new marine reptile record for the Upper Cretaceous of the Chatham Islands is described from the Takatika Grit. Incomplete mosasaur and plesiosaur remains represent the first record of marine reptiles from the Chatham Islands and wider New Zealand. Present among the myriad bones are elasmosaurid plesiosaurs and mosasaurine mosasaurs. This assemblage is comparable to the New Zealand marine reptile record and represents apex predators that flourished in a zone of upwelling in a Late Cretaceous southern high-latitude ecosystem.     

A stegosaur tooth (Dinosauria, Ornithischia) from the Early Cretaceous of southwestern France

The spotty nature of the terrestrial fossil record for the Mesozoic hinders a more complete understanding of dinosaur diversity. For stegosaurs (Ornithischia), the plated dinosaurs, only a few and fragmentary remains are reported from the Early Cretaceous of Europe. A recent revision concluded that only a partial vertebra of the nomen dubium Craterosaurus (?Aptian, England) could be considered as stegosaurian. Here we report on a stegosaur tooth from the Early Cretaceous (Berriasian) Purbeckian deposits of Cherves-de-Cognac (Charente), southwestern France. This tiny tooth was examined in detail using microtomography. Comparisons being limited by the rarity of stegosaur tooth rows material (e.g., from the skull of the holotype of Stegosaurus stenops) and dental material, notably from Europe, we observed new material of cf. Stegosaurus armatus and Hesperosaurus mjosi from the Upper Jurassic Morrison Formation of Wyoming (USA). The tooth shows the most similarities to the Late Jurassic genera Stegosaurus and Hesperosaurus, but differs in having a distinctive downwardly arched (V-shaped) cingulum on the ?lingual face (maxillary tooth hypothesis). It is referred to as Stegosauria indeterminate, a medium-sized quadrupedal herbivore that inhabited an emerged land between the Armorican Massif and the Massif Central. This finding is the first evidence of a stegosaur from the Early Cretaceous of France and a welcome addition to the meagre European record of that time. In addition, it is the second stegosaurian tooth crown reported from Europe. The assemblage of ornithischians of Cherves-de-Cognac shares some similarities with that of the Early Cretaceous (Berriasian) of the Purbeck Limestone Group, southern England. The relative rarity of ornithischian osteological remains in both Purbeckian environments suggests that most of these dinosaurs were mainly inhabitants of inland terrestrial palaeoenvironments.     

High‐resolution sequence stratigraphy of the Maastrichtian‐Ypresian succession along the eastern scarp face of Kharga Oasis,southern Western Desert,Egypt

A thick Maastrichtian‐Ypresian succession, dominated by marine siliciclastic and carbonate deposits of the regionally recognized Nile Valley and Garra El‐Arbain facies associations, is exposed along the eastern escarpment face of Kharga Oasis, located in the Western Desert of Egypt. The main objectives of the present study are: (i) to establish a detailed biostratigraphic framework; (ii) to interpret the depositional environments; and (iii) to propose a sequence stratigraphic framework in order to constrain the palaeogeographic evolution of the Kharga sub‐basin during the Maastrichtian‐Ypresian time interval. The biostratigraphic analysis suggests the occurrence of 10 planktonic zones; two in the Early Maastrichtian (CF8b and CF7), four in the Palaeocene (P2, P3, P4c and P5) and four in the Early Eocene (E1, E2, E3 and E4). Recorded zonal boundaries and biostratigraphic zones generally match with those proposed elsewhere in the region. The stratigraphic succession comprises seven third‐order depositional sequences which are bounded by unconformities and their correlative conformities which can be correlated within and outside Egypt. These depositional sequences are interpreted as the result of eustatic sea‐level changes coupled with local tectonic activities. Each sequence contains a lower retrogradational parasequence set bounded above by a marine‐flooding surface and an upper progradational parasequence set bounded above by a sequence boundary. Parasequences within parasequence sets are stacked in landward‐stepping and seaward‐stepping patterns indicative of transgressive and highstand systems tracts, respectively. Lowstand systems tracts were not developed in the studied sections, presumably due to the low‐relief ramp setting. The irregular palaeotopography of the Dakhla Basin, which was caused by north‐east to south‐west trending submerged palaeo‐highs and lows, together with the eustatic sea‐level fluctuations, controlled the development and location of the two facies associations in the Kharga Oasis, the Nile Valley (open marine) and Garra El‐Arbain (marginal marine).     

Dinosaur remains from the type Maastrichtian: an update

Isolated cranial and post-cranial remains of hadrosaurid dinosaurs have been collected from various outcrops in the type area of the Maastrichtian stage during the last few years. In the present contribution, dentary and maxillary teeth are recorded from the area for the first time. Post-cranial elements comprise a newly collected, fragmentary, large right metatarsal III and a broken ?right humerus, recently recognised in the collections of Teylers Museum (Haarlem). Unfortunately, none of these remains can be identified to species level. The available material suggests, however, that more than one taxon of non-lambeosaurine hadrosaurid and a possible euhadrosaurian are represented. Most of the new finds are stratigraphically well documented, which means that they may be linked to the recently published sequence-stratigraphic interpretation of the type Maastrichtian. Dinosaur remains recorded previously from the Maastrichtian type area are tabulated.

     

An abelisaurid from the latest Cretaceous (late Maastrichtian) of Morocco,North Africa

During the latest Cretaceous, distinct dinosaur faunas were found in Laurasia and Gondwana. Tyrannosaurids, hadrosaurids, and ceratopsians dominated in North America and Asia, while abelisaurids and titanosaurians dominated in South America, India, and Madagascar. Little is known about dinosaur faunas from the latest Cretaceous of Africa, however. Here, a new abelisaurid theropod, Chenanisaurus barbaricus, is described from the upper Maastrichtian phosphates of the Ouled Abdoun Basin in Morocco, North Africa on the basis of a partial dentary and isolated teeth. Chenanisaurus is both one of the largest abelisaurids, and one of the youngest known African dinosaurs. Along with previously reported titanosaurian remains, Chenanisaurus documents the persistence of a classic Gondwanan abelisaurid-titanosaurian fauna in mainland Africa until just prior to the end-Cretaceous mass extinction. The animal is unusual both in terms of its large size and the unusually short and robust jaw. Although it resembles South American carnotaurines in having a deep, bowed mandible, phylogenetic analysis suggests that Chenanisaurus may represent a lineage of abelisaurids that is distinct from those previously described from the latest Cretaceous of South America, Indo-Madagascar, and Europe, consistent with the hypothesis that the fragmentation of Gondwana led to the evolution of endemic dinosaur faunas during the Late Cretaceous.     

Springs in Egypt

Examples of springs in Egypt deal only with examples of natural springs producing potable water. None of the natural springs producing highly mineralized thermal water for therapeutical are considered. No water from natural springs in Egypt is bottled. Egyptian standards state that the total dissolved solids in potable water should not exceed 1000 ppm, except in Siwa, where the only available source for water for human consumption is from springs that have water containing more than 2000 ppm TDS. Six natural springs in Egypt provide typical examples for the Sinai and the Western Desert: Ain Furtaga in the southern pre-Cambrian province of Sinai Peninsula; Ain El Gudeirat in the sedimentary plateau of North Sinai; and Ain El Bishmo, Ain El Bousa, and Ain El Gabal in the Western Desert Oases of Bahariya, Kharga, and Dakhla. They discharge from the Nubian Sandstone aquifer system. The sixth spring, Ain El Arayes, is a spring in Siwa Oasis.     

Palynostratigraphy and paleobiogeography of the Jurassic – Lower Cretaceous succession in Kabrit-1 well,northeastern Egypt

Well-preserved and abundant Jurassic–Early Cretaceous palynomorph assemblages were recorded from the Kabrit-1 well, north Eastern Desert, Egypt. Thirty-one rock-cutting samples were analyzed and six rock units were differentiated. Seventy-one palynomorph species were identified from the productive samples. Six palynozones were differentiated, and they covered all the studied succession except for a palynomorph barren interval present in the uppermost part. These palynozones arranged in ascending order are as follows: two palynozones were recorded from the Middle-Upper Jurassic (Gonyaulacysta jurassica – Lithodinia jurassica Assemblage Zone and Klukisporites pseudoreticulatus- Systematophora penicillata – Escharisphaeridia pocockii Assemblage Zone) and the other four palynozones characterize the Lower Cretaceous deposits (Pilosisporites trichopapillosus – Cribroperidinium orthoceras Assemblage Zone; Dicheiropollis etruscus Interval Zone; Murospora florida – Afropollis operculatus Assemblage Zone and Afropollis jardinus Range Zone). We infer open marine conditions during deposition of the lower part of the Khalig El Ayoun Formation (Oxfordian-Kimmeridgian) and within the Kharita Formation (Albian), and shallow marine or coastal environments during deposition of the remainder of the studied succession. Paleobiogeographically, Late Jurassic index terrestrial palynomorph species documented here are in common with those previously recorded from North Africa, while marine dinocyst species are common to those documented from the West European and North American province. During the Early Cretaceous, the recorded terrestrial microfloral species were similar to those previously recorded from the West African-South American Province, while the marine dinocyst species were still related to the West European and North American marine palynofloral province.     

Planktic foraminiferal biostratigraphy of the Maastrichtian-Paleocene succession at the north Farafra Oasis,Western Desert,Egypt

Seven planktic foraminiferal zones are distinguished in the Maastrichtian-Paleocene succession at the north Farafra Oasis. These are the Rugoglobigerina hexacamerata (CF8b), Gansserina gansseri, and Contusotruncana contusa zones in the Maastrichtian topped by a well-known unconformity across the Cretaceous/Paleogene (K/Pg) boundary. The Danian is subdivided into two biozones: Globanomalina compressa/Praemurica inconstans-Praemurica uncinata Subzone (P1c) and Praemurica uncinata–Morozovella angulata (P2) Zone. The Late Paleocene is divided into two zones: Morozovella angulata-Globanomalina pseudomenardii (P3) and Globanomalina pseudomenardii (P4). A minor hiatus between the Danian/Selandian and Selandian/Thanetian boundaries are also recorded. These time gaps across the stage boundaries may be related to the tectonic events that affected the sedimentation regime throughout the Upper Cretaceous–Lower Paleogene interval in the Farafra Oasis.     

Palynostratigraphy and palaeoenvironmental significance of the Cretaceous palynomorphs in the Qattara Rim-1X well,North Western Desert,Egypt

Palynological and palynofacies analyses were carried out on some Cretaceous samples from the Qattara Rim-1X borehole, north Western Desert, Egypt. The recorded palynoflora enabled the recognition of two informal miospore biozones arranged from oldest to youngest as Elaterosporites klaszii-Afropollis jardinus Assemblage Zone (mid Albian) and Elaterocolpites castelainii–Afropollis kahramanensis Assemblage Zone (late Albian–mid Cenomanian). A poorly fossiliferous but however, datable interval (late Cenomanian–Turonian to ?Campanian–Maastrichtian) representing the uppermost part of the studied section was also recorded. The palynofacies and visual thermal maturation analyses indicate a mature terrestrially derived organic matter (kerogen III) dominates the sediments of the Kharita and Bahariya formations and thus these two formations comprise potential mature gas source rocks. The sediments of the Abu Roash Formation are mostly dominated by mature amorphous organic matter (kerogen II) and the formation is regarded as a potential mature oil source rock in the well. The palynomorphs and palynofacies analyses suggest deposition of the clastics of the Kharita and Bahariya formations (middle Albian and upper Albian–middle Cenomanian) in a marginal marine setting under dysoxic–anoxic conditions. By contrast, the mixed clastic-carbonate sediments of the Abu Roash Formation (upper Cenomanian–Turonian) and the carbonates of the Khoman Formation (?Campanian–Maastrichtian) were mainly deposited in an inner shallow marine setting under prevailing suboxic–anoxic conditions as a result of the late Cenomanian and the Campanian marine transgressions. This environmental change from marginal to open (inner shelf) basins reflects the vertical change in the type of the organic matter and its corresponding hydrocarbon-prone types. A regional warm and semi-arid climate but with a local humid condition developed near/at the site of the well is thought to have prevailed.     

An Upper Cretaceous (Campanian-Maastrichtian) actinopterygian fish assemblage from the marginal marine Adaffa Formation of Saudi Arabia

Actinopterygian remains have been recovered from Upper Cretaceous (lower Campanian to lower Maastrichtian) marginal marine deposits of the Adaffa Formation in northwestern Saudi Arabia. The fossils comprise gars (Lepisosteidae), pachycormids (cf. Protosphyraena sp.), indeterminate pycnodontiforms, enchodontid teleosts (cf. Enchodus sp.) and other indeterminate Teleostei. This assemblage is significant because it includes a novel occurrence for the Middle East (Pachycormidae) together with taxa (Lepisosteidae, Pycnodontiformes, Enchodontidae) that have been previously recorded from Late Cretaceous faunas elsewhere in the Mediterranean Tethyan region.     

First postcranial remains of Multituberculata (Allotheria,Mammalia) from Gondwana

Multituberculates (Allotheria) are generally regarded as the evolutionarily most successful and longest-lived (Middle Jurassic to late Eocene) clade of Mesozoic and early Paleogene mammals. Despite this “reputation” and the fact that the group is particularly well represented in both taxonomic diversity and relative abundance on Laurasian landmasses during the Cretaceous and Paleocene, multituberculates are exceedingly poorly represented on the southern supercontinent Gondwana. Previous records on Gondwanan landmasses have been based on fragmentary dental remains and all except the three most recently published (each represented by a single isolated tooth or fragment of tooth) have been disputed and allocated to either Haramiyida or Gondwanatheria. Furthermore, several previous records, disputed or not, are based on fragmentary dental remains of a type (plagiaulacoid) that has evolved independently several times in mammalian evolution.Here we place on record a multituberculate femur from the Upper Cretaceous (Maastrichtian) Maevarano Formation of the Mahajanga Basin, Madagascar. This specimen, although fragmentary as well, exhibits a number of features common to all multituberculate femora: neck cylindrical in cross section and set apart from shaft; greater trochanter prominent, extending proximally beyond head, inclined dorsally, and separated from neck by deep incisure; lesser trochanter prominent and protruding ventrally; posttrochanteric fossa present on ventral aspect, lateral to lesser trochanter; subtrochanteric tubercle present on dorsal aspect, distal to incisure between greater trochanter and neck; diaphysis straight, elliptical in cross section (slightly compressed dorsoventrally); and third trochanter absent. Three of these features (prominent, ventrally placed lesser trochanter; presence of posttrochanteric fossa; presence of subtrochanteric tubercle) are regarded as autapomorphies of Multituberculata. This specimen therefore not only independently and conclusively confirms the presence of the clade on Madagascar—previously based on a small molar fragment—but on the entire supercontinent as well.