New species of Saurichthys (Actinopterygii: Saurichthyidae) from Middle Triassic (Anisian) of Yunnan Province, China

Abstract: Saurichthys dawaziensis sp. nov., from the third member of the Gejiu Formation (Anisian of Middle Triassic) in the vicinity of Luoping, Yunnan Province, is a small to medium- sized saurichthyid fish. It is characterized by the efferent pseudobranchial arteries penetrating on the parasphenoid and the mandible sensory canal with a branch in the angular. The new species is more derived than the Lower Triassic species in having a relatively narrow postorbital region, short preopercular, single ossification of the sclerotic ring, small haemal arches between the pelvic fins and the anal fin without ossified spines, and in the absence of quadratojugal. It is more primitive than the Middle and Upper Triassic Saurichthys in having fringing fulcra and numerous segments in the unpaired fins, and the presence of both prae- and post-zygapophyses in the neural arches. Some anatomic changes related to the vertebral column and unpaired fins of the genus are also discussed.     

记云南中三叠世新的裂齿鱼类(英文)

记述了产自云南中三叠世的裂齿鱼类3个新属种——邓氏富源裂齿鱼(Fuyuanperleidus dengi gen.et sp.nov.)、苏氏罗平裂齿鱼(Luopingperleidus sui gen.et sp.nov.)和小齿滇东裂齿鱼(Diandongperleidus denticulatus gen.et sp.nov.)。邓氏富源裂齿鱼区别于其它裂齿鱼的特征,包括上颌骨形状、上下颌牙齿大小及形状、头骨纹饰、上颌骨与第一眶下骨愈合、高鳞片。苏氏罗平裂齿鱼区别于其它裂齿鱼的特征,包括三角形的鳃条骨、腹鳍前有4列水平方向的高鳞片、3个臀鳞。小齿滇东裂齿鱼区别于其它裂齿鱼的特征,包括胸鳍、腹鳍、背鳍和臀鳍前面鳍条的前缘具有小锯齿,鳞片后缘具有许多小锯齿。裂齿鱼类的新发现不仅丰富了中国华南裂齿鱼类的多样性,而且提供了裂齿鱼类在三叠纪全球辐射的新信息。     

First Record of Protorosaurid Reptile (Order Protorosauria) from the Middle Triassic of China

A new genus and species of the family Tanystropheidae, Dinocephalosaurus orientalis gen. et sp. nov., is described based on a nearly complete skull. This is the first record of the order Protorosauria from China. It also represents the only known occurrence of Tanystropheidae outside Europe, the Middle East and North America. Dinocephalosaurus is quite similar to Tanystropheus from Monte San Giorgio, Switzerland. Primarily it is distinguished from Tanystropheus in the shape of the premaxilla, maxilla, jugal and parietal. Although the family Tanystropheidae is now referred to the order Protorosauria, the new material from China indicates that the archosauromorph affinities of tanystropheids need further investigation. The discovery of Dinocephalosaurus provides new clues for the study of the evolution and radiation of Protorosauria and Tanystropheidae. It is also important for the study of the eastern Tethyan Fauna and the paleobiogeographical relationship between Europe and southern China in the Triassi     

The discovery of a new family Sunopteridae fam.nov.(Insecta: Protorthoptera) from Shaanxi Province,China

The fossil specimens discussed in this paper were collected from the grayish green mudstone and shale in the upper part of Lower Member of Middle Triassic Tongchuan Formation (T2t) in Shaanxi Province, China. Venationary feature correlation with old families and taxonomic discussion show that, among them, one specimen can be assigned to a new family - Sunopteridae fam. nov., which includes a new genus and species and can be referred to Order Protorthoptera Handlirsch, 1906 in taxonomic position, thus belonging to a new member of the Tongchuan Entomoassemblage of Shaanxi Entomofauna (belonging to Shaanxi Biota). The discovery of the new family and its new genus and species has certain reference values as follows: ① The discovery of the new family and its new genus and species fills firstly the vacancy in the protorthopterous field of China; ② In the meantime, it fills also the gaps of the valuable Middle Triassic protorthopterous fossil specimens between Late Carboniferous and Late Triassic Epochs in the world; ③ Judging from venationary features and taxonomic position, especially in the course of the formation of the merged vein of M and Cu, it formed a new unique and particular style of merged vein M+Cu. It not only is unique and simple, but also can be distinguished from the above mentioned 10 old families, thus providing important taxonomic evidence.     

A new species of the genus Besania Brough 1939 from the Middle Triassic of Canton Grisons (Switzerland) with a discussion of the phylogenetic status of the taxon

Recent excavations in the Middle Triassic Prosanto Formation (Ladinian, Ct. Grisons, Switzerland) have produced new material of the genus Besania Brough 1939. These specimens show many previously unrecorded details. Some features differ from those of the type species Besania micrognathus, and we therefore erect a new species, Besania schaufelbergeri sp. nov. This new material has prompted a re-evaluation of the phylogenetic interrelationships of the genus. The taxon seems to be closely related to basal Halecostomi. An erratum to this article is available at .     

云南罗平地区中三叠世龙鱼化石的发现

记述了龙鱼属一新种,即云南龙鱼Saurichth ysyunnanensis sp.nov.。该种以具有长椭圆形的鳃盖骨、躯干部披6列鳞片为特征与其他种区别。化石产于云南省罗平县大凹子村中三叠世安尼期关岭组中。     

A new basal actinopterygian fish from the Late Ladinian of Monte San Giorgio (Canton Ticino, Switzerland)

A new taxon of Late Ladinian basal actinopterygian is described on the basis of a single specimen from the Meride Limestone (Kalkschieferzone) of Meride (Canton Ticino, Switzerland). The species is characterized by a single plate-like branchiostegal, a suboperculum larger than the operculum, a maxillary fixed and posteriorly expanded, a large and rectangular dermopterotic and a large triangular dermosphenotic; the scales are rectangular to rhomboidal, the lepidotrichia of median fins are segmented from their base and the caudal fin shows epaxial rays. These features resemble the condition of the Redfieldiiformes, a freshwater fish group, whose presence in Europe is still controversial, owing to poor preservation of specimens found so far.     

The first record of redfieldiiform fish (Actinopterygii) from the Upper Triassic of Korea: Implications for paleobiology and paleobiogeography of Redfieldiiformes

A new genus and species of redfieldiiform fish, Hiascoactinus boryeongensis gen. et sp. nov., is described based on a nearly complete specimen from the Upper Triassic Amisan Formation of South Korea. Hiascoactinus is distinguished from other redfieldiiforms by having a barely ornamented dermal skull surface except for the snout region, two heteromorphic suborbitals arranged vertically behind the postorbital, a pistol-shaped suprascapular, and dorsal and anal fins with divided fin membranes between rays. The morphological features of the snout region and dorsal and anal fins of Hiascoactinus provide important clues to understand its feeding and swimming behavior. All previously proposed Asian redfieldiiform fossils are inaccurately classified, and consequently, Hiascoactinus is regarded as the only valid redfieldiiform taxon in Asia. Hiascoactinus is assigned to the basal group of redfieldiiforms primarily based on the presence of an antopercle. The basal phylogenetic position of Hiascoactinus indicates that basal redfieldiiforms dispersed from the southern Gondwanaland to the easternmost Laurasia through the terrestrial water system, and these two landmasses were connected during the Late Triassic.     

云南罗平中三叠统一海龙类新材料

海龙类前肢化石标本(YIGM SPC V 0832-2)采自于云南省罗平县新寨一带中三叠统安尼阶(Pelsonian亚阶)关岭组二段顶部.其不仅为海龙类在罗平生物群中的首次发现, 而且为长颈型海龙的最早记录.该海龙类新材料虽然保存不全, 但是仍具有一些独特的特征: 桡骨远端两侧边缘平行; 尺骨远端扩展; 11个硬骨化腕骨; 第三掌骨近端内侧缘发育横突; 指式为2-3-4-4-4(?).虽然上述特征尚不足以建立一海龙类新属种, 但是仍然反映了部分海龙类单元在向海洋适应辐射的过程保留有原始爬行类的特征.这些原始特征暗示着部分海龙类可能以营陆生生活为主.      

中华原始软骨硬鳞鱼之再描述及其地质时代

鲟形鱼类是一类脊椎骨尚未完全骨化、鳞片极度退化的原始辐鳍鱼类。本文重新系统描述了该类别中时代最古老的中华原始软骨硬鳞鱼(Eochondrosteus sinensis Lu,Li et Yang, 2005),补充并修订了其古生物学特征。该鱼类具有扇形的下鳃盖骨、多块条形鳃条骨、较大的喉板骨、具齿的上、下颌,且全身仅在尾鳍上叶具有鳞片等特征,可以区别于鲟形鱼类其他属种。初步的支序系统发育分析表明,Eochondrosteus可能是鲟形鱼类中最原始的类群,与所有其他鲟形鱼类的祖先类群构成一个姐妹群;同时其具有一些古鳕类才普遍拥有的特征,如相对较大的喉板骨和眶下骨等,有可能是原始辐鳍鱼类向鲟形鱼类进化过程中的过渡类型。这一鲟形鱼类化石产于甘肃省肃北县马鬃山镇西北约50 km的方山口组深灰色粉砂质页岩中。根据野外实地调查和岩石地层学、生物地层学对比,该鱼类的时代有可能是早三叠世,而非命名时认为的晚二叠世,但仍是迄今已发现的时代最古老之鲟形鱼类。     

A New Family of Moths from the Middle Jurassic (Insecta: Lepidoptera)

<正>Three lepidopteran species,from the Middle Jurassic Daohugou beds(inner Mongolia,China),are described in a new family,Mesokristenseniidae,and new genus,Mesokristensenia,which could represent the sister group of the Micropterigidae.Mesokristensenia differs from all extant Lepidoptera,but one genus(Agathiphaga, Agathiphagidae),in retaining four median veins in the forewing,a plesiomorphy also present in many Trichoptera. Evidence for placing Mesokristensenia in the Lepidoptera includes four traits,notably a previously unrecorded autapomorphy of this insect order:beyond stem M1+2,vein M1 is bent and connected to cross-vein r-m(in both wing pairs).Among 24 characters taken into account to assess the systematic position of Mesokristensenia,12 are considered informative for a cladistic analysis involving this fossil taxon and the four suborders recognized in present-day Lepidoptera(Zeugloptera,Aglossata,Heterobathmiina,and Glossata).     

Additional late Middle Cambrian trilobites from Karsha Formation (Haimanta Group) Zanskar region of Zanskar-Spiti basin, Northwest Himalaya

Additional latest Middle Cambrian polymerid trilobites are recorded from the Teta Member (Karsha Formation) along the Tangze-Yogma-Kuru section, Kurgiakh Valley, southeastern Zanskar, Northwest Himalaya. It includes record of a new genus Pianspis Saito & Sakakuru, 1936 of family Lisianiidae and others associated fauna comprises of Neoanomocarella asiatica (Lisianiidae), Parablackwelderia sp., (Damesellidae) and two undetermined pygidium and one unidentified cranidia. All these specimens are collected from the level representing the Lejopyge acantha Zone of Guzhangian Stage (Cambrian Series 3) of the Middle Cambrian. The fauna is mainly correlated with the South China, North Korea, Kazakhastan, Uzbekistan, Siberia and Australia.     

New Fossils of Eoptychopteridae (Diptera)from the Middle Jurassic of Northeastern China

Three new species of the extinct genus of Eoptychopterina from the Eoptychopteridae family, Eoptychopterina antica sp. nov., Eoptychopterina adnexa sp. nov., and Eoptychopterina mediata sp. nov., are described and illustrated. These three new species are established based on fossil specimens with bodies and complete wings. All were collected from the Middle Jurassic Jiulongshan Formation of Daohugou in eastern Inner Mongolia, China. Based on the new materials, the name of two species in Eoptychopterina from China--Eoptychopterina elenae Ren and Krzeminski and Eoptychopterina gigantea Zhang--is sysnonymum junius.     

New Isophlebioid Dragonflies from the Middle Jurassic of Inner Mongolia, China (Insecta: Odonata: Isophlebioptera:Campterophlebiidae)

Three new species of fossil dragonflies assigned to Sinokaratawia Nel, Huang and Lin in family Campterophlebiidae, i.e. S. daohugouica sp. nov., S. magica sp. nov. and S. gloriosa sp. nov., and new materials of male S. prokopi Nel, Huang and Lin, 2007 are described from the Middle Jurassic of Daohugou, Inner Mongolia, China. An emended diagnosis of genus Sinokaratawia was proposed.     

Strengthening Western Gondwanan correlations: A Brazilian Dicynodont (Synapsida,Anomodontia) in the Middle Triassic of Namibia

Terrestrial Middle Triassic strata occur throughout continental Africa and are particularly well exposed in South Africa, Tanzania, Zambia, and Namibia. The youngest age for all these African deposits is widely accepted as early Middle Triassic (Anisian). Fossils collected recently from the uppermost strata of the upper Omingonde Formation in Namibia highlighted the presence of Chiniquodon, a carnivorous cynodont previously only found in Ladinian-Carnian aged rocks of South America. In addition, work in progress indicates that a large archosaur, originally reported as Erythrosuchus, also discovered from levels close to the top of this unit, is in fact a rauisuchian, a group of archosaurs well known from Ladinian-Carnian beds of southern South America. Here we present the first record of the tuskless dicynodont Stahleckeria potens from the top of the upper Omingonde Formation in central Namibia. This taxon was up until now only known from the Ladinian Dinodontosaurus Assemblage Zone of the Santa Maria Formation in southern Brazil. Thus, compelling evidence for a Ladinian age for the upper levels of the upper Omingonde Formation is provided by two therapsid and one archosaur taxa. The tetrapod fauna of the upper Omingonde Formation partially fills the gap of the well-documented hiatus (Ladinian gap), prevalent throughout the Karoo basins of south and central Africa. The presence of the same therapsid taxa in the Namibian Waterberg Basin and the Paraná Basin of Brazil during Middle Triassic suggests that these basins were biogeographically linked through a series of interconnecting lowlands, with no major ecological, climatic and/or physical barriers.     

New Isophlebioid Dragonflies from the Middle Jurassic of Inner Mongolia,China(Insecta:Odonata:Isophlebioptera:Campterophiebiidae)

Three new species of fossil dragonflies assigned to Sinokaratawia Nel,Huang and Lin in family Campterophlebildae,i.e.S,daohugouica sp.nov.,S.magica sp.nov.and S.gloriosa sp.nov.,and new materials of male S.prokopi Nel,Huang and Lin,2007 are described from the Middle Jurassic of Daohugou,Inner Mongolia,China.An emended diagnosis of genus Sinokaratawia was proposed.     

New Fossils of Eoptychopteridae (Diptera) from the Middle Jurassic of Northeastern China

Three new species of the extinct genus of Eoptychopterina from the Eoptychopteridae     

New Species of Cimbrophlebiidae (Insecta: Mecoptera) from the Middle Jurassic of Northeastern China

Four new species of three genera in the family Cimbrophlebiidae, Telobittacus decorus sp. nov., Bellicimbrophlebia heteroneura sp. nov., Cimbrophlebia amoena sp. nov., and Cimbrophlebia gracilenta sp. nov., are described and illustrated based on eleven well-preserved fossil specimens. These specimens were collected from the latest Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia, China. This is the first record of the genus Cimbrophlebia in the Jurassic extending its age from the Lower Cretaceous to the Middle Jurassic. Due to same venational characters, we propose that an undescribed fossil specimen of Cimbrophlebiidae from the Jurassic of Germany is attributed to C. gracilenta sp. nov. Our new findings demonstrate an evolutionary trend of venational change from 185 to 50 Ma that the level of the bifurcation of Rs is gradually moving toward the base of wing in reference to the bifurcation of M.     

原长翅蝎蛉亚目化石在中国陕西铜川地区首次发现

记述了原长翅亚目化石（Protomecoptera Tillyard,1917）在中国陕西铜川中三叠统中的首次发现,填补这个亚目化石在中国的空白.根据脉序特征,确定是2个新属、种：Glytochorista martynovae gen.et sp.,Phyllochorista orientis gen.et sp.nov.,属这个亚目的汤姆蝎蛉科Tomichoristidae O.Martynova,1958,从而将这个科的化石推向最高层位,即从早二叠世推向中三叠世,跨时5000多万年.从这个意义来讲,对研究该科从古生代晚期向中生代早期的演化有一定的重要参考价值.化石采自铜川中三叠世铜川组下段上部灰绿色泥页岩.新属、新种属于陕西昆虫群（系中三叠世陕西动物群或陕西生物群的一个化石门类）铜川昆虫组合的新成员,时代相当于欧洲拉丁尼期（Ladinian stage）.     

Early Triassic (Late Griesbachian) gastropods from South China (Shanggan, Guangxi)

An Early Triassic (Griesbachian) gastropod fauna is reported from South China (Shanggan, Guangxi) and consists of four species: Bellerophon abrekensis, Wannerispira shangganensis Kaim & Nützel sp. nov., Naticopsis sp., and Palaeonarica guangxinensis Kaim & Nützel sp. nov. The taxon Wannerispira Kaim & Nützel nom. nov. replaces Pagodina Wanner non Van Beneden. This is the first report of Bellerophon abrekensis from China. Previously, it was only known from its type locality in Far East Russia. Wannerispira shangganensis sp. is the first certain Triassic report of the Permian subfamily Neilsoniinae and represents a holdover taxon. The neritimorph Palaeonarica is reported for the first time from the Early Triassic and this is the oldest occurrence of this genus. Compared with other Griesbachian gastropods, the present material is relatively well preserved so that the taxonomy rests on rather firm ground. Very few nominal taxa have been reported from the Griesbachian and therefore the present report presents substantial additional information about gastropods from the aftermath of the end-Permian mass extinction event. The gastropod association from Shanggan shares one species with Primorye, Far East Russia (B. abrekensis). Two species, W. shangganensis and P. guangxinensis, closely resemble specimens reported from the Griesbachian of Oman. This could suggest that Griesbachian gastropod faunas of the Tethys were rather homogenous although the data are still scarce.