湘中佘田桥剖面上泥盆统弗拉斯阶地层研究新进展

湘中佘田桥剖面的佘田桥阶大致与弗拉斯阶范围一致 ,其下为吉维阶泥质灰岩 ,含腕足类 Emanuella tak-wanensis等及竹节石 Viriatellina kienelensis等。佘田桥阶自下而上分别为榴江组、蒸水河组、老江冲组。榴江组主要是紫红色粉砂质泥岩 /粉砂岩互层、底部为黑色薄层硅质岩。蒸水河组下部以灰黑色及暗灰色钙质泥页岩为主 ,含介形虫 Bertillonella erecta、B.cicatricosa?、Franklinella calcarata、Entomozoe(N ehdentomis) tenera及菊石 Meso-beloceras?sp.;上部泥质灰岩、似瘤状灰岩、粉砂质泥岩含牙形石 Palmatolepis cf.semichatovae、大型菊石 Mantico-ceras sp.及介形虫 Entomozoe(N ehdentomis) pseudorichterina。老江冲组以泥页岩为主 ,顶部附近则出现一些泥质灰岩、似瘤状泥灰岩层 ,产丰富的小嘴贝类、无洞贝类及弓石燕类等腕足动物 ,少量海绵 (?)及珊瑚等。佘田桥阶的顶界与弗拉斯阶的顶界一致 ,即以无洞贝类和大量四射珊瑚的绝灭为标志 ,而 Yannanellina腕足动物群的出现开始于锡矿山阶 (或法门阶 )。综合各类化石资料 ,佘田桥剖面的榴江组大致相当于牙形石 falsiovalis带— punctata带 ,蒸水河组相当于 L ower hassi带— L ower rhenana带 ,老江冲组相当于 U pper rhenana带— linguiform     

贵州南部上泥盆统弗拉斯阶望城坡组的无洞贝类腕足动物——华南弗拉斯期的绝灭方式

贵州南部独山弗拉斯期的望城坡组含有两种无洞贝类腕足动物组合：下部为出现于贺家寨段底部的Atryparia（Costatrypa）dushanensis动物群，大致相当于牙形石falsiovalis至transitans带；上部为出现于卢家寨段的Radiatrypa yangi动物群，大致与hassi至上rhenana牙形石带相当。独山剖面无洞贝类及其他腕足动物资料表明，贺家寨段为弗拉斯期地层。     

弗拉斯—法门期之间的生物绝灭事件

晚泥分世弗拉斯—法门期(Frasnian—Famennian)之间存在一次全球性生物绝灭事件(简称F—F事件).本文简介了F—F事件的生物绝灭、事件界线、事件成因及近年国内研究情况弗拉斯末期,大量的海生无脊椎动物,尤其是热带、亚热带浅水区造礁及附来礁生物绝灭.绝灭的生物包括层孔虫、珊瑚、腕足类、竹节石、三叶虫、菊石及浮游植物等门类中的很大一部分成员.F－F件发生在牙形石的palmatolepis gigas带和Palmatolepis triangularis带之间,最可能发生在Palmatolepis gigas最上亚带.事件发生的原因有各种不同解释:1)海平面变化说;2)古气候变冷说;3)天文事件说等.目前国内对F—F事件的研究较少,侯鸿飞、王士涛(1985)曾总结了华南等地F—F界线附近的生物群变化.笔者等研究了西秦岭中带F—F事件,探讨了这次事件的可能成因.     

四川龙门山区中-上泥盆统底栖群落取代与海平面波动

四川龙门山区中—上泥盆统吉维特-弗拉斯阶观雾山组—土桥子组中可以识别出12个以腕足动物为主的底栖群落,群落变化可划分为两种群落取代类型：一种是突变取代,另一种是渐进取代。突变取代发生在弗拉斯阶土桥子组中Leiorhyrwhus群落和Zhonghuacoelia Striatopugnax群落之间。渐进取代发生在中泥盆统吉维特阶的观雾山组内,取代过程由Independatrypa lemma Uncinulus heterocostellis Ernanuella tokwanensis群落经过Sinospongophyllum irregulare Pseudomicroplasma fongi群落到Clathrocoilona spissa Hexagonaria复合礁群落完成。     

湖南中部早石炭世腕足动物组合及几个问题的讨论

对湘中下石炭统腕足动物群的系统研究表明,该地区的腕足动物群自下而上可划分为6个组合带;1.“plicatifera” tenuiplicaia Hou Ass.（Lower Toumaisian）,2.Unispiri-fer tournaicensis（Kon.）—Eochoristites alatus（Ching）Ass.（Middle Tournaisian）; 3．Finosp-irifer shaoyangensis（Ozaki）—Marginatia vaughani（Muir--Wood）Ass.（Upper Tournasian）;4.（?）itiliproductus groberi（Kren.）——Pugilis hunanensis（Ozaki）Ass.（Lower Visean）;5.Para-kansuella subrhomboidea Sp·nov.——Kansuella——Gigantoproductus maximus （M’coy）Ass．（UpperVisean）;6.Latiproductus latissimus（Sow.）—Gigantoproductus edelburgensis（Phillips）Ass.（Lower Namurian or Serpukhovian）.这些腕足动物大部分是特提斯区广泛分布的属种,能够与西欧,苏联等地的同时期动物群进行很好的对比.对邵东组腕足动物的研究表明,该组的地质时代属晚泥盆世末期.其所产腕足动物能很好地与西欧及苏联同期地层的腕足动物比较.研究表明,在湘中杜内与维宪阶的生物地层界线,根据腕足动物,应置于Finospirifer-marginatia动物群和Delepinea subcarinata—Megachonetes zimmermanni组合带之间,而不是过去认为的Eochoristives—Martiniella动     

桂北地区早石炭世杜内期的长身贝类的发现及其生物古地理意义

在广西北部兴安县高尚镇金子山剖面下石炭统鹿寨组底部，新发现了长身贝类腕足化石，包括1新种、1个相似种、6个未定种和2个属种未定类型。这些长身贝类以Productellidae和Productidae科的分子为主，另有少量Echinoconchidae和Monticuliferidae科的分子。这些长身贝类全部属于生活在静水环境中的薄壳类型，无一与华南同期地层中已知类型相同。它们之间的差异可能反映了不同沉积环境条件对腕足动物群属种的控制。在鹿寨组底部发现的长身贝类以及近年在桂林南边村和黔南睦化剖面腕足动物群的发现显示出华南台缘斜坡―盆地相区杜内阶中的腕足动物群具有极高的属种分异度，不但有大量见于世界其他地方同期动物群中的分子，也包含有不少原本认为只在维宪阶或更高层位出现的类型或全新的类型，说明华南台缘斜坡―盆地相区不但与世界其他地区的腕足动物群之间存在密切的生物地理区系联系，而且华南台缘斜坡―盆地相区是杜内期腕足动物起源和辐射演化的重要策源地之一。许多属种可能是在这里起源，并迁移扩散到其他地方。     

广西香田弗拉斯-法门阶界线事件标志——碳、氧稳定同位素异常特征

本文对广西香田晚泥盆世弗拉斯-法门阶界线剖面进行了系统的碳、氧稳定同位素研究。发现随着弗拉斯-法门期之交的生物大量死亡和绝灭,在弗拉斯-法门阶界线层中,碳、氧稳定同位素组成出现了明显的异常。这和白垩—第三系界线灾变事件中出现的异常颇为相似。碳同位素异常达-6.6‰左右,氧同位素出现正异常达2‰。此外,在界线层中出现铱异常为230×10^-12g/g。这些表明,上述同位素地球化学异常是地球历史中生物绝灭灾变事件的重要标志。     

西秦岭中带弗拉斯—法门期生物绝灭事件的研究

晚泥盆世弗拉斯—法门期(Frasnian—Famennian)之间存在一次全球性生物绝灭事件(简称F—F事件)。这次事件在西秦岭地区有所表现。西秦岭中带西成地区吉维特到弗拉斯期为碳酸盐台地和生物礁沉积。法门期以暗灰色和灰绿色板岩、粉砂质板岩沉积为主。弗拉斯期末(相当于gigas最上亚带),该区大量的层孔虫、珊瑚、腕足及兰绿藻等生物绝灭。沉积相、矿物及地球化学特征都表明该事件是山于海水急剧加深引起的。事件发生的时间为gigas上亚带和最上亚带之间。     

晚泥盆世弗拉斯期—法门期之交海水温度变化的同位素证据

位于我国湖南省冷水江地区锡矿山矿区的老江冲剖面为发育良好的浅水台地相沉积，出露著名的上泥盆统弗拉斯-法门阶（Fr/Fa)界线，多年的地层古生物学工作已详细地了解了该区的古生物面貌，本研究采用了基不可一世保存完好的腕足动物Cyrospirifer uernemili壳体进行了稳定C、O同位素地球化学分析，δ^18O值为-8.4‰-9.6‰，δ^13C值为-2.2‰-0.5‰，二值在弗拉斯-法门阶交界处均发生了异常波动，将此值与其他地区的研究结果进行比较，可以认为，弗拉斯-法门期交界时所发生的剧烈而大范围的事件所造成的同位素波动可以作为大区域地层对比的标志，经过计算，推断从弗拉斯期末至法门期初，我国华南地区的古海洋发生了降温事件。     

黔南宾夕法尼亚亚纪礁环境中腕足动物古生态研究

黔南地区宾夕法尼亚亚纪逍遥阶礁相地层中腕足动物普遍发育,主要产长身贝类和石燕贝类。本文主要研究该区腕足动物与造礁生物叶状藻和Fomitchevella 珊瑚之间的生态关系。在叶状藻礁内,小个体腕足动物常附着在藻叶上营假漂浮生活; 但藻叶密集处,小个体腕足动物也难见,可能藻丛造成水流循环不畅使腕足动物难以获得充足的食料; 藻叶稀疏处,腕足动物个体加大,数量增加。在大型Fomitchevella 珊瑚格架礁内,腕足动物与Fomitchevella 之间不存在食物上的竞争。腕足动物介壳层为Fomitchevella 幼虫定殖提供了生物成因的硬质基底,在此基础之上发育了大型珊瑚礁。腕足动物在 Fomitchevella 进入统殖阶段由于生态空间受到局限而退居次要地位。     

Atrypid Brachiopods from the Upper Devonian Wangchengpo Formation (Frasnian) of southern Guizhou, China — Extinction Patterns in the Frasnian of South China

The Wangchengpo Frasnian section of Dushan County contains two atrypid brachiopod assemblages. The lower is characterized by the Atryparia （Costatrypa） dushanensis fauna that appears at the base of the Hejiazhai Member approximately in the falsiovalis to transitans conodont zones： the upper is characterized by the Radiatrypa yangi fauna, which appears in the Lujiazhai Member approximately correspondent with the hassi to Upper rhenana conodont zones. Atrypid brachiopods, together with other brachiopods from the Dushan section show that the Hejiazhai Member is of Frasnian age. Preliminary analyses of Frasnian atrypid brachiopods from sections of South China indicate that there are nine genera and subgenera including Atryparia （Costatrypa）, Kyrtatrypa, Spinatrypa, lsospinatrypa, Spinatrypina, lowatrypa, Desquamatia （Desquamatia）. Desquamatia （Seratrypa）, and Radiatr）pa. Atrypid species diversity did not change much through the Frasnian. On a regional scale in South China, most atrypid species went extinct prior to the Frasnian/Famennian boundary. At any specific locality or section, these atrypids became extinct about 20-40 m below the Frasnian/Famennian （F/F） boundary, within the linguiformis conodont Zone, marking this as the major extinction level. Three new atrypid species are described： Atryparia （Costatrypa） dushanensis, lowatrypa pseudobodini, and Radiatrypa yangi.     

Microbial mounds prior to the Frasnian‐Famennian mass extinctions,Hantang, Guilin,South China

Late Frasnian mounds of the Yunghsien Formation, Guilin, South China, developed as part of the Guilin platform, mostly in reef‐flat and platform margin settings. Microbial mounds in platform margin settings at Hantang, about 10 km west of Guilin, contain Frasnian biota, such as Stachyodes and Kuangxiastraea and, thus, occur below the Frasnian‐Famennian mass extinction boundary. Platform margin facies were dominated by microbes, algae and receptaculitids. Massive corals and stromatoporoids are not common and rarely show reef‐building functions as they did in Givetian time. The margin mounds are composed of brachiopod‐receptaculitid cementstone, and a variety of boundstones that contain Rothpletzella, Renalcis, thrombolite and stromatolite. Other microbial communities include Girvanella, Izhella, Ortonella and Wetheredella. Solenoporoid algae are abundant locally. Zebra structures and neptunian dykes are well‐developed at some intervals. Pervasive early cementation played an important role in lithification of the microbial boundstones and rudstones. Frasnian reefs of many regions of the world were constructed by stromatoporoids and corals, although a shift to calcimicrobe‐dominated frameworks occurred before the Famennian. However, the exact ages of many Frasnian margin outcrops are poorly constrained owing to difficulties dating shallow carbonate facies. The Hantang mounds represent a microbe‐dominated reef‐building community with rare skeletal reef builders, consistent with major Late Devonian changes in reef composition, diversity and guild structure occurring before the end of the Frasnian. A similar transition occurred in the Canning Basin of Western Australia, but coeval successions in North America, Western Europe and the northern Urals are either less well‐known or represent different bathymetric settings. The transition in reef‐building style below the Frasnian‐Famennian boundary is documented here in the two best exposed successions on two continents, which may have been global. Set in the larger context of Late Devonian and Mississippian microbial reef‐building, the Hantang mounds help to demonstrate that controls on microbial reef communities differed from those on larger skeletal reef biota. Calcimicrobes replaced stromatoporoids as major reef builders before the Frasnian‐Famennian extinction event, and increasing stromatoporoid diversity towards the end of the Famennian did not result in a resurgence of skeletal reef frameworks. Calcimicrobes dominated margin facies through the Famennian, but declined near the Devonian‐Carboniferous boundary. Stromatolite and thrombolite facies, which occurred behind the mound margin at Hantang, rose to dominate Mississippian shallow‐water reef frameworks with only a minor resurgence of the important Frasnian calcimicrobe Renalcis in the Visean when well‐skeletonized organisms (corals) also became volumetrically significant frame builders again.     

Cephalopod limestone deposition on a shallow pelagic ridge: the Tafilalt Platform (upper Devonian, eastern Anti-Atlas, Morocco)

As a result of early Variscan tectonic movements and of differential subsidence, a platform and basin topography was created along the northern margin of the Sahara Craton during the late Devonian. In the Moroccan Anti-Atlas Mountains, the Tafilalt Platform is an approximately N-S running ridge which developed since the late Middle Devonian. It separated a slowly subsiding shallow basin in the east (Tafilalt Basin) from a rapidly subsiding furrow in the west (Mader Basin). Platform deposits are characterized by highly reduced thicknesses, shallow subtidal to supratidal deposits in the late Frasnian and by unconformities at the Lower/Upper Frasnian and the Frasnian/Famennian boundaries. After a local transgression over emergent areas in the north, water depth probably never reached more than several tens to about 100 m in the lower Famennian. Cephalopod limestones of this age, deposited on the platform, represent a very diverse facies pattern comprising quartz-rich brachiopod coquinas, crinoidal limestones, thick-bedded cephalopod limestones and nodular limestones. Sedimentation rates ranged from 1 to 5 mm/ 1000 yr. In the late Famennian more uniform marl and nodular limestone facies suggest slightly deeper environments. Platform margins are characterized by higher rates of subsidence, debris flow deposits and slump structures. In the relatively shallow Tafilalt Basin, marls with intercalated nodular limestones were deposited. In the Mader Basin, sandy and calcareous turbidites suggest deeper water conditions in the late Devonian. During the Strunian/Tournaisian the whole area was overwhelmed by a thick deltaic sequence. The general facies distribution is in agreement with depositional models of other Upper Devonian and Lower Carboniferous cephalopod limestones in the European Variscan orogenic belts. In all these cases, condensed cephalopod limestones occupy a distinct palaeogeographic position in predictable facies sequences that reflect pre-orogenic phases in the Variscan geodynamic cycle. Moreover, close parallels exist with condensed sequences in the Triassic and Jurassic that occur in a very similar position within the Alpine orogenic cycle.     

泥盆系弗拉阶/法门阶之交事件沉积和海平面变化

在华南板块南部被动大陆边缘和西伯利亚板块南部活动大陆边缘弗拉阶/法门阶之交识别出7种物理事件相: 细粒钙屑浊积岩相、钙质角砾岩相、软变形灰岩相、含微球粒的粒泥灰岩相、凝灰质砾岩相、黑色页岩相和火山喷溢相.它们在法门阶下Palmatolepis triangularis带底部和/或上Palmatolepis rhenana带底部具有广泛的分布, 可能是两次陨击事件记录, 是良好的高分辨率年代地层对比标志.弗拉期/法门期之交的海平面变化不具有同步性和一致性, 可分辨出2种型式(阶跃型和渐进型) 和5种状态(弗拉期最末期下降→法门期最初期上升、弗拉期最末期上升→法门期最初期下降、弗拉期最末期上升→法门期最初期上升、弗拉期最末期下降→法门期最初期下降、弗拉期最末期与法门期最初期海平面变化不明显).弗拉阶/法门阶之交生物的阶梯状绝灭可能与从上Palmatolepis rhenana带底部至Palmatolepis crepida带不均匀分布的6次陨击事件密切相关.      

广西玉林早泥盆世早期腕足动物生物地层研究

在广西玉林樟木地区早泥盆世北均塘组、良合塘组、樟木组３５属４８种腕足动物演化和地层分布特征研究的基础上,建立了６个腕足动物生物带（自下而上）：１．Ｑｕａｄｒｉｔｈｙｒｉｎａｅｘｐａｎｓａ－Ｐｒｏｄａｖｉｄｓｏｎｉａｍｉｒａｂｉｌｉｓ带;２．Ｇｒａｙｉｎａｍａｇｎｉｆｉｃａｇｕａｎｇｘｉｅｓｉｓ－Ｓｃｈｉｚｏｐｈｏｒｉａｓｕｌｃａｔｉｎａ带;３．Ｒｅｔｉｃｕｌａｒｉｏｐｓｉｓｃｏｓｔａｔｕｓ带;４．Ｇｒａｙｉｎａｍａｇｎｉｆｉｃａｓｉｎｉｃａ带;５．Ｑｕａｄｒｉｔｈｙｒｉｓｒｏｂｕｓｔａ－Ｂｏｕｃｏｔｉａ带;６．Ｑｒｉｅｎｔｏｓｐｉｒｉｆｅｒｗａｎｇｉ带。依据国际对比和与笔石的层位关系认为：第１－３带为晚洛赫柯夫期,并且１、２、３带很可能分别相当晚洛赫柯夫早、中、晚期;第４带为早布拉格早期;第５、６带分别相当中、晚布拉格期。     

湖北崇阳二叠纪—三叠纪之交生物灭绝和沉积微相演化

二叠纪—三叠纪之交,湖北崇阳地区处于浅水碳酸盐岩台地环境.二叠纪末的全球事件在该剖面的沉积微相和生物演化上均留下了清楚的记录.二叠纪末生物大灭绝之前,崇阳地区为典型的正常浅海台地环境,生物种类多样,数量丰富,主要生物化石有钙藻、有孔虫、腕足、棘皮类和海绵等.生物大灭绝之后,钙藻、(筳)类、棘皮类、海绵、绝大部分有孔虫开始消失,取而代之的是个体微小的腹足、介形虫和大量的蓝细菌化石.大灭绝界线之上,首先出现的是25 cm厚的纹层状的微生物岩,含较丰富的种类单调的有孔虫化石.之后逐渐相变为花斑状微生物岩和穹隆状微生物岩,厚度分别为6.4,2.3m.不同类型微生物岩在结构构造和生物组成上存在差别.微生物岩沉积结束之后,则相变为浅滩相鲕粒灰岩.共划分出3种沉积相,即开阔台地相、潮坪相和浅滩相.崇阳剖面的生物灭绝和沉积微相变化是二叠纪—三叠纪之交浅水台地环境生物与环境过程的典型代表,为认识二叠纪末浅海沉积相演化和全球事件提供了新材料.     

贵州紫云晚石炭世叶状藻礁灰岩微相特征与沉积环境的研究

紫云晚石炭世叶状藻礁发育在碳酸盐岩台地边缘内。研究以碳酸盐岩微相分析为主要手段,对叶状藻礁剖面中岩石的微相特征和沉积环境研究。主要总结出6个微相类型,为生物碎屑粒泥灰岩、生物碎屑泥粒灰岩、球粒生屑泥粒灰岩、叶状藻粒泥灰岩—叶状藻泥粒灰岩、腕足泥粒灰岩、生物碎屑颗粒灰岩。总结了一个在开阔台地与台地边缘过渡带上的局部相模式,由较深水的开阔台地相、叶状藻礁相、台内斜坡相和台内较浅水的斜坡相、较浅水的开阔台地相、台内浅滩相过渡,分析总结了叶状藻礁的沉积环境。微相组合及其在剖面纵向分布规律显示,剖面的总体沉积环境为潮下带浅水开阔台地环境。      

贺兰山北段石炭纪和二叠纪植物群

贺兰山北段石炭纪和二叠纪植物化石经鉴定,计有39属104种。讨论了植物群的性质,划分了5个组合,即(1)晚石炭世早期Bothrodendroncirculare—Mesocalamitescistiformis组合;(2)晚石炭世中期Lepidodendronsubrhombicum—Conchophyllumrichthofenii组合;(3)晚石炭世晚期Lepidodendronszeianum—Neuropterisovata组合;(4)早二叠世早期Lepido dendronposthumii—Callipteridiumkoraiense组合和(5)早二叠世晚期Caulopteriswudaensis—Paratingiadatongensis组合。这5个组合的代表岩组分别为红土洼组、羊虎沟组、太原组下部、太原组中上部和山西组。其地质时代大致相当于纳缪尔B—C期、维斯发期、斯蒂芬期、阿赛尔期、萨克马尔期和阿丁斯克期。植物群含有大量的华夏型分子,为典型的华夏植物群。此外还讨论了植物群的演化。     

西藏申扎地区奥陶纪鹦鹉螺化石

申扎地区塔尔玛桥东－扎扛一带的奥陶系是藏北地区有可靠古生物化石依据的第二个出露区。地层发育好,含化石门类多,数量大,时代依据可靠,界线清楚,是藏北地区目前已知研究奥陶系（纪）的最理想地区。经鉴定,该区鹦鹉螺化石有29种,10未定种,分别隶属3目8科15属,其中3新属9新种。这为该区奥陶纪地层划分、对比及研究古地理格局,探讨藏北地质构造演化提供了重要资料。文中仅描述了1新属6新种,老属种描述从略。     

重庆北碚长兴期腕足动物群落的划分及其地层意义

重庆北碚地区晚二叠世长兴期地层出露良好,剖面位于北碚代家沟东北部,经过详细调查研究,在厚度为110.5m的长兴组石灰岩地层中发现有丰富的腕足动物,共有28属46种。按照识别与划分化石群落的5个基本条件,本区长兴期腕足动物可以划分为3个化石群落,本区长兴期生态地层建立3个化石群落带,自上而下为:Uncinunellina-Peltichia带、Haydenella-Perigeyerella带、Oldhamina-Meekella带。从而将本区长兴期地层三分。