Vertical versus horizontal growth strategies of coral frameworks (Tulamben, Bali, Indonesia)

Two different coral framework structures located in a shallow subtidal area on the east coast of Bali are described in this study. One structure is a typical coral carpet with a distinct internal succession of coral taxa and growth forms. It starts with a variety of coral species exhibiting massive, tabular, branching, and platy growth forms settling on volcanic boulders and cobbles. The main body of the coral carpet is composed almost monospecifically of Acropora cf. vaughani, which has filled all accommodating spaces up to the low-water sea level. Mostof this carpet died during the bleaching event of 1998 and the resultant dead Acropora framework is now capped by a platy Montipora assemblage. Some of the Acropora branches within the dead carpet, however, are still alive and display active growth. The Montipora cover protects the dead Acropora framework against mechanical and biological destruction. The few still growing Acropora branches may also contribute to the strength of the framework. The second coral framework is made up almost monogenerically of Montipora. One species of Montipora is of a laminar growth form and produces whorl-like colonies. Within this framework, only part of the Montipora colonies are dead; however, these are intensively fragmented. The fragments have been rapidly settled by a platy Montipora species, which has stabilized the fragments. In this case, the fragment shedding of the Montipora offers the opportunity for progradation of the framework on these fragments. Concerning the Acropora carpet, similar examples from the fossil record of the Miocene era of Spain and Austria have been reported.     

Ludlow (Silurian) stromatoporoid biostromes from Gotland, Sweden: facies, depositional models and modern analogues

Stacked stromatoporoid‐dominated biostromes of the Ludlow‐age Hemse Group (Silurian) in eastern Gotland, Sweden, are 0·5–5 m thick and a few tens of metres to >1 km in lateral extent. They form one of the world's richest Palaeozoic stromatoporoid deposits. This study compiles published and new data to provide an overall facies model for these biostromes, which is assessed in relation to possible modern analogues. Some biostromes have predominantly in‐place fossils and are regarded as reefs, but lack rigid frameworks because of abundant low‐profile non‐framebuilding stromatoporoids; other biostromes consist of stromatoporoid‐rich rudstones interpreted here as storm deposits. Variation between these two `end‐members' occurs both between interlayered biostromes and also vertically and laterally within individual biostromes. Such variation produces problems of applying established reef classification terms and demonstrates the need for the development of terminology that recognizes taphonomic destruction of reef fabrics. An approach to such terminology is found in all four categories of a recent biostrome classification scheme that are easily recognized in the Hemse biostrome facies: autobiostromes (>60% in place); autoparabiostromes (a mixture of in‐place and overturned reef‐building organisms, 20–60% in place); parabiostromes (builders are overturned and damaged, <20% in place); and allobiostromes (transported and detrital reef material, nothing in place). These categories provide a broad taphofacies scheme for the Hemse biostromes, which are mostly autoparabiostrome to allobiostrome. The biostromes developed on crinoidal grainstone sheets and expanded laterally across relatively flat substrates in a marine setting of low siliciclastic input. Planar erosion surfaces commonly terminate biostrome tops. Three broadly similar modern analogues are identified, each of which has elements in common with the Hemse biostromes, but none of which is an exact equivalent: (a) laterally expanded and coalesced back‐barrier patch reefs behind the Belize barrier, an area influenced by limited accommodation space; (b) a hurricane‐influenced shelf, interpreted for Grand Cayman, where reef cores consist of rubble and lack substantial framework; the wide distribution of rounded pebbles and cobbles of stromatoporoids in the Hemse biostromes most probably resulted from hurricanes; (c) coral carpets in 5–15 m water depth of the northern Red Sea, where lateral expansion of low‐diversity frames dominated by Porites coral has produced low‐profile biostromes up to 8 m thick and several km long. Such carpets accumulated large amounts of carbonate, with little export, as in the Hemse biostromes, although the latter did not build frameworks because of the nature of growth of the stromatoporoids. The notable lack of algae in the Hemse biostrome facies is also a feature of Red Sea coral carpets; nevertheless, coral carpets are ecologically different. Hemse biostromes lack evidence of a barrier reef system, although this may not be exposed; the facies assemblage is consistent with either a storm/hurricane‐influenced mid‐ to upper ramp or back‐barrier system.     

Quaternary coral-reef terraces from Kish and Qeshm Islands, Persian Gulf: new radiometric ages and tectonic implications

A survey of raised coral reef terraces in locations along the southern coast of Iran was carried out with the aim of assessing regional Late Quaternary tectonic uplift rates influenced by salt doming. Two islands were studied: Kish, where no previous data were available, and Qeshm, where a previous survey had already attributed the lowest step in two sequences of raised marine terraces to the Last Interglacial.Twenty-five ESR and seven Th/U ages were obtained from Kish Island. The results show that this flat, about 32 m high, gently domed island was totally submerged during the last two interglacial periods. Corals ascribed to MIS 5 and MIS 7 have been dated at the same elevations, near the present sea level, and in the uppermost, inner part of the island, giving evidence of a polycyclic origin for the island surface deposits. Following a discussion on the possible position of eustatic peaks during MIS 7, a maximal average uplift rate of 0.20±0.02 mm/yr has been deduced from the above data.Furthermore, the survey obtained some new results from Qeshm, where seven ESR ages confirmed the MIS 5 age of the lowest raised marine terrace (that also corresponds to an uplift rate of about 0.2 mm/yr), failing, however, to date older steps, due to significant coral-sample re-crystallisation. In the western part of Qeshm, five new radiocarbon ages of elevated beach material demonstrated the variability of uplift rates even along short distances around an active salt dome.     

An integrate analysis of an Hauterivian coral biostrome from the Agrio Formation,Neuquén Basin,west-central Argentina

A coral biostrome from the lower Hauterivian of the Neuquén Basin, west-central Argentina, was examined in detail. Taxonomic analysis established it as a monospecific assemblage of the scleractinian colonial coral Stereocaenia triboleti (Koby). Palaeoecological analysis of the species allows us to infer that it was an opportunistic species, capable of thriving in mixed carbonate-siliciclatic environments, on a soft substrate under a moderate to low sedimentation rate showing great regenerative capacity. Taphonomic analysis, together with lithofacies and microfacies analysis, allowed us to establish that the coral biostrome was deposited in a middle ramp setting under open marine and well-oxygenated conditions. Corals colonized a soft substrate during a transgressive phase and reaching its climax in the maximum flooding zone under lowest clastic input. They managed to form a low coral meadow of ramose forms of up to 30 cm in height. Coral remains were exposed on the interface after death and thus were subject to severe encrustation and bioerosion. The coral biostrome is interpreted as a mixed biogenic-sedimentologic skeletal concentration deposited by an interplay of an initial biogenic concentration through later episodes of physical reworking, probably storms, which yielded a largely parautochthonous fossil assemblage with minimum lateral transport. As sedimentation rate increased during a highstand interval, the coral-dominated community could not cope with it and was finally buried. The described coral biostrome recorded at the base of a shallowing-upwards cycle closely resembles coral biostromes described from the Hauterivian of Peru and Turkmenistan, and differs significantly from framework reef facies recorded in the Hauterivian of the Tethys region from the Paris Basin to Crimea.     

Southern dispersal and Palaeoecological implications of woolly rhinoceros (Coelodonta antiquitatis): review of the Iberian occurrences

Cold-adapted large mammal populations spread southward during the coldest and driest phases of the Late Pleistocene reaching the Iberian Peninsula. Presence of woolly rhinoceros (Coelodonta antiquitatis) can be identified from 23 Iberian sites, which is compiled and analyzed herein, and the fossil specimens from seven of these sites are described here for first time.Morphological and biometrical analyses demonstrate that the Iberian woolly rhinoceros did not significantly differ from individuals of other European populations, but represent the westernmost part of a continuous Eurasian belt of distribution.The first presence of woolly rhino in the Iberian Peninsula has been identified during the late Middle Pleistocene and early Late Pleistocene. However, the highest abundance of this species is recorded during MIS 3 and 2. The latest Iberian occurrences can be dated around 20 ka BP. The presence of woolly rhinoceros in the Iberian Peninsula correlates with periods of extreme dry and cold climatic conditions documented in Iberian terrestrial and marine sediment sequences.From a palaeobiogeographic point of view, the maximum southern spread of C. antiquitatis on the Iberian Peninsula was registered during the late Middle Pleistocene or early Late Pleistocene, reaching the latitude of Madrid (about 40°N). Subsequently, during MIS 3 and 2, all Iberian finds were restricted to the Northern regions of Iberia (Cantabrian area and Catalonia). The southern expansion of C. antiquitatis during the Late Pleistocene in the Iberian Peninsula reached similar latitudes to other Eurasian regions.The ecological composition of fossil assemblages with presence of woolly rhinoceros was statistically analyzed. Results show that temperate ungulate species are predominant at Iberian assemblages, resulting in a particular mixture of temperate and cold elements different of the typical Eurasian cold-adapted faunal associations. This particular situation suggests two possible explanations: a) Eventual migrations during the coldest time spans, resulting in a mixing of cold and temperate faunas, instead a faunal replacing; b) Persistence of woolly rhinoceros populations in the Iberian Peninsula during interglacial episodes confined at cryptic southern refugia.     

Microencruster-microbial-cement framework of the Upper Jurassic reef developed on the slope of the intra-Tethyan carbonate platform (Bulgaria)

The carbonate succession in the Lyalintsi section of the western Moesian Platform (western Bulgaria) displays a shallowing-upward trend. Growth of the Tithonian–Valanginian coral biostromes and low-relief bioherms was preceded by Oxfordian–Kimmeridgian sedimentation of fine-grained peloidal-bioclastic limestones and Saccocoma-bearing limestones on the homoclinal ramp and the carbonate platform slope. In the late Kimmeridgian, boundstones with very rare corals, but with easily recognisable biohermal morphology, were developed. The main components of this reef are encrusting microorganisms, microbial crusts and synsedimentary cements. Microencrusters Labes atramentosa, Crescentiella morronensis, Perturbatacrusta leini and Radiomura cautica, as well as thin crusts of calcified sponges (sclerosponges), are the main biotic components. Corals (almost exclusively microsolenids) are sparse, whereas photophilic microencrusters (e.g., “Lithocodium–Bacinella”), are absent, although they are common in the overlying shallow-water part of the Lyalintsi sequence. Microbialites and synsedimentary cements provided additional support for the reef framework. The framework, especially the biotic components, and the reefal facies position within the sedimentary succession, implies that the high-energy upper slope of the carbonate platform was the depositional setting of the microencruster-microbial-cement reef studied. Encrusting microorganisms, except for C. morronensis and sponges, are only known from the intra-Tethyan platforms. This study supports conclusion of studies of coeval Alpine reefs that the presence of the microencruster-microbial-cement framework provides insight into the palaeobathymetry, palaeogeography and tectonic configuration of the intra-Tethyan carbonate platforms.     

Lithic assemblage from the Jingshuiwan Paleolithic site of the early Late Pleistocene in the Three Gorges,China

The Jingshuiwan Paleolithic site lies in the second terrace of the Changjiang (Yangtze) River and has a complete geomorphological section. Archaeological materials from early Late Pleistocene fluvial deposits of silt and sand are dated by optically stimulated luminescence (OSL) to ca. 70 ka. The stone assemblage from layer 7 includes retouched tools (118), cores (304), flakes (281), flake fragments (101), stone hammers (four) and chunks (102). Artifacts were made from lithic sources locally available from the former riverbed. The main type used was silicarenite; quartzite, hypabyssal intrusive rock, extrusive rock and volcanic breccia were also used. The principal flaking technique was direct hammer percussion without prepared striking platforms. Major blanks for tool fabrication were complete flakes (67.0%), followed by cores and incomplete flakes. Most tools were large. Chopper-chopping tools and scrapers were the dominant tool types, followed by points and notches. Modified tools were mostly retouched unifacially on the surface of blanks by direct hammer percussion. Jingshuiwan provides evidence that South China was occupied during MIS 4. Because of the similarity of the stone tool assemblage with earlier ones associated with Homo erectus, it may also provide indirect evidence that H. erectus persisted into the early Late Pleistocene.     

Organic (coral) buildups in the carboniferous at the southwestern margin of the East European Platform

Organic buildups (biostromes and bioherms) were established for the first time in the coal-bearing formation of the Lvov-Volynsk coal basin. Colonial rugose corals of the Siphonodendron genus represented the main bioherm-builders. Coral buildups were formed under shallow marine conditions and located within the active tectonic region at the southwestern margin of the East European Platform. The region represents a northwest-trending band (10–12 km wide) extending over more than 70 km. The studied late Visean coral bioherms and biostromes are similar to coeval ones from some European regions.     

Mesophotic coral buildups in a prodelta setting (Late Eocene,southern Pyrenees,Spain): a mixed carbonate–siliciclastic system

Lower Priabonian coral bioherms and biostromes, encased in prodelta marls/clays, occur in the Aínsa‐Jaca piggyback basin, in the South Central Pyrenean zone. Detailed mapping of lithofacies and bounding surfaces onto photomosaics reveals the architecture of coral buildups. Coral lithosomes occur either isolated or amalgamated in larger buildups. Isolated lithosomes are 1 to 8 m thick and a few hundred metres wide; clay content within coral colonies is significant. Stacked bioherms form low‐relief buildups, commonly 20 to 30 m thick, locally up to 50 m. These bioherms are progressively younger to the west, following progradation of the deltaic complex. The lowermost skeletal‐rich beds consist of bryozoan floatstone with wackestone to packstone matrix, in which planktonic foraminifera are abundant and light‐related organisms absent. Basal coral biostromes, and the base of many bioherms, consist of platy‐coral colonies ‘floating’ in a fine‐grained matrix rich in branches of red algae. Corals with domal or massive shape, locally mixed with branching corals and phaceloid coral colonies, dominate buildup cores. These corals are surrounded by matrix and lack organic framework. The matrix consists of wackestone to packstone, locally floatstone, with conspicuous red algal and coral fragments, along with bryozoans, planktonic and benthonic foraminifera and locally sponges. Coral rudstone and skeletal packstone, with wackestone to packstone matrix, also occur as wedges abutting the buildup margins. Integrative analysis of rock textures, skeletal components, buildup anatomy and facies architecture clearly reveal that these coral buildups developed in a prodelta setting where shifting of delta lobes or rainfall cycles episodically resulted in water transparency that allowed zooxanthellate coral growth. The bathymetric position of the buildups has been constrained from the light‐dependent communities and lithofacies distribution within the buildups. The process‐product analysis used here reinforces the hypothesis that zooxanthellate corals thrived in mesophotic conditions at least during the Late Eocene and until the Late Miocene. Comparative analysis with some selected Upper Eocene coral buildups of the north Mediterranean area show similarities in facies, components and textures, and suggest that they also grew in relatively low light (mesophotic) and low hydrodynamic conditions.     

A new record of Pleistocene hippopotamus from River Severn terrace deposits, Gloucester, UK—palaeoenvironmental setting and stratigraphical significance

A new Pleistocene vertebrate assemblage from fluvial deposits of the River Severn in Gloucester, England, has yielded the remains of hippopotamus (Hippopotamus amphibius), a new record for this terrace system, with additional material from probable bison (cf. Bison priscus) and elephant (Elephantidae sp.). The presence of these taxa indicates fully temperate climatic conditions and the occurrence of hippopotamus, a significant biostratigraphical indicator for the British Late Pleistocene, suggests an age for the assemblage within MIS 5e (the Last Interglacial). This would contradict the older MIS 7-6 age for the gravel body that is currently accepted on the basis of deposit mapping and imply a more complex mode of deposition than presently envisaged in the valley.     

Heavy metals contamination of the Quaternary coral reefs, Red Sea coast, Egypt

In order to assess pollutants and impact of environmental changes along the Egyptian Red Sea coast, seven recent and Pleistocene coral species have been analyzed for Zn, Pb, Mn, Fe, Cr, Co, Ni, and Cu. Results show that the concentration of trace elements in recent coral skeletons is higher than those of Pleistocene counterpart except for Mn and Ni. In comparison with recent worldwide reefs, the present values are less than those of Central America coast (iron), Gulf of Aqaba, Jordan (lead, copper), Gulf of Mannar, India (chromium, zinc, manganese), Costa Rica, Panama (chromium, nickel), North-west coast of Venezuela and Saudi Arabia (copper). The present values are higher than those of Gulf of Aqaba, Jordan (iron, zinc, manganese), Gulf of Mannar, India (lead, cobalt, nickel), North-west coast of Venezuela (lead, zinc, chromium, manganese), Australia (copper, nickel, zinc, manganese). The highest values were recorded in Stylophora pistillata (iron, lead and copper), Acropora cytherea (cobalt), Pocillopora verrucosa (zinc) and the lowest concentrations were recorded in Goniastrea pectinata (iron, chromium, copper and nickel), Favites pentagona (lead, zinc and manganese), and Porites lutea (cobalt). The differences in metals content among the studied species are attributed to differences in microstructure and microarchitecture.     

Recognizing cryptic environmental changes by using paleoecology and taphonomy of Pleistocene bivalve assemblages in the Oga Peninsula,northern Japan

Multivariate analyses applied to Pleistocene bivalve assemblages from the Oga Peninsula (northern Japan) discriminate three distinct assemblages. The assemblages and their taphonomy were used to recognize environmental settings and changes. The Astarte–Cyclocardia–Glycymeris assemblage indicates shelf environment (below the storm wave base) where gravels and shells are transported from shallower settings. Supply of the exotic coarse sediment probably enabled epifaunal bivalves to inhabit the sea floor. The Glycymeris assemblage is characterized by dominance of G. yessoensis and represents current-swept shoreface environment (above the storm wave base). The Moerella assemblage is characterized by bivalves inhabiting bay to open-marine conditions and diverse deposit-feeders, indicating a moderately land-locked environment, such as an open bay or a bay mouth. Fine-grained substrata rich in organic matters in the bay were probably suitable for the deposit-feeders. Ordination also shows the assemblages along two environmental gradients, a bathymetrical one and the other related to open-marine and bay conditions. The environmental changes are explained mainly by glacio-eustatic sea-level changes and alternation of coastal geomorphology caused by local crustal movements. This study also suggests that fossil assemblages can be a powerful tool to reconstruct environments and depositional dynamics even in intensely bioturbated sedimentary facies.     

Geoarchaeology of Pleistocene open-air sites in the Vila Nova da Barquinha-Santa Cita area (Lower Tejo River basin, central Portugal)

This paper aims to provide insight into human occupation and landscape change during the Pleistocene in a central area of the Lower Tejo basin (Portugal). Detailed geomorphological mapping, coupled with lithostratigraphy, sedimentology and luminescence dating, supports the identification of a complete terrace staircase sequence. It consists of six gravely terraces located below the culminant (Pliocene) basin unit. A chronological framework for the sedimentary sequences and associated human industries is proposed and correlated with marine oxygen isotope stages (MIS): T1 terrace, not dated; T2, not dated; T3, >300 ka; T4, ∼300-160 ka (MIS8, MIS7 and MIS6); T5, ∼136-75 ka (MIS5); T6, ∼62-30 ka (MIS3); colluvium and aeolian sands, ∼30-14 ka (MIS2); valley fill deposits, ∼14 ka to present (MIS1). The oldest artefacts were found at the base of the T4 terrace, with the local stratigraphic level dated to ≥175 ± 6 ka (Middle Pleistocene). The lithic assemblages collected from distinct stratigraphic levels (T4, T5 top, T6 terraces and colluvium) are characterized by the predominance of opportunistic technological choices, a feature that can be attributed partly to the preferential exploitation of the available raw material, dominated by local-sourced quartzites and quartz pebbles. The adaptation to local raw material (texture and volume), together with subsistence patterns and behaviours, could explain the rarity of Acheulian types (handaxes and cleavers) and picks in the T4 terraces of the Tejo tributaries; this is in contrast to the same terrace of the Tejo valley, in which these types are found. Interpretation of the environmental conditions (controlled by climate and glacio-eustatic sea-level changes) affecting the hunter-gatherer human groups is also presented.     

Late Quaternary environmental and human events at En Gedi, reflected by the geology and archaeology of the Moringa Cave (Dead Sea area, Israel)

The Moringa Cave within Pleistocene sediments in the En Gedi area of the Dead Sea Fault Escarpment contains a sequence of various Pleistocene lacustrine deposits associated with higher-than-today lake levels at the Dead Sea basin. In addition it contains Chalcolithic remains and 5th century BC burials attributed to the Persian period, cemented and covered by Late Holocene travertine flowstone. These deposits represent a chain of Late Pleistocene and Holocene interconnected environmental and human events, echoing broader scale regional and global climate events. A major shift between depositional environments is associated with the rapid fall of Lake Lisan level during the latest Pleistocene. This exposed the sediments, providing for cave formation processes sometime between the latest Pleistocene (ca. 15 ka) and the Middle Holocene (ca. 4500 BC), eventually leading to human use of the cave. The Chalcolithic use of the cave can be related to a relatively moist desert environment, probably related to a shift in the location of the northern boundary of the Saharo-Arabian desert belt. The travertine layer was U-Th dated 2.46 ± 0.10 to 2.10 ± 0.04 ka, in agreement with the archaeological finds from the Persian period. Together with the inner consistency of the dating results, this strongly supports the reliability of the radiometric ages. The 2.46-2.10 ka travertine deposition within the presently dry cave suggests a higher recharge of the Judean Desert aquifer, correlative to a rising Dead Sea towards the end of the 1st millennium BC. This suggests a relatively moist local and regional climate facilitating human habitation of the desert.     

Environmental changes and growth history of a cold-water carbonate mound (Propeller Mound,Porcupine Seabight)

On- and off-mound sediment cores from Propeller Mound (Hovland Mound province, Porcupine Seabight) were analysed to understand better the evolution of a carbonate mound. The evaluation of benthic foraminiferal assemblages from the off-mound position helps to determine the changes of the environmental controls on Propeller Mound in glacial and interglacial times. Two different assemblages describe the Holocene and Marine Isotope Stage (MIS) 2 and late MIS 3 (∼31 kyr BP). The different assemblages are related to changes in oceanographic conditions, surface productivity and the waxing and waning of the British Irish Ice Sheet (BIIS) during the last glacial stages. The interglacial assemblage is related to a higher supply of organic material and stronger current intensities in water depth of recent coral growth. During the last glaciation the benthic faunas showed high abundances of cassidulinid species, implying cold bottom waters and a reduced availability of organic matter. High sedimentation rates and the domination of Elphidium excavatum point to shelf erosion related to sea-level lowering (∼50 m) and the progradation of the BIIS onto the shelf. A different assemblage described for the on-mound core is dominated by Discanomalina coronata, Gavelinopsis translucens, Planulina ariminensis, Cibicides lobatulus and to a lower degree by Hyrrokkin sarcophaga. These species are only found or show significantly higher relative abundances in on-mound samples and their maximum contribution in the lower part of the record indicates a higher coral growth density on Propeller Mound in an earlier period. They are less abundant during the Holocene, however. This dataset portrays the boundary conditions of the habitable range for the cold-water coral Lophelia pertusa, which dominates the deep-water reefal ecosystem on the upper flanks of Propeller Mound. The growth of this ecosystem occurs during interglacial and interstadial periods, whereas a retreat of corals is documented in the absence of glacial sediments on-mound. Glacial conditions with cold intermediate waters, a weak current regime and high sedimentation rates provide an unfavourable environmental setting for Lophelia corals to grow. A Late Pleistocene decrease is observed in the mound growth for Propeller Mound, which might face its complete burial in the future, as it already happened to the buried mounds of the Magellan Mound province further north.     

A new site from the Spanish Middle Pleistocene with cold-resistant faunal elements: La Parte (Asturias,Spain)

The La Parte (Asturias) northern Spain site contains a cold-adapted mammal faunal assemblage that corresponds to a level radiometrically dated to a minimum age of 150 ka. It represents the most ancient site with cold resistant fauna in the Iberian Peninsula.Among the species recorded in La Parte, Coelodonta antiquitatis and Rangifer tarandus represent the typical cold-adapted large mammal association.The presence of the woolly rhino at La Parte provides relevant environmental information about the Mammoth Steppe. This species was not as abundant in the Middle Pleistocene as during the Late Pleistocene, when its cold-adapted features can be studied from an evolutionary perspective. The remains attributed to R. tarandus suggest an open steppe ecosystem.The presence of Marmota sp. is also confirmed in La Parte. A first interpretation would suggest an alpine or periglacial ecosystem, but based on recent results which do not support the correlation between phylogeny and climatic tolerance for extant species of Marmota (López and Cuenca, 2002. Palaeogeography, Palaeoclimatology, Palaeoecology 186, 311), we propose the presence of this rodent suggests an open landscape with cold conditions.The rest of the taxa included in the La Parte assemblage (Crocuta crocuta, Panthera leo, Cervus elaphus, Megaloceros cf. giganteus, Bison priscus and Equus caballus) are usually associated with typical cold-adapted faunas, but they are also found in woody temperate zones so they do not characterize by themselves a cold period. The faunal association from La Parte suggests a combination of steppic and open woodland ecological conditions.     

Comparison of ESR and TIMS U/Th dating of marine isotope stage (MIS) 5e, 5c, and 5a coral from Barbados—implications for palaeo sea-level changes in the Caribbean

Electron spin resonance (ESR) dating of coral has become an efficient geochronological tool in supporting morphostratigraphic studies carried out on Barbados during the last 10 years. The newly developed approach for D_E determination (D_E–D_max plot procedure) improves the precision of ESR dating of Pleistocene coral, and therefore permits differentiation between the main marine isotope stages (MIS) 5, 7, 9 and 11 and also between sub-stages 5e, c and a. This study compares results of ESR and TIMS Uranium series dating (U/Th) of emergent Last Interglacial coral reef terraces from Barbados, and presents some implications for the timing and extent of sea-level changes during marine isotope stages (MIS) 5e, c and a. Both dating methods indicate a distinct formation of up to three coral reef terraces during MIS 5e, at approximately 132 ka (ESR) to 128 ka (U/Th), at c. 128 ka (ESR) and at c. 120 ka (U/Th) to 118 ka (ESR). It is also highly probable that three reef terraces were developed during MIS 5c between c. 103 ka (U/Th) and 105 ka (ESR). The formation of two separate coral reefs during MIS 5a is recognized for the first time on Barbados, with an age estimate for the older MIS 5a-2 reef of 85 ka (ESR) or 84 ka (U/Th), and an age estimate for the younger MIS 5a-1 reef terraces of 74 ka (ESR) or 77 ka (U/Th). Assuming a constant uplift rate of 0.276 m/ka at the south coast of Barbados, sea-level reached its maximum during MIS 5e-3 and MIS 5e-2 between 132 and 128 ka ago. After this, sea-level declined reaching a level of c. −11 m below present sea level approx. 118–120 ka ago (MIS 5e-1). During the substage 5c sea-level was generally lower than in substage 5e. It reached relative maxima at c. −13, −20 and −25 m during MIS 5c (approx. 105 ka) and formed three distinct coral reef terraces probably in relative short time intervals. For the first time, a double sea-level oscillation is recognized on Barbados during MIS 5a: an early MIS 5a-2 (c. 85 ka) with a sea-level places at approx. −21 m, and a late MIS 5a-1 sub-stage (c. 74 or 77 ka) with a sea-level at approx. –19 m below present sea level.     

Carbon and oxygen stable isotope compositions of late Pleistocene mammal teeth from dolines of Ajoie (Northwestern Switzerland)

Fossils of megaherbivores from eight late Pleistocene ¹⁴C- and OSL-dated doline infillings of Ajoie (NW Switzerland) were discovered along the Transjurane highway in the Swiss Jura. Carbon and oxygen analyses of enamel were performed on forty-six teeth of large mammals (Equus germanicus, Mammuthus primigenius, Coelodonta antiquitatis, and Bison priscus), coming from one doline in Boncourt (~ 80 ka, marine oxygen isotope stage MIS5a) and seven in Courtedoux (51–27 ka, late MIS3), in order to reconstruct the paleoclimatic and paleoenvironmental conditions of the region. Similar enamel δ¹³C values for both periods, ranging from − 14.5 to − 9.2‰, indicate that the megaherbivores lived in a C₃ plant-dominated environment. Enamel δ¹⁸O_PO4 values range from 10.9 to 16.3‰ with a mean of 13.5 ± 1.0‰ (n = 46). Mean air temperatures (MATs) were inferred using species-specific δ¹⁸O_PO4–δ¹⁸O_H2O-calibrations for modern mammals and a present-day precipitation δ¹⁸O_H2O-MAT relation for Switzerland. Similar average MATs of 6.6 ± 3.6°C for the deposits dated to ~ 80 ka and 6.5 ± 3.3°C for those dated to the interval 51–27 ka were estimated. This suggests that these mammals in the Ajoie area lived in mild periods of the late Pleistocene with MATs only about 2.5°C lower than modern-day temperatures.     

Vegetation and climate changes during the Eemian and Early Weichselian in the Upper Volga region (Russia)

Ples is the key-section that represents the Mikulino (Eemian) interglaciation and early part of the Valdai (Weichselian) glacial epoch in the north-central Russia. Pollen profile of this section reflects spread of the broad-leaved forest dominated by Quercus, Ulmus, and Tilia with participation of Acer, Fraxinus and Carpinus in the optimum phase of the interglaciation. A gradual cooling and increasing humidity of climate brought about a decline in the broad-leaved forest and development of dark-coniferous communities (Picea-Abies-Pinus sibirica) in the end of the interglaciation. It was followed by Scots pine and birch forest, and then replaced by communities of tundra shrubs with patches of the open birch forest. During the first post-Eemian cooling the climatic conditions were not uniform, as a short climatic amelioration separated it into two stages. This minor warming is indicated by an increase in birch woodland in complex plant cover. This first Early Weichselian cold stage was followed by a warm interstade, when mixed spruce-birch, and later larch-pine and spruce-pine forests occupied the region. This interstade, known as Verkhnevolzhskiy (Upper Volga) in the climatostratigraphic subdivision of the Late Pleistocene in the Russian Plain (after Grichuk [1961. Fossil floras as the basis for the Quaternary stratigraphy. In: Markov, K.K. (Ed.), Rel`ef i Stratigrafiya Chetvertichnykh Otlozheniy Severo-zapada Russkoi Ravniny, Izdatel'stvo AN SSSR, Moscow, pp. 25–71]), corresponds to the Brörup interstade in the northern Europe, or to Marine Oxygen Isotope Substage (MIS) 5c (e.g. Mangerud [1989. Correlation of the Eemian and the Weichselian with deep sea oxygen isotope stratigraphy. Quaternary International 3, 1–4.]). On the whole, we correlate the sediment sequence at Ples to the MIS from 5e to 5c.     

Stratal,sedimentary and faunal relationships in the Coniacian 3rd-order sequence of the Iberian Basin,Spain

The Coniacian 3rd-order sequence in the Iberian Basin is represented by a carbonate ramp-like open platform. The biofacies is mainly dominated by nekto-benthic (such as ammonites) and benthic organisms (such as bivalves, mainly rudists) with scarce solitary corals (hermatypics are absent), showing major differences among the Transgressive System Tract (TST) and Highstand Normal Regression (HNR). During the TST, platform environments were dominated by Pycnodonte, other oysters and molluscs (with only subordinate rudists) and ammonites, which were represented by ornamented platycones (Tissotioides and Prionocycloceras), and by smooth oxycones (Tissotia and Hemitissotia). During the HNR, shallow water depositional areas were occupied by rudist-dominated associations. Storm- and wind-induced currents and waves acting on these associations produced large amounts of loose bioclastic debris that covered outer platform areas. This facies belt graded landwards into protected, lower-energy settings (inner platform, lagoon and littoral environments). Rudist biostromes were preserved in seaward areas of these protected shallow environments of overall moderate to low hydrodynamic gradient, which was punctuated by storms. In this environment and landwards, large areas of marly substrate favoured the presence of gastropods, other bivalves, echinoderms, benthic foraminifera and solitary corals. Because of the input of siliciclastics and, probably, the lack of nutrients in suspension, the establishment of rudist communities was difficult in more landward areas of the lagoon and in tidal environments. This heterozoan carbonate factory was thus controlled by warm-water conditions and high energy levels, which were responsible for high-nutrient contents in suspension.