人工增殖蓝点马鲛作为利用鳀鱼资源另一途径的分析

一、前言 本文提出一个雏议,旨在引起有关方面对开发利用鳀鱼Engraulis japonicus Temminck & Schegel资源的另一途径问题进行讨论,以期取得对这个问题的充分认识和作出可行性判断。 我国近海和外海拥有的鳀鱼资源非常丰富。根据著者等的调查,1983年夏季在渤海上层的鱼类生物量中,鳀鱼占四分之三(6月)和三分之一(7月)的高比例。朱德山、王为祥报道的调查结果表明,1984年11月—1985年4月,黄东海外海鳀鱼资源量达200万     

Changes of nutrient concentrations and N: P: Si ratios and their possible impacts on the Huanghai Sea ecosystem

To investigate the impacts of nutrient concentrations and N:P:Si ratios on the ecosystem of the Huanghai Sea (Yellow Sea), the current status and long-term variation of nutrients concentrations and ratios as well as phytoplankton community structure in the Huanghai Sea were collected and analyzed. The results reveal great annual and seasonal fluctuations in the nutrient concentrations and N:P:Si ratios during 1998-2008 with no clear pattern observed in the whole region. Yet on a seasonal scale of spring and in the coastal regions such as the Jiaozhou Bay and Sanggou Bay, the increase of DIN concentration and N:P ratio as well as the decrease of phosphate and silicate concentrations and Si:N ratios were relatively significant. Many pelagic ecosystem changes have occurred concurrent with these changes of the nutrient regime, such as the recent increase of primary production, changes of phytoplankton chlorophyll a biomass and abundance, an increase of eutrophication, and occurrence of HABs. In addition, new trends in the variation of nutrients seem to be developing in some particular transect such as 36°N, which suggests that long-term and systematic ecosystem monitoring in the Huanghai Sea is necessary.     

Geographical Variation of Body Size of Neocalanus cristatus, N. plumchrus and N. flemingeri in the Subarctic Pacific and Its Marginal Seas: Implications for the Origin of Large Form of N. flemingeri in the Oyashio Area

Size distributions of Neocalanus cristatus, N. flemingeri and N. plumchrus were investigated in the eastern and the western subarctic gyres and three marginal seas of the North Pacific during the diapause period to examine the geographical variation in body size of Neocalanus species and to clarify the origin of the large biennial N. flemingeri which has been observed in the Oyashio region. There were significant among region variations in body sizes for all three species of Neocalanus. Generally, the body sizes of the copepods were larger in the marginal seas and marginal areas of the open ocean. In the open ocean, the body sizes increased westward. These patterns of variation in the body sizes roughly correlated with local food availability. Distribution of biennial N. flemingeri was restricted to the Sea of Japan, the Okhotsk Sea and the Oyashio region. The large-sized biennial N. flemingeri were abundantly observed in the Okhotsk Sea, and the medium-sized biennial individuals were observed in the Sea of Japan. These facts strongly suggest that the large biennial N. flemingeri in the Oyashio region are advected from the Okhotsk Sea.     

Spatial and temporal characteristics of trophic structure of the southern Huanghai Sea fish communities

The spatial and temporal characteristics of trophic structure of fish communities in the southern Huanghai Sea were examined based on the data sampled from bottom trawl surveys conducted during the autumn of 2000 and the spring of 2001. Hierarchical agglomerative cluster method and bootstrap randomization were used to identify significant trophic groups for each fish assemblage in the southern Huanghai Sea. A total of six major trophic groups were identified within this system, which classified predators based upon location in the water column or prey size （ i. e. , benthic to pelagic predators or fish to small invertebrate prey predators）. The similarity level used to identify significant trophic groups in each assemblage ranged from 24% to 34%. Although planktivores were the dominant trophic group in each assemblage （60% - 79% ）, there were spatial and temporal variations in the trophic structure, which reflected the differences in the abundance and availability of dominant preys. Simplified food webs were constructed to evaluate the most important trophic relationships between the dominant prey taxa and the fishes in each assemblage within this system. Although there were some differences in the key prey species among different food webs, pelagic prey items （mainly euphausiids and copepods） represent the most important energetic link between primary producers and higher trophic level predators. The trophic level for most fishes was between 3 and d, and the weighted mean trophic level for each assemblage ranged from 3.3 to 3.4. Compared with previous study in the mid-1980s, there was an obvious downward trend in the trophic level for most fish species, which resulted mainly from the fluctuation in key prey species in the Huanghai Sea. The decrease in the importance of Japanese anchovy seems to be offset by other abundant prey species such as Euphausia pacifica and copepods （ mainly Calanus sinicus ） .     

Reproductive cycle and minimal length at sexual maturity of Engraulis encrasicolus (L.) in the Zrmanja River estuary (Adriatic Sea, Croatia)

The reproductive cycle of anchovy, Engraulis encrasicolus (L.), was studied from monthly random samples of purse seine catches. A total of 1477 anchovy specimens were collected from January to December 2003 in the Zrmanja River estuary (Novigrad Sea). The analysis was based on the temporal evolution of gonadosomatic index, mass and stage of gonads. The total length of anchovy ranged from 4.5 to 14.5 cm and mass from 0.56 to 19.80 g. Sex ratio was slightly different from 1:1; the females were insignificantly predominated (♂/♀ = 0.99). The period of reproductive activity was from April to September coinciding with the most developed stages of gonads as well as with the highest gonad weights, and gonadosomatic indices. To estimate the length at maturity, a sub sample of 454 anchovy was taken from May to July (peak of anchovy spawning period). The length at which 50% of anchovy were mature (L₅₀) was calculated to be 8.2 cm. The length–weight relationship of anchovy was described by the expression: W = 3.51 × 10⁻³ L_T^3.211 (r² = 0.998). The relationships between total length–standard length and total length–fork length are L_T = 1.1405L_S + 0.2420 and L_T = 1.0425 L_F + 0.3944, respectively.     

Transport and distribution of nutrients in anchovy spawning ground to the southern waters of Shandong Peninsula

INTRODUCTIONNutrients (N ,PandSi)areessentialforphytoplanktongrowthinmarineecosystem .Thetransportandcycling ,distributionandvariationofnutrientsaffectnotonlyonthestructureandfunctionofecosystem ,butalsoonthevariationofthebiologicalresourcesinthesea .ThesouthofShondongPeninsulaisoneofthecentralzonesofurbanandagriculturalactivities.Al thoughthereisnodirectlargeriverinput,seasonalvariationofchemicalenvironmentisobviousintheHaizhouBay .Inspringandsummer,thedistributionsofnutrientsareinflu…     

Interannual variations of the occurrence of epipelagic fish in the diets of the seabirds breeding on Teuri Island, northern Hokkaido, Japan

The diets of breeding seabirds can be a good monitor of marine environmental changes. From 1984 to 2001 we monitored the diets of black-tailed gulls (Larus crassirostris) (“surface foragers”), rhinoceros auklets (Cerorhinca monocerata) (“epipelagic divers”), and Japanese cormorants (Phalacrocorax filamentotus) (“bottom divers”) that breed on Teuri Island at the northern boundary of the Tsushima Warm current in the Sea of Japan/East Sea. Between 1984 and 1987, both the gulls and the auklets foraged on the sardine (Sardinops melanostictus), but after 1992, they switched to the anchovy (Engraulis japonica). This change might reflect the collapse of the sardine stock in the late 1980s. In the 1990s, the year-to-year variations of the percentage of anchovy in the diets of the three seabird species showed similar trends: High in 1994 and 1998–2001; and low in 1992–1993 and 1995–1997. The estimated stock size of the anchovy population in the Tsushima Current area was positively correlated with the percentage of mass of anchovy in the seabirds’ diets. Thus, the short-term annual changes of the total anchovy availability, which might reflect SST or the volume transport of Tsushima Current, possibly affected the seabirds diets on this island.     

东中国海环流及其季节变化的数值模拟

关于东中国海环流的研究，国内外学者已做了大量的工作。早期科学家们主要依赖于对温盐资料和少数测流资料的分析研究对渤、黄、东海的环流结构有了较系统和深入的认识。东中国海环流是由一个气旋式的“流涡”组成，东侧主要是北上的黑潮-对马暖流-黄海暖流及其延伸部分；西侧为南下的沿岸流系。黑潮对东中国海环流的影响是如此之大，以致于除了某些局部区域外，上述海域主要流系的冬、夏季分布形式比较相似而无本质上的差异(胡敦欣等，1993)。但本文所研究海域正处于世界上最显著的季风区，冬、夏季盛行风向基本相反，过渡季节(春、秋季)风向多变，风力减弱；海洋热盐结构季节变化明显(如冬季混合强，而夏季层化明显等)，这些因素都使得东中国海环流存在着较明显的季节变化。 自20世纪80年代以来，东中国海环流的数值模拟工作逐步展开，并已成为研究环流结构及其形成机制的强有力工具。但由于数值模式本身以及计算方案的缺陷(如有些学者用固定的风场、温盐场对东中国海环流进行诊断模拟等)和观测资料的不足，数值模拟的结果难以得到验证，渤、黄、东海的环流研究中仍有大量的问题存在争议，以待澄清。例如，台湾暖流的来源、流径;对马暖流的来源;夏季黄海暖流的流径以及黄海冷水团环流等均有不同的论述。对黄、东海环流季节变化的数值模拟工作也较少，多用冬、夏典型月份的风场强迫积分至稳定态，给出冬、夏季环流，这种做法值得商榷。三维环流模式很难在1个月内达到稳定态，尤其是夏季层化明显、风力减弱的情况下，非常定风场的影响更应引起人们的重视。 本文采用比较符合实际的计算方案，用年循环风场和海面热通量场为外强迫，对渤、黄、东海的环流及其季节变化进行了模拟，并对一些争议问题进行了探讨。     

黄、东海地理环境与环流系统分析——长江口及济州岛邻近海域综合调查研究报告(第二章)

简要介绍了黄海和东海的地理环境概况,着重分析调查海域的环流系统。有如下一些初步看法与结论。 台湾暖流的前缘混合水,可从长江冲淡水底层穿越而影响到苏北沿岸,直到32°N以北的浅水区域。对马暖流西侧的水体是东海混合水,而其东侧为黑潮分支。黄海暖流的流向在不同季节具有规律的摆动。黄海底层冷水团属于季节性水团,其强盛及消衰与温跃层的形成及消亡紧密相关。黄海底层冷水团与中部底层冷水并非每年彼此独立,它们的共同特征甚至比其差异更明显。夏季东海冷水不能借助爬升侵入黄海底层冷水团内部。在济州岛南部区域,中层的逆温、逆盐现象,是由黄海密度环流的扩散效应与东海冷水沿黄海底层冷水团边界的爬升这两个原因而形成的。     

Geographic structure of European anchovy: A nuclear-DNA study

Atlantic–Mediterranean anchovies were genetically characterized at two polymorphic nuclear loci (intron 6 of two creatine-kinase genes) and compared to reference Engraulis albidus and E. encrasicolus samples from the northern Western Mediterranean to provide new insights into their geographic structure. Northeastern Atlantic anchovy, represented by one sample from the Canary archipelago and one sample from the Alboran Sea, were genetically distinct from Mediterranean E. encrasicolus (Weir and Cockerham's ^θ = 0.027–0.311), indicating geographic isolation from either side of the Almería–Oran oceanographic front. Generally smaller genetic differences were evident among anchovy populations from different sub-basins in the Mediterranean (^θ = − 0.019–0.116), the genetic differences between Black Sea and Ionian Sea/Aegean Sea anchovies being the strongest (^θ = 0.002–0.116). There was no evidence of the presence of E. albidus in our samples outside Camargue (northern shore of the Western Mediterranean). However, a sample from the southern Western Mediterranean appeared to be genetically intermediate between E. albidus and Mediterranean E. encrasicolus, indicating possible hybridization. Anchovy from the Benguela current system off southern Africa possessed allele frequencies characteristic of E. albidus at one locus and Northeastern Atlantic anchovy at the other locus, suggesting past introgression.     

冬季黄海暖流西偏机理数值探讨

利用海洋数值模式(MITgcm)模拟了冬季黄海流场并对冬季黄海暖流西偏的机理进行了探讨。冬季黄海流场模拟试验表明,黄海暖流由济州岛以西约32.5°N,125°E附近进入黄海,然后沿着黄海深槽西侧70 m等深线附近向北偏西运动;海面高度调整对黄海暖流路径具有重要影响,沿着黄海暖流路径的海面高度梯度比周围海区大,由海面高度梯度产生的地转流引起的北向体积输运占总的北向体积输运的78%。狭长海湾地形控制试验表明,单纯的黄海地形分布不足以引起黄海暖流西偏。黄海典型断面试验与渤海、黄海、东海地形控制试验说明,黄海暖流进入黄海的地理位置对流场分布有重要影响,黄海暖流进入黄海的位置恰好位于深槽西侧地形坡度较大区域,在位涡守恒的约束下黄海暖流受地形捕获沿70 m等深线附近向北偏西运动;试验还表明,黄海暖流进入黄海的位置与东海北部环流和地形分布有关,在冬季风的作用下东海北部环流的一部分沿着地形陡坡进入黄海形成黄海暖流。由此认为,黄海、东海环流在其特殊地形的约束下对冬季风的响应和调整,是引起黄海暖流西偏的主要原因。     

Response of Japanese anchovy (Engraulis japonicus) to the Pacific Decadal Oscillation in the Yellow Sea over the past 400 a

Quantitative identification of long-term changes in the abundance of Japanese anchovy (Engraulis japonicus) in the Yellow Sea is particularly important for understanding evolutionary processes of the Yellow Sea ecosystem. Unfortunately, the driving mechanisms of climate variability on the anchovy are still unclear due to the lack of long-term observational data. In this study, we used the fish scale deposition rate in the central Yellow Sea to reconstruct the time series of the anchovy stock over the past 400 a. On this basis, we further explored the impacts of the Pacific Decadal Oscillation (PDO) on the anchovy. Our results show that the anchovy stock is positively correlated with the PDO on a decadal time scale. In addition, anchovy abundance was relatively high during 1620–1860 AD (the Little Ice Age, LIA), though in a state of constant fluctuation; anchovy abundance maintained at a relatively low level after ~1860 AD. In particular, followed by overfishing since the 1980s, the anchovy stock has declined sharply. Based on these findings, we infer that fluctuations of the anchovy stock may be regulated by basin-scale “atmosphere–ocean” interactions. Nevertheless, the role of overfishing should not be ignored.     

黄东海近代厌氧沉积物中鳀鱼鱼鳞沉积信息的空间分布及重建潜力

Many studies have revealed that anchovy has exhibited large variability in population size on decadal tim-escales. However, such works concerning anchovy population are mainly based on short historical catch records. In order to understand the causes of variability in fish stocks （natural and/or anthropogenic） and calibrate the error between catches and standing stocks, it is essential to develop long-term time series of fish stocks from the time when human impacts are minimal or negligible. Well preserved fish scales from sediment record are regarded as useful indicators revealing the history of fish population dynamics over the last centuries. Anchovy scales was first analyzed over the Yellow Sea and East China Sea and the largest abundance was found in the central South Yellow Sea where is regarded as the largest overwintering ground for Japanese anchovy （Engraulis japonicas）. Thus in the central South Yellow Sea, two cores covering the last 150 years were collected for estimating fish scale flux. The scale deposition rate （SDR） records show that the decadal scale SDRs were obviously coherent between cores with independent chronologies. Thecalibration of downcore SDRs to the standing stocks of anchovy further validated that SDR is a reliable proxy to recon-struct the long-term anchovy population dynamic in the central South Yellow Sea where anoxic conditions prevail in the sediment. When assembled with other productivity proxies, it would be expected that SDR could be associated with changes in oceanic productivity and may make a contribution to determine the forcing factors and elucidate the mechanism of the process in future.     

南黄海西部36°N断面生态环境特征及其季节变化

重点研究了南黄海西部36°N断面水文、生源要素的四季变化特征,阐明了该断面诸参数四季变化的主控因素,并以此反映和指示了与其相关海域的生态环境特征.结果表明:黄海冷水团及黄海暖流具有季节性演替变化规律;秋季跃层以下水体是一部分夏季冷水团残留水和黄海暖流水的混合水;断面西侧在四季均存在小范围的低盐冷水现象(即青岛冷水团),...     

自然环境下虾夷扇贝(Patinopecten yessoensis)体内麻痹性贝毒的净化研究

为探究在自然环境中虾夷扇贝Patinopecten yessoensis体内麻痹性贝毒(PSTs)的净化过程,从黄海北部大窑湾海域采集了受PSTs污染的虾夷扇贝,将其转移至黄海的棋盘磨水域,该水域未曾有PSTs污染的报道。本研究采用高效液相色谱-荧光检测法(HPLC-FLD)检测30天净化过程中虾夷扇贝体内PSTs的组分和水平。研究结果发现,在第9天,虾夷扇贝软体中的PSTs的含量下降到一个相对较低的水平。而且,初期扇贝体内PSTs的净化率高于后期。扇贝各组织器官中PSTs含量分析发现,消化腺中的PSTs含量最高,这对于人类健康和扇贝养殖具有重要的意义。在本研究中还发现,虾夷扇贝的死亡率与PSTs的水平有一定的关系。     

The relationship between tidal current field and sediment transport in the Huanghai Sea and the Bohai Sea

-Based on a two-dimensional numerical M2 tidal model of the Huanghai and Bohai Seas, sediment transport of particles with different sizes in the computational area is given. It is concluded that tidal currents play an important role in sediment transport and deposition in the Huanghai and Bohai Seas.     

A new spceies of Nassarius (Gastropoda, Nassariidae) from the northern China seas

A new species, Nassarius (Zeuxis) humeratus n. sp. of Nassariidae was described based on the specimens collected in the Huanghai Sea (Yellow Sea) and Bohai Sea during the expedition from 1958 to 1959. The new species was distinguished from similar species Nassarius (Zeuxis) bolangoi (Ladd, 1976) and Nassarius (Varicinassa) variciferus (A. Adams, 1852) in shell characteristics. Some relative discussion was provided. The new species mainly distributes in the subtidal zone of 30-53 m depth, in muddy and sandy bottom in the Bohai Sea and northern Huanghai Sea.     

Nutrient and carbon removal ratios and fluxes in the Ross Sea, Antarctica

Net community production (NCP) and nutrient deficits (Def(X)) were calculated using decreases in dissolved CO₂ and nutrient concentrations due to biological removal in the upper 200 m of the water column during four cruises in the Ross Sea, Antarctica along 76°30′S in 1996 and 1997. A comparison to excess dissolved and particulate organic carbon showed close agreement between surplus total organic carbon (TOC) and NCP during bloom initiation and productivity maximum; however, when TOC values had returned to low wintertime values NCP was still significantly above zero. This seasonal NCP, 3.9±1 mol C m⁻², must be equivalent to the particle export to depths greater than 200 m over the whole productive season. We estimate that the annual export was 55±22% of the seasonal maximum in NCP. The fraction of the seasonal maximum NCP that is exported through 200 m is significantly higher than that measured by moored sediment traps at a depth of 206 m. The removal of carbon, nitrate and phosphate (based on nutrient disappearance since early spring) and their ratios showed significant differences between regions dominated by diatoms and regions dominated by the haptophyte Phaeocystis antarctica. While the ΔC/ΔN removal ratio was similar (7.8±0.2 for diatoms and 7.2±0.1 for P. antarctica), the ΔN/ΔP and ΔC/ΔP removal ratios for diatoms (10.1±0.3 and 80.5±2.3) were significantly smaller than those of P. antarctica (18.6±0.4 and 134.0±4.7). The similarity in ΔC/ΔN removal ratios of the two assemblages suggests that preferential uptake of phosphate by diatoms caused the dramatic differences in ΔC/ΔP and ΔN/ΔP removal ratios. In contrast to low ΔC/ΔP and ΔN/ΔP removal ratio in diatom-dominated areas early in the growing season, deficit N/P and C/P ratios in late autumn indicate that the elemental stochiometry of exported organic matter did not deviate significantly from traditional Redfield ratios. Changes in biologically utilized nutrient and carbon ratios over the course of the growing season indicated either a substantial remineralization of phosphate or a decrease in phosphate removal relative to carbon and total inorganic nitrogen over the bloom period. The species dependence in C/P ratios, and the relative constancy in the C/N ratios, makes N a better proxy of biological utilization of CO₂.     

南黄海和东海毛颚类生态特征的研究Ⅰ.数量分布的特征

根据国家海洋局1977年12月份至1978年11月份在黄海南部和东海(28°~34°N,120°30'~127°00'E)广大海域进行周年逐月份的12航次综合调查资料,对该海域毛颚类数量变动、时空分布及其与环境因子的关系进行了分析.结果表明,本调查区毛颚类的数量以6~9月份较高,7月份为年最高值.1~4月份为年低值期,尤以4月份为最低谷.其中在数量上取主导作用的种类是肥胖箭虫(Sagitta enflata)、强壮箭虫(S.crassa)拿卡箭虫(S.nagae)和百陶箭虫(S.bedoti)等种类.在平面分布上,长江口以北毛颚类数量一般高于长江口以南的海域,在南黄海,除夏季外,其他季节均有出现高值.在东海高密集区一般出现在不同水系交汇区,尤其在长江冲淡水、沿岸水和外海高温高盐水交汇的锋面最为密集.这对于不同水系在研究区内运动和消长有一定的指示意义.     

Study on fishery biology features of spawning group of Pseudopleuronectes yokamae(Günther) in the Sheath Bay

Pseudopleuronectes yokamae (Günther) is one of the most important economic fish species in the genus living specially in the northwest Pacific, and is distributed in the southern area of the far east sea of Russia, Japan, Korea, the Huanghai Sea, the Bohai Sea and the northern area of the East China Sea. Usually they live in nearshore waters of these areas as geographic subpopulations. P. yokamae in the Sheath Bay belongs to a local group of the Huanghai-Bohai Sea subpopulation and is distributed in the mouth and adjacent waters of the bay all the year round, and the catch of its spawning group has important economic value in the area. In this paper the fishery biology features of spawning group of P. yokamae in the Sheath Bay is systematically studied to give biology evidence for proper utilization and the multiplification of its resources.