Pelagic food web interactions and hypolimnetic oxygen depletion: Results from experimental enclosures and lakes

Large lakes enclosures were used to examine the influence of nutrient (P, N) enrichment and planktivorous fish (1 + yellow perch) predation on hypolimnetic oxygen depletion. Results were compared to similar data for lakes with high (Lake St. George) and low (Haynes Lake) abundances of planktivorous fish. In both the unfertilized and fertilized enclosures, fish predation on large cladocerans increased the biomasses of pico- and nanoplankton (0.2–20 µm), phytoplankton (chlorophyll a) and total phosphorus (TP), reduced sedimentation, water clarity, and hypolimnetic oxygen concentrations (AHO). Fertilized enclosures without fish had highest TP and sedimentation rates, but the AHO were low. The high planktivore lake had higher pico- and nanoplankton, higher chlorophyll a, reduced water clarity, and lower AHO than the low planktivore lake. Areal hypolimnetic oxygen depletion (AHOD) rates were strongly related with Secchi depth and plankton size-distribution (r ² = 0.77, and 0.79, respectively), but not as strongly with TP, chlorophyll a, and sedimentation rates (r ² = 0.25, 0.53, and 0.02, respectively). Such observations are useful in forming a generalized hypothesis that lakes with low planktivory and high water clarity have lower oxygen depletion because 1) plankton that are settling are larger and spend less time in the hypolimnetic water column before reaching the sediment, and therefore undergo less decomposition, and 2) the euphotic depth extends into the hypolimnion and production of oxygen can take place.     

Community structure in boreal lakes with recurring blooms of the nuisance flagellate Gonyostomum semen

Blooms of the nontoxic raphidophyte Gonyostomum semen have shown a recent increase in frequency and distribution in the Fennoscandian region. Due to large cell size and several grazer-avoidance strategies, G. semen is hypothesized to be inedible for most zooplankton species and therefore may constitute a bottleneck for the transfer of energy and nutrients in pelagic food webs. Repression of other phytoplankton through increased competition and induced mortality could further exacerbate this effect. In a field study of four lakes with recurring blooms of G. semen and four lakes without blooms, we found significant differences in community structure between the two lake groups during the bloom period. Bloom-lakes had lower biovolumes of small chrysophytes and chlorophytes and zooplankton assemblages were predominated by small, potentially bacterivorous cladocerans, suggesting a limited availability of edible phytoplankton and an increased importance of microbial pathways during G. semen blooms. Low biovolumes of large cladocerans in bloom-lakes may be due to interference of G. semen with filter feeding. Moreover, high abundances of the phantom midge Chaoborus flavicans in bloom-lakes suggest that the flow of energy and nutrients is directed more towards this invertebrate predator than fish. This could have negative impacts on fish populations, especially if bloom periods are prolonged.     

The effects of nutrients and zooplankton community structure on phytoplankton growth in a subtropical Australian reservoir: An enclosure study

Patterns of zooplankton–phytoplankton interactions in subtropical lakes of the Southern Hemisphere may deviate from those established for north-temperate lakes. We tested the responses of phytoplankton growth to different community structures of zooplankton and nutrient enrichment in a subtropical Australian reservoir for the prediction of potential outcomes of lake biomanipulation. Two zooplankton communities were created in lake enclosures over 4 weeks: a rotifer-dominated community developed in the presence of planktivorous fish (Hypseleotris spp.) and a Ceriodaphnia-dominated community developed in the absence of fish. Biomass gradients of both communities were established in 20 L containers and several separate containers received no additions (controls) or were enriched with nitrogen and/or phosphorus. The growth rate of total phytoplankton significantly increased in response to nutrient enrichment, indicating nutrient limitation. Most phytoplankton taxa were not markedly affected by grazing of either zooplankton community. However, both communities had significant stimulatory effects on the growth of inedible chlorophytes. The ability of zooplankton grazing to negatively affect phytoplankton growth during the summer was counteracted regardless of zooplankton community structure, possibly by nutrients regenerated by zooplankton. We hypothesise that in the subtropical system studied, changes in food web nutrient recycling may be more important for the outcome of biomanipulation than grazing impacts.     

Effects of fish manipulation on the plankton community in small hypertrophic lakes from the Pampa Plain (Argentina)

Trophic cascade hypotheses predict that fish will affect the structure and biomass of pelagic plankton communities. In order to investigate trophic cascade effects from fish down to phytoplankton, whole-lake studies were performed in five hypertrophic (mean total phosphorus (TP) concentrations higher than 1000 mg m⁻³) shallow lakes located in the Pampa Plain. The main climatic characteristic of this region is the alternation between periods of drought and flood, with corresponding changes of lake depth and conductivity of lake water. All lakes were studied from April to December 2000. Samples were taken of their physical and chemical characteristics and biotic communities, focusing on the zooplankton community. Fish were manipulated in four lakes (Capurro, Longinotti, Vedia 1, Vedia 2), while the fifth (Lake Vedia 3) was left undisturbed as a reference system. High abundance of planktivorous minnows (Jenynsia multidentata and Cheirodon interruptus) dominated the fish community in the reference lake. In the manipulated lakes, fish stocks were largely reduced in late autumn (May). During winter, Capurro, Longinotti and Vedia 1 were stocked with a visual planktivore, the pampean silverside (Odontesthes bonariensis, Atherinidae). Fish stocking was 24, 33 and 19 kg ha⁻¹, respectively. In contrast, no fish were stocked in Lake Vedia 2. Following fish removal, large Daphnia appeared in these lakes and grazed intensively on the phytoplankton. In contrast, no Daphnia were found in the reference lake (Vedia 3). The stocking of O. bonariensis in lakes Capurro, Longinotti and Vedia 1 led to a decrease in the percentage of large cladocerans and a rise in the phytoplankton biomass:TP ratio. Moreover, the lakes mentioned were stocked with different quantities of silversides over different periods of time. These differences were reflected temporarily in the plankton dynamics, affecting mainly larger sized zooplankton. Nevertheless, the presence of Daphnia was short lived in the lake where fish had been removed and no O. bonariensis were stocked. Competition for resources and recruitment of remaining fish probably caused a collapse in the zooplankton biomass. Our results support the idea that fish predation on zooplankton and its effect on phytoplankton is very intense in small pampean lakes. Fish removal was short lived, however. This could be because in small pampean lakes fish recolonization is favored, and minnows are highly prolific. Moreover, if manipulation of the trophic structure of lakes is undertaken in the pampean region, high nutrient loading from the watershed, climate and hydrology should also be taken into account.     

Zooplankton community structure and ecology of the tropical-highland Lake Hayq, Ethiopia

Lake Hayq, a highland lake in Ethiopia, was stocked with Tilapia fish (Oreochromis niloticus) in late 1970s, offering an opportunity to study the effect of fish predation in a natural lake. Since 1930s, some limnological surveys have been done sporadically documenting a change in zooplankton composition including the disappearance of cladocerans, hypothesizing the stocked planktivorous fish could be a cause. Nevertheless, no detailed research was conducted to identify potential effects of fish stocking predominantly due to its remote location. The article presents data about zooplankton composition, abundance and biomass done between October 2007 and January 2009 on short-time intervals including the underlying limnological variables. The zooplankton community was depauperate comprising two copepods, three cladocerans, and six rotifers taxa, as typical for tropical lakes. Total mean standing biomass of all crustacean zooplankton was 237 mg dry mass m⁻³, which gave Lake Hayq an intermediate position when compared with other tropical lakes. Of copepods, Thermocyclops ethiopiensis was almost an exclusive species, and its temporal variation was influenced by food supply and water temperature. We refute the hypothesis that Tilapia was the cause for the seasonal disappearance of cladocerans, and attribute it to the adverse effect of episodic mixing. Nevertheless, the planktivorous fish probably plays a key role in structuring the cladocerans in particular the large-sized Daphnia magna. In January 2008, we observed a massive planktivorous fish mortality that triggered high algal biomass, which was later grazed by large-sized D. magna demonstrating the trophic cascade hypothesis in a natural ecosystem.     

Limnology of the neotropical high elevation shallow lake Yahuarcocha (Ecuador) and challenges for managing eutrophication using biomanipulation

Temperate and tropical shallow lakes differ in several fundamental aspects with respect to management of eutrophication. High altitude tropical shallow lakes are a special case, showing similarities with temperate and tropical lakes. We studied the ecology of the eutrophic high-altitude tropical lake Yahuarcocha in the Ecuadorian Andes and evaluated the potential of biomanipulation to control eutrophication. With a toxin-producing Cylindrospermopsis bloom, low Secchi depth and low submerged macrophyte cover, Yahuarcocha is clearly in a turbid ecosystem state. Relatively low nutrient concentrations should theoretically allow for a shift to a clear water state through biomanipulation. Top-down control of phytoplankton by zooplankton, however, is complicated by the (1) absence of predatory fish, (2) fish community dominated by small poecelid species, (3) lack of a refuge for zooplankton from fish predation within the macrophytes, and (4) persistent, grazing resistant bloom of the cyanobacterium Cylindrospermopsis. In these aspects, lake Yahuarcocha is more similar to tropical shallow lakes, probably because water temperature is high relative to the mean air temperature and because of the absence of a cold season. The fish and macrophyte communities consisted almost entirely of exotic species. The exotic fish species probably stabilized the turbid state in the lake.     

Size- and depth-dependent variation in habitat and diet of the common carp (Cyprinus carpio)

The habitat and diet variation of the common carp (Cyprinus carpio) were studied in Lake Banyoles (Catalonia, Spain). Carp was the second most abundant species offshore and used more the littoral in spring and deep bottoms in winter. The diet of carp was based on detritus, amphipods (Echinogammarus sp.), phantom midge larvae (Chaoborus flavicans), diatom mucilages, and plant debris. Amphipods and phantom midge larvae were much more important in diet than previous studies found, because of their greater availability in this lake. Among the carp inhabiting deep waters, there was size-dependent variation in diet, with smaller carp selecting more meiobenthos (cladocerans, ostracods, and small chironomids) and larger carp preying on profundal macrobenthos (phantom midge larvae and large chironomids). Roach (Rutilus rutilus) and carp dominated in abundance the non-littoral zone of the lake and showed resource partitioning, with roach being a more efficient zooplanktivore and carp being more able to prey on hard material (plant seeds, mollusks, and ostracods).     

Basic limnological characteristics of the shallow eutrophic lake Grimnitzsee (Brandenburg, Germany)

The medium shallow lake Grimnitzsee (maximum depth: 9.9 m; mean depth: 4.6 m; area: 7.7 · 10⁶ m²) which is situated in the biosphere reserve “Schorfheide-Chorin” in northern Brandenburg (Germany) was studied in 1994 and 1995. A bathymetric map of Grimnitzsee is given for the first time. The lake is usually polymictic although in 1994 and 1995 relatively long summer stratification was observed due to very high global radiation input. Nutrient concentration, light climate, oxygen status, phytoplankton biomass and the species composition of littoral diatoms characterize the lake as eutrophic. Special features deducible from the lake's polymictic character were the multiple development of aerobic or anaerobic strata above the sediment, the fast recovery of silicon concentration in the water column after diatom sedimentation, the importance of resuspension for the success of planktonic diatom populations, and an only moderate correlation between chlorophyll a concentration and light attenuation as well as seston dry weight probably due to the influence of suspended particles.     

Subfossil and modern diatom plankton and the paleolimnology of Rotsee (Switzerland) since 1850

The diatom biostratigraphy of the topmost sediment meter of Rotsee, Central Switzerland, is characterized by a major change fromCyclotella comensis-dominated toStephanodiscus hantzschii/S. parvus-dominated assemblages. A comparison between old phytoplankton samples, taken between 1910–1930, and subfossil diatom assemblages is used for dating the upper 35 cm of the core. There is evidence that the change in dominant diatoms occurred in 1919/20, thus before the opening of an artificial inlet in 1922, and is due to increasing eutrophication. Furthermore, the sedimentary carbonate content can be used as a good indicator for past phytoplankton productivity in Rotsee.     

Modeling Energy Fluxes in Lake Ecosystems as a Means of Hydrobiological Studies

A mass-balance model was developed for the prediction of the biomass and annual production of phytoplankton, macrophytes, epiphytes, and phytobenthos (primary producers), bacterioplankton and bacteriobenthos (reducers), herbivorous and predator zooplankton, zoobenthos, planktivorous fish, benthivorous fish, and predatory fish (consumers). The model is controlled by a small number of parameters.__________Translated from Vodnye Resursy, Vol. 32, No. 3, 2005, pp. 361–370.Original Russian Text Copyright © 2005 by Boulion.     

以鲢、鳙养殖为主的长江中下游武山湖浮游植物群落结构特征

武山湖是紧邻长江的通江型富营养化湖泊,是国家级湿地公园.为切实了解该湖在以鲢、鳙养殖为主的情况下浮游植物结构特征,于2017年9月-2018年8月对其浮游植物群落结构特征及水质开展了监测.监测结果表明武山湖水质全年处于轻度富营养到重度富营养水平之间;12次共采集浮游植物7门100种（属）,浮游植物优势种共有23种,其中蓝藻门有9种,绿藻门有8种,硅藻门和隐藻门各有3种.夏季和秋季蓝藻门优势种最多且优势度高,冬季和春季绿藻门和硅藻门优势种多且优势度高.武山湖浮游植物每月优势度最大的种类主要有蓝藻门微囊藻和细小平裂藻、绿藻门小球藻以及硅藻门小环藻.浮游植物生物量峰值出现在6月,达34.77 mg/L;丰度峰值出现在7月,达341.46×10⁶ cells/L.冗余分析（RDA）和线性相关分析均表明浮游植物丰度和生物量与总磷、温度和pH均呈显著正相关,且蓝藻门生物量和丰度以及优势属与总磷和温度均呈显著正相关.研究结果表明武山湖浮游植物丰度和生物量在夏季均很高,发生蓝藻水华的风险较大.相对于氮,磷是更重要的限制浮游植物生长的营养元素.     

Harmful algal blooms (HABs) in Daya Bay, China: An in situ study of primary production and environmental impacts

A month-long investigation of phytoplankton biomass and primary production (PP) was carried out during a harmful algal bloom (HAB) in Daya Bay, China, in 2003. During the bloom, the phytoplankton community was dominated by Scrippsiella trochoidea and Chattonella marina. The phytoplankton biomass (Chl a) and PP reached peak levels of 519.21 mg m⁻³ and 734.0 mgC m⁻³ h⁻¹, respectively. Micro-phytoplankton was the key contributor to Chl a and PP in a cage-culture area and in the adjacent HAB-affected waters, with percentages of up to 82.91% and 84.94%, respectively. The HAB had complicated relationships with hydrological and meteorological factors in Daya Bay. However, the water around the cage-culture area always showed statistically greater phytoplankton biomass and nutrient loadings than in adjacent waters, suggesting that this was the “trigger area” of the bloom. The spatial and temporal distribution of diverse HABs in Daya Bay, their ecological characteristics, and their environmental impacts are also discussed in this paper.     

Crustacean zooplankton distribution patterns and their biomass as related to trophic indicators of 29 shallow subtropical lakes

A comparative limnological study was carried out to present a snapshot of crustacean zooplankton communities and their relations to environmental factors to test whether there is a consistent relationship between crustacean biomass and trophic indicators among lake groups with similar trophic conditions. The study lakes showed a wide range of trophic status, with total phosphorus (TP) ranging from 0.008 to 1.448 mg L⁻¹, and chlorophyll a from 0.7 to 146.1 μg L⁻¹, respectively. About 38 species of Crustacea were found, of which Cladocera were represented by 25 taxa (20 genera), and Copepoda by 13 taxa (11 genera). The most common and dominant species were Bosmina coregoni, Moina micrura, Diaphanosoma brachyurum, Cyclops vicinus, Thermocyclops taihokuensis, Mesocyclops notius and Sinocalanus dorrii. Daphnia was rare in abundance. Canonical correspondence analysis showed that except for four species (D. hyalina, S. dorrii, C. vicinus and M. micrura), almost all the dominant species had the same preference for environmental factors. Temperature, predatory cyclopoids and planktivorous fishes seem to be the key factors determining species distribution. TP was a relatively better trophic indicator than chlorophyll a to predict crustacean biomass. Within the three groups of lakes, however, there was no consistent relationship between crustacean biomass and trophic indicators. The possible reason might be that top-down and bottom-up control on crustaceans vary with lake trophic state. The lack of significant negative correlation between crustacean biomass and chlorophyll a suggests that there was little control of phytoplankton biomass by macrozooplankton in these shallow subtropical lakes.     

Pelagic trophic transfer efficiency in an oligotrophic, dimictic deep lake (Lake Stechlin, Germany) and its relation to fisheries yield

We investigated trophic transfer efficiency in the pelagic food chain of deep, oligotrophic Lake Stechlin (Germany) by analyses of the primary, secondary, and fish production. Primary production between April and November 2000 was estimated at 78 g C m⁻², pelagic secondary production at 14 g C m⁻², and production of the main planktivorous fish species [European cisco, Coregonus albula (L.)] at 0.77 g C m⁻². Thus, trophic transfer efficiency between primary and pelagic secondary production was around 18%, whereas between pelagic and fish production around 6%. The high efficiency at the first step of the chain is discussed to be due to the high food quality in oligotrophic lakes due to the dominance of Bacillariophyceae and Chlorophyceae rich in essential fatty acids. In turn, the relatively low trophic transfer efficiency between the secondary and the fish production is mainly explained by the avoidance of calanoid copepods as food source by the ciscoes. Concerning the trophic transfer efficiency, results from this study support the general assumption of a 10% transfer between neighbouring trophic levels within ecosystems.     

Geographic variations in shell growth rates of the mussel Diplodon chilensis from temperate lakes of Chile: Implications for biodiversity conservation

The Chilean lake district includes diverse lentic ecosystems along ca. 700 km of the country (36°–43°S), including the “Nahuelbutan lakes”, “Araucanian lakes” and “Chiloe lakes”. This area is recognized as an important “hot spot” of benthic freshwater biodiversity in Southern South America. In Chilean temperate lakes, increased nutrient loads of P and N caused eutrophication, particularly in the Nahuelbutan Lakes. The freshwater Hyriidae mussel Diplodon chilensis (Gray, 1828) which is one of the most abundant species in Chilean temperate lakes, is known to be very susceptible to eutrophication. This species presents a clear reduction in its geographic ranges and is considered to be a threatened species in many Chilean lakes. In this study, we used a correlative approach to determine how eutrophication-driven changes in the food supply and in geographical parameters of different Chilean lakes affected the shell growth rates of D. chilensis. The results obtained from sclerochronological analyses of the mussel shells suggest an association with a group of environmental variables, including geographical types (negative), such as latitude and altitude, and limnological types (positive), especially phosphorous and turbidity. However, the D. chilensis populations under extreme conditions of turbidity in eutrophic and hypertrophic lakes are extinct or nearly so. The high positive correlation of the mean D. chilensis growth rates with orthophosphate (R=0.76; P<0.05), in relation to dissolved inorganic nitrogen, suggests that P is the major limiting factor of the primary productivity in Chilean temperate lakes. We discuss some implications of our results in terms of the conservation of biodiversity in temperate lake ecosystems at different taxonomic levels.     

Public participation in monitoring programmes as a tool for lakeshore monitoring: the example of Lake Pyhäjärvi, Karelia, Eastern Finland

Lake Pyhäjärvi, on the border between Finland and Russia in Karelia, is a very valuable clear-water lake of the Lobelia type. It belongs to the European Union's Natura 2000 programme in Finland, and has been included in regional and national monitoring programmes since the 1960s. The main monitoring station is situated near the outlet of the lake. Deterioration of its water quality was suspected already in the 1980s because of decreasing Secchi depths (transparency) and increasing chlorophyll a.The occurrence of algal blooms on the lakeshores is monitored weekly during each summer at one site on Lake Pyhäjärvi (site 1). This is a part of nationwide intensive algae monitoring programme organised by the environmental authorities together with voluntary observers at some 270 lake sites in Finland since 1998. Since 1997, Secchi depth observations have been carried out by volunteers biweekly or monthly at 17 sites on the lake. In the vicinity of one of these transparency observation sites (station 100), intensive monitoring of algae has been carried out. At this lakeshore monitoring site 69 algal observations were made, ten of which recorded algal blooms during the study period 1998–2002. The observed algal blooms were caused by algae of the Anabaena species, mainly by Anabaena lemmermannii. At Lake Pyhäjärvi the number of algal bloom observations received from the public have decreased from the 1990s to the 2000s. The range of Secchi disc transparency was 5.0–8.4 m with a mean value of 6.2 m at station 100 and 4.3–7.7 m (mean 6.1 m) at the main monitoring station 2 during the open water periods in 1998–2002. During this study period, the maximum values at site 100 seem to have increased slightly, which might indicate some improvement in the water quality due to decreased point source loading.We conclude that the intensive algal monitoring results of 5 years at the lakeshore site and the transparency results — both compiled by trained volunteers — reflect an improvement in the state of Lake Pyhäjärvi in Karelia. This conclusion is in accordance with the long-term water quality and short-core studies of sedimentary diatoms in Lake Pyhäjärvi. We suggest that the intensive algal observations and transparency measurements are both suitable methods for the monitoring of lakeshores and lakes, and that both are suitable for voluntary monitoring. We found public participation a good tool for monitoring lakes and lakeshores.     

云南云龙天池枝角类群落演替对1950s以来人类活动影响信号的识别

从1950s开始,云南地区部分湖泊受到了水文调控(如筑坝)和鱼类引入等流域开发活动的直接影响,湖泊水环境与生态系统结构已出现明显改变.开展浮游动物群落的长期生态响应评价有助于认识气候波动和人为胁迫的影响模式.本文选取云龙天池进行了沉积物记录分析,在重建过去100年环境变化历史的基础上开展了枝角类群落的多指标分析(物种组成、个体大小、生物量等),进一步结合多变量分析识别了枝角类群落构建的关键驱动因子.结果显示,云龙天池在过去100年间经历了明显的水位波动,约1962年以前水位较低,1950s起的水文调控(筑坝)导致湖泊水位波动上升,2006年以来略有下降.枝角类群落随水位波动呈现由底栖种向浮游种占优转变的模式,并在2006年以来底栖种略有增加.总体上,低水位时期底栖枝角类占优,高水位时期浮游枝角类占优.水体营养水平也对枝角类群落产生了较为显著的影响.在沉积物总氮和有机质通量上升时,长额象鼻溞(Bosmina longirostris)相对丰度和枝角类浓度都明显上升.枝角类象鼻溞个体大小表明,1969年鱼类引入后象鼻溞的壳长、壳刺长度、触角长度显著减小,反映了鱼类捕食压力上升的影响.本研究...     

富营养化初期湖泊放养鲢(Hypophthalmichthys molitrix)、鳙(Aristichthys nobilis)控藻生态效果的初步评估

放养鲢（Hypophthalmichthys molitrix）、鳙（Aristichthys mobilis）控制富营养化湖泊蓝藻水华暴发是当前最有效的手段之一.为评估富营养化初期湖泊鲢、鳙控藻的生态效果,2016年在洱海红山湾开展原位围栏鲢、鳙控藻实验,对围栏内外的浮游生物群落结构及水环境因子的变化进行对比分析,同时测定了围栏内鲢、鳙的生长及摄食节律.结果表明：围栏内鲢、鳙呈现匀速生长,围栏生存环境适宜.根据Egger''s模型估算鲢、鳙的日摄食量分别为24.96%和18.18%,鲢的日粮高于鳙;滤食率结果表明,鲢对浮游植物的摄食率高于鳙（分别为3.01和2.19 L/（g·h）,而鳙对浮游动物的滤食率高于鲢（分别为18.61和13.54 L/（g·h））.研究期间,围栏内外水体理化因子无显著差异;但围栏内外微囊藻生物量差异明显,并且在7月达到生物量峰值（围栏内为1.7 mg/L,围栏外为2.4 mg/L）.鲢、鳙对浮游植物的优势种类——微囊藻和隐藻有明显的削减效果,6-8月逐渐上升并在8月达到最高（64%）;当浮游植物优势种由微囊藻替换为硅藻门和绿藻门的种类时,鲢、鳙的削减效率不明显（10-11月为负值）.此外,鲢、鳙放养显著降低了浮游甲壳动物的总生物量和枝角类生物量.因此,结合实验结果得出,鲢具有更高的控藻能力,尤其对微囊藻水华,但同时具备了较高对浮游甲壳动物的削减能力,需要进一步分析富营养化初期湖泊——洱海中实施非经典生物操纵的适宜性.     

九寨沟国家级自然保护区长海夏季浮游植物群落结构及生态评价

高山湖泊对于全球气候变化及人类影响是一个极为敏感的参照系统.九寨沟国家级自然保护区长海作为一个独特的高山湖泊,研究其浮游植物群落结构及其与环境的关系,评估其水质现状及影响因素,有着重要的意义.本文于2014年7月对长海浮游植物群落结构进行了研究.全湖共布设12个采样点,并在中心采样点进行了垂直分层采样.本次调查共发现浮游植物6门38属63种,平均丰度为6.98×10~5cells/L,平均生物量为0.31 mg/L.浮游植物的水平分布差异不大;在垂直分布上,浮游植物的丰度从表层0.5 m至水下50 m呈现先增加后减少的趋势,在20 m水深处达到最大.长海浮游植物的优势种是长海小环藻(Cyclotella changhai)和飞燕角甲藻(Ceratium hirundinella),长海小环藻数量较多,飞燕角甲藻生物量较大.长海浮游植物多样性指数较低,综合各类水质评价方法,可以得出九寨沟长海处于贫-中营养状态.     

A laboratory investigation into the effects of slope on lava flow morphology

In an attempt to model the effect of slope on the dynamics of lava flow emplacement, four distinct morphologies were repeatedly produced in a series of laboratory simulations where polyethylene glycol (PEG) was extruded at a constant rate beneath cold sucrose solution onto a uniform slope which could be varied from 1° through 60°. The lowest extrusion rates and slopes, and highest cooling rates, produced flows that rapidly crusted over and advanced through bulbous toes, or pillows (similar to subaerial “toey” pahoehoe flows and to submarine pillowed flows). As extrusion rate and slope increased, and cooling rate decreased, pillowed flows gave way to rifted flows (linear zones of liquid wax separated by plates of solid crust, similar to what is observed on the surface of convecting lava lakes), then to folded flows with surface crusts buckled transversely to the flow direction, and, at the highest extrusion rates and slopes, and lowest cooling rates, to leveed flows, which solidified only at their margins. A dimensionless parameter, Ψ, primarily controlled by effusion rate, cooling rate and flow viscosity, quantifies these flow types. Increasing the underlying slope up to 30° allows the liquid wax to advance further before solidifying, with an effect similar to that of increasing the effusion rate. For example, conditions that produce rifted flows on a 10° slope result in folded flows on a 30° slope. For underlying slopes of 40°, however, this trend reverses, slightly owing to increased gravitational forces relative to the strength of the solid wax. Because of its significant influence on heat advection and the disruption of a solid crust, slope must be incorporated into any quantitative attempt to correlate eruption parameters and lava flow morphologies. These experiments and subsequent scaling incorporate key physical parameters of both an extrusion and its environment, allowing their results to be used to interpret lava flow morphologies on land, on the sea floor, and on other planets.