New records of Jurassic petrified wood in Jianchang of western Liaoning,China and their palaeoclimate implications

Diverse and rich fossil wood records have been documented from the Mesozoic of Liaoning Province, NE China. The Tiaojishan Formation(mainly distributed in Beipiao, Chaoyang, Fuxin and Jianchang regions) is one of the most significant horizons for the Jurassic petrified wood in Liaoning Province. Previously, wood fossils of this formation were mainly reported in Beipiao and Chaoyang regions, whereas fossil wood record was merely known in Jianchang region. Here we describe new fossil wood specimens from the Tiaojishan Formation in Jianchang County, western Liaoning. Two fossil wood taxa, i.e., Protaxodioxylon jianchangense Tian et Wang sp. nov. and Xenoxylon peidense Zheng et Zhang were recognized on the basis of anatomical features. These fossil wood records add new data for understanding the fossil wood diversity, floral composition and palaeoclimate of the Tiaojishan Formation. The occurrence of Xenoxylon and Protaxodioxylon implies a cool temperate, wet and seasonal climate condition with interannual variations during the Middle to Late Jurassic transition in western Liaoning region.     

The oldest angiosperm—a tricarpous female reproductive fossil from western Liaoning Province, NE China

The fossil plant female reproductive organs each with a tricarpous were collected from the Yixian Formation, Chaoyang district, western Liaoning Province. The geological age of the Yixian Formation here is believed to be Late Jurassic, which places this fossil as the oldest angiosperm currently known. Its morphological characters are distinct from other known angiosperm reproductive organs as it has 3 ovaries, each with a terminal style and an apical stigma. 9 such organs compose a cyme. As this morphological arrangement is unique to this plant, a new genus and species has been established, namedChaoyangia liangii gen. et sp. nov. Project supported by the Special Finances of Biological Sciences and Technology, Chinese Academy of Sciences (the special item of the Ministry of Finance of the People’s Republic of China).     

Pseudofrenelopsis fossils from Cretaceous gypsum beds in Guixi,Jiangxi Province,China and their geological significance

Compressed fossil plants were found in the gypsum beds of the Zhoujiadian Formation in Guixi City, Jiangxi Province, China. The cuticular features of two species of Pseudofrenelopsis (Cheirolepidiaceae) were examined by scanning electron microscopy in this study: Pseudofrenelopsis papillosa Cao ex Zhou and Pseudofrenelopsis guixiensis Bainian Sun et Dai, sp. nov. A detailed analysis of the internode epidermis, and the adaxial and abaxial leaf cuticles of the two species, was performed, extending our knowledge of the morphological and anatomical characteristics of the extinct Pseudofrenelopsis. A new species, Pseudofrenelopsis guixiensis sp. nov., is proposed based upon comparison of its cuticular features. It differs from previously published species in that it has hairs on the abaxial surfaces of its leaves, sometimes shares subsidiary cells on the surfaces of internodes, lacks a hypodermis, and has no papillae on its epidermal cells. Because the distribution of fossil Pseudofrenelopsis is restricted to the Lower Cretaceous, the Early Cretaceous age of the Zhoujiadian Formation is affirmed. The fossils of Pseudofrenelopsis show obvious xeromorphic characteristics, such as apparent thickening of the cuticles, subsidiary cells with obvious papillae, and anticlinal walls with strong cutinization, and they are found associated with gypsum beds and close to red beds. All of these features indicate an arid climate.     

Klukiopsis jurassica—A new Jurassic schizaeaceous fern from China

A new Jurassic schizaeaceous fernKlukiopsis jurassica gen. et sp. nov. from Yima, Henan Province, China is described. The new fern is characterized by the abaxial sori arranged in two rows, apical and complete annulus and more than 800 smooth trilete spores in each sorus.     

Klukiopsis jurassica--A new Jurassic schizaeaceous fern from China

A new Jurassic schizaeaceous fern Klukiopsis jurassica gen. et sp. nov. from Yima, Henan Province, China is described. The new fern is characterized by the abaxial sori arranged in two rows, apical and complete annulus and more than 800 smooth trilete spores in each sorus.     

Constraints on the age of the Tuchengzi Formation by LA-ICP-MS dating in northern Hebei-western Liaoning,China

Accurately determining the age of the Tuchengzi Formation has direct influence on confirming the boundary between the Jurassic and the Cretaceous systems in northern Hebei-western Liaoning, and on related geological problems in China. However, the Tuchengzi Formation mainly consists of sedimentary rocks, with a poor fossil record and especially lack of index fossils. The Tuchengzi Formation is also lack of the type of volcanic rocks that can provide an isotopic age. Therefore, the age of the Tuchengzi Formation has been uncertain. Based on our systematic dating of the tuff interbedded in the Tuchengzi Formation of Chengde and Jinlingsi-Yangshan basins in northern Hebei-western Liaoning, combined with the dating results of previous researchers, here we suggest that the age range of the Tuchengzi Formation in northern Hebei-western Liaoning is from 147 Ma to 136 Ma. It implied that the Tuchengzi Formation was mainly formed in the Early Cretaceous. Supported by National Natural Science Foundation of China (Grant No. 90714010)     

New fossil eccoptarthrids (Coleoptera: Curculionoidea) from the Yixian Formation of western Liaoning, China

One new genus and three new species of the weevil family Eccoptarthridae (Curculionoidea), Leptocar polychaetus gen. et sp. nov., Abrocar macilentus sp. nov., Cretonanophyes punctatus sp. nov., are described and illustrated. They pertain to the Upper Jurassic or Lower Cretaceous Yixian Formation of western Liaoning Province, China. The host plants to the living eccoptarthrids and the floras in Yixian Formation indicate that those archaic eccoptarthrids possibly lived on conifers with a phylogenetic closeness to Cupressaceae. Besides, the early diversification of eccoptarthrids is discussed: origi- nated in Central or East Asia in the Late Jurassic, spread into Western Europe and South America during the Early Cretaceous, underwent a dramatic decline after the Early Cretaceous which likely was caused by competitive pressure and the displacement of their host plants.     

New discovery of Palaeontinid fossils from the Middle Jurassic in Daohugou, Inner Mongolia (Homoptera, Palaeontinidae)

Three new species of fossil Palaeontinidae are described from Daohugou Village, Inner Mongolia, China: Daohugoucossus shii sp. nov., D. parallelivenius sp. nov., D. lii sp. nov. The diagnosis of Dao- hugoucossus Wang, Zhang and Fang, 2006 is revised. These materials are the first complete fossil palaeontinids in the Middle Jurassic of the world. Based on observation of these new specimens, Sc of this new genus has the following combined characters: joined with R at base, diverged near M separa- tion, coalesced with R before the nodal line, extended beyond the nodal line and fused with R1 before distal end. Due to the Sc characteristics mentioned above, the genus Liaocossus Ren, Yin and Dou, 1998 can be well distinguished from “Ilerdocossus-complex”.     

Phylogenetic relationships of the Chinese fossil species of the genusEquus (Perissodactyla, Equidae)

The Chinese fossils ofEquus began to appear from the beginning of the Quaternary at 2.5 Ma B.P., and the extant species ofEquus still live in China under natural conditions at the present, which is unique in the world. 12 valid fossil species ofEquus have been discovered in China, including the stenonid, caballoid and hemione representatives ofEquus. The origin and evolution of the Chinese fossil species and other relevant species ofEquus are discussed in detail. The pmposed phylogenetic relationships and temporal distribution sequences including all the Chinese fossil species ofEquus are established. Some mistakes in the earlier researches about the Chinese fossils ofEquus are corrected. Project supported by the French-Chinese Cooperation (Grant No. 203720C).     

Two new species of Vestrogothia (Phosphatocopina, Crustacea) of Orsten-type preservation from the Upper Cambrian in western Hunan, South China

Although fossils of Orsten-type preservation represented by Skaracarida and Phosphatocopina were first reported from the Middle and Upper Cambrian in western Hunan, South China in 2005, diversified phosphatocopine species have never been appropriately described and elucidated in terms of their evolutionary relationships. Here, we described two new species of Phosphatocopina, Vestrogothia anterispinata sp. nov. and V. bispinata sp. nov. The evolutionary relationship among all the valid species of Phosphatoco...     

Geological age of the Yokawa Formation of the Kobe Group (Japan) on the basis of terrestrial mammalian fossils

Abstract   The age of the Yokawa Formation of the Cenozoic Kobe Group distributed in Hyogo Prefecture of western Japan based on mammalian fossils is discussed. Two fossil dental specimens of terrestrial mammals discovered from the lowest part of the Yokawa Formation in the Sanda area are described. These two fossils described here are: (i) a right mandibular fragment with p2–m3 of Bothriodon sandaensis sp. nov. (selenodont anthracotheriid artiodactyl), which appears to be the most primitive among the species of the genus; and (ii) right m1–m3 of cf. Hyrachyus sp. (primitive rhinocerotoid perissodactyl). In the lower part of the Yokawa Formation, Zaisanamynodon (amynodontid perissodactyl) was previously reported. The morphology ('evolutionary stage') of B. sandaensis is indicative of the latest Middle to Late Eocene, that of cf. Hyrachyus sp. is indicative of the Early to Middle Eocene, and Zaisanamynodon is indicative of the Late Middle to Late Eocene. Therefore, the fossil mammals of the Yokawa Formation indicate an latest Middle Eocene ( ca . 38 Ma) correlation for the lower part of the formation, as a working hypothesis. Although the resolution of the geological age based on these mammalian fossils is relatively low compared to that based on marine index fossils, this result is concordant with the recent radiometric correlation of the lower part of the Yokawa Formation in the Sanda area.     

Latest Middle Miocene fauna and flora from Kumkol Basin of northern Qinghai-Xizang Plateau and paleoenvironment

We report 7 mammalian and 16 pollen species from the top of the Shimagou Formation in Kumkol Basin of northern Qinghai-Xizang Plateau, whose age is latest Middle Miocene(about 12.5 Ma). We erect a new species of fossil rodent,Spermophilinus kumkolensis sp. nov. The micromammalian fauna from the Kumkol Basin shares high similarities with contemporary faunas from Europe and northern China, indicating the existence of active zoogeographic exchange between Europe and northern Asia. Trees are dominant in pollen fossils, and herbs and shrubs are minorities. Broadleaved Quercus and Fagus predominate in tree pollens. The flora reconstructed by pollens displays a mixture of evergreen broadleaved forest and dry and warm steppe landscape. Compared to the modern vegetation in Yunnan region, the latest Middle Miocene flora in the Kumkol Basin represents a distinct vertical distribution. The ecotypes of the latest Middle Miocene fauna and flora in the Kumkol Basin,as a whole, are markedly different from those of modern animals and vegetation, the past climate being more suitable than today.     

Description de deux nouvelles Cyclotelles (Bacillariophyceae) de milieux alcalins et eutrophes:Cyclotella costei nov. sp. etCyclotella wuethrichiana nov. sp.

Two new species of the genus Cyclotella are presented which have been discovered during ecological and paleoecological research studies in the littoral zone of two small alkaline eutrophic lakes in France (Lake Paladru) and in Switzerland (Lake Le Loclat). These new species are namedCyclotella costei nov. sp. andCyclotella wuethrichiana nov. sp.     

Radiolarian fauna from the Chiungchussuan Shuijingtuo Formation(Cambrian Series 2) in Western Hubei Province,South China

Little is known regarding characteristics and evolution of radiolarian fossils in the early Cambrian due to its rarity and poor preservation. Here we report a new radiolarian fauna from the Cambrian Shuijingtuo Formation corresponding to the Chiungchussuan Stage(Cambrian Series 2) in western Hubei, China. It contains over 300 radiolarian tests and these fossils belong to 4 morphotypes, including Paraantygopora porosa gen.et sp. nov., Braunosphaera sinensis gen.et sp. nov., Tetrasphaera? sp. and incertae spherical radiolarians. In taxonomy, Spumellaria predominates in the fauna and comprises 92% of the total. In shell structure, about 73% of all specimens are similar to the early Ordovician radiolarian fossils, with the shell walls characterized by perforated plate structures. The spherical radiolarians with latticed shells constitute another feature of the early Cambrian radiolarian fauna. The radiolarian fauna reported here contains many fossil specimens and covers different taxonomical taxa. These specimens usually consist of distinctive concentric multi-layers with complex structure. These characteristics indicate that Radiolaria had already thrived during the Cambrian Chiungchussuan Stage(Series 2, Stage 3), which may have occurred synchronously with the explosion and prosperity of macrobiota recorded in the strata deposited in shallower water condition.     

Jurassic Progonocimicidae (Hemiptera) from China and phylogenetic evolution of Coleorrhyncha

The fossil record of Progonocimicidae of Hemiptera suborder Coleorrhyncha from China is reviewed. Ovicimex laiyangensis Hong et Wang, 1990, from the Lower Cretaceous of Laiyang, is excluded from Coleorrhyncha. The available generic name Mesocimex Hong, 1983 is resurrected from synonymy and proposed for replacement of the preoccupied name and junior homonym Mesoscytina Hong, 1983, non Mesoscytina Tillyard, 1919. It resulted in following new combinations: Mesocimex abditus (Yu. Popov, 1982) comb. n., Mesocimex ambiguus (Yu. Popov, 1985) comb. n., Mesocimex brunneus (Hong, 1983), comb. n., Mesocimex fidus (Yu. Popov, 1982) comb. n., Mesocimex intermedius (Yu. Popov, 1985) comb. n., Mesocimex kuzbasicus (Yu. Popov, 1985) comb. n., Mesocimex liliputus (Yu. Popov, 1988) comb. n., Mesocimex minutus (Yu. Popov, 1982) comb. n., Mesocimex modestus (Yu. Popov, 1985) comb. n., Mesocimex paulinus (Yu. Popov, 1982) comb. n., Mesocimex cognatus (Yu. Popov, 1982) comb. n. An annotated list of species of Mesocimex is given. In addition, a new species, Mesocimex lini sp. nov., is described based on well-preserved specimens with wings and bodies from the Middle Jurassic of Daohugou, China. This discovery supports the Middle Jurassic age of Daohugou Lagerstatte, since all species of Mesocimex are confined to the Lower-Middle Jurassic. The phylogenetic evolution of Coleorrhyncha is discussed: Cicadocorinae, Karabasiinae, Hoploridiinae, and Peloridiidae are monophyletic clades whereas Progonocimicinae and Karabasidae are clearly paraphyletic groups.     

Fossil nothofagaceous leaves from the Eocene of western Antarctica and their bearing on the origin, dispersal and systematics of Nothofagus

Fossil leaves resembling Nothofagaceae have been investigated from the Eocene of western Antarctica and a new form genus Nothofagofolia is proposed for these kinds of fossils. Some new specimens belonging to this form genus are described. They were collected from the Fossil Hill locality of Fildes Peninsula, King George Island, South Shetland Islands, western Antarctica. Two new species, two new combinations and an unnamed species are reported. A number of published Nothofagus leaf fossils from the same locality are discussed and revised. As a result of these studies of Nothofagus leaf morphology, we conclude that (1) Nothofagus probably originated in high latitudes of the Southern Hemisphere during the middle-late Late Cretaceous and diversified, dispersed gradually to the lower latitudes of the same hemisphere; (2) leaf morphological characters are significant for the systematics of the family Nothofagaceae, especially at the intrageneric level; and (3) extant species of Nothofagus known from southern temperate areas have more primitive leaf morphological characters and lower leaf ranks than those from tropical mountains as well as those of the Fagaceae and Betulaceae. Supported by the Foundation of State Key Laboratory of Palaeobiology & Stratigraphy, Nanjing Institute of Geology and Palaoentology, Chinese Academy of Sciences (Grant No. 013106), the National Natural Science Foundation of China (Grant No. 30670159) and the Fund of Innovation Program by Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences     

Response to paleoatmospheric CO<Subscript>2</Subscript> concentration of <Emphasis Type="Italic">Solenites vimineus</Emphasis> (Phillips) Harris (Ginkgophyta) from the Middle Jurassic of the Yaojie Basin,Gansu Province,China

Solenites vimineus (Phillips) Harris (Ginkgophyta) specimens with well-preserved cuticles were collected from five different beds in the oil shale member of the Middle Jurassic Yaojie Formation in the Yaojie Basin, Gansu Province, northwestern China. Gross morphology and fine structures of the fossil leaves were studied, and stomatal parameters were analyzed, according to which, the paleoatmospheric CO₂ concentration can be deduced as 1512–1896 ppm that would have caused an increase in mean temperature of about 6.5–7.4°C. Carbon isotopes from the fossils indicate that the carbon isotopic discrimination and water use efficiency (WUE) of S. vimineus were raised nearly obtaining the ideal physiologic state in increased CO₂ concentrations and temperature conditions, suggesting that S. vimineus records the paleoenvironmental information of the Middle Jurassic Yaojie Basin and can be used for environmental reconstruction.     

安徽省重要古生物化石资源及其意义

安徽省地处华北、华南板块的交变地带,沉积单元和生物群落均非常复杂,自晚太古代以来的各时代地层发育较全并含有丰富的无脊椎动物、脊椎动物、植物化石和遗迹化石等。该省的古生物化石资源以分布广泛,类型众多,保存完好,科学意义重大为主要特征。本文重点探讨了安徽省古生物化石资源的分布特征、赋存特征及具有地方特色的古生物化石类型,对该省古生物化石资源的意义进行了初步评价,并对安徽省古生物化石资源的深入研究和合理开发利用的方向进行了述评。     

Fossil leaves of Berhamniphyllum (Rhamnaceae) from Markam,Tibet and their biogeographic implications

A new occurrence of buckthorn fossil leaves is reported from the upper Eocene strata of Markam Basin,southeastern Tibet,China.The leaf margin is either entire or slightly sinuous.Secondary veins are regularly spaced,forming eucamptodromous venation.These secondaries exist as straight lines from midvein to near margin and then arch abruptly upward and enter into a margin vein.The tertiary veins are densely spaced and parallel,and are percurrent to secondary veins.This leaf architecture conforms with Berhamniphyllum Jones and Dilcher,an extinct fossil genus reported from America.Our fossils are characterized by their dense secondaries,with secondary veins on the upper half portion of the blade accounting for over 40%of all secondaries.A new species,Berhamniphyllum junrongiae Z.K.Zhou,T.X.Wang et J.Huang sp.nov.,is proposed.Further analysis shows that confident assignment among Rhamnidium,Berchemia,and Karwinskia cannot be made based on leaf characters alone.Berhamniphyllum might represent an extinct common ancestor of these genera.In this study,several fossil Berchemia from Yunnan and Shandong are emended and reassigned to Berhamniphyllum.A new complex,namely the Berchemia Complex,is proposed based on morphology,molecular evidence,and the fossil record.This complex contains the fossil leaves of Rhamnidium,Karwinskia,Berchemia,and Berhamniphyllum.The historical biogeography of the Berchemia Complex is also discussed in this paper.This complex might have originated in the late Cretaceous in Colombia,South America,and dispersed to North America via Central America during the Eocene.Subsequently,the complex moved from North America to East Asia via the Bering Land Bridge no later than the late Eocene.Besides,the complex migrated from North America to Europe via the North Atlantic Land Bridge and then migrated further to Africa.In East Asia,it first appeared in Markam on the Qinghai-Tibetan Plateau,and then dispersed to other regions of Asia.     

The unique nature in ore composition,geological backgroudd and metallogenic mechanism of non-conventional superlarge ore deposits: A preliminary discussion

The new plants documented here, including a representative of the trimerophytesPsilophyton primitiuum sp. nov., a questionable rhyniophyte or trimerophyteHedeia sinica sp. nov., a prelycopodBragwanathia sp. and two species of zosterophyllophytes,Zosterophyllum australianum Lang and Cookson 1930 and2. sp. 1, from the Posongchong Formation of southeastern Yunnan, China, add to the known floral diversity of the Early Devonian of this region. Two sections of the Posongchong Formation, Changputang section of Wenshan district and Gegu section of Mengzi district also are introduced. After comparing the plants with those of the coeval flora of Australia and considering the data of recent paleocontinental reconstructions, the authors suggest that there is a northeastern Gondwana phytogeographic unit during the early Devonian comprising Australia, South China Block and perhaps Shan-Thai Block. The similarity of the floral component between Australia and South China is discussed. Because both Australia and South China also have dominant or endemic taxa, each might be recognized as a subunit, separately characterized byHedeia for Australia andEophyllophyton for South China. Project supported by the National Natural Science Foundation of China (Grant Nos. 49372075 and 49742004)