Impacts of climate change on primary production and carbon sequestration of boreal Norway spruce forests: Finland as a model

The aim of this study was to estimate the potential impacts of climate change on the spatial patterns of primary production and net carbon sequestration in relation to water availability in Norway spruce (Picea abies) dominated forests throughout Finland (N 60°–N 70°). The Finnish climatic scenarios (FINADAPT) based on the A2 emission scenario were used. According to the results, the changing climate increases the ratio of evapotranspiration to precipitation in southern Finland, while it slightly decreases the ratio in northern Finland, with regionally lower and higher soil water content in the south and north respectively. During the early simulation period of 2000–2030, the primary production and net carbon sequestration are higher under the changing climate in southern Finland, due to a moderate increase in temperature and atmospheric CO₂. However, further elevated temperature and soil water stress reduces the primary production and net carbon sequestration from the middle period of 2030–2060 to the final period of 2060–2099, especially in the southernmost region. The opposite occurs in northern Finland, where the changing climate increases the primary production and net carbon sequestration over the 100-year simulation period due to higher water availability. The net carbon sequestration is probably further reduced by the stimulated ecosystem respiration (under climate warming) in southern Finland. The higher carbon loss of the ecosystem respiration probably also offset the increased primary production, resulting in the net carbon sequestration being less sensitive to the changing climate in northern Finland. Our findings suggest that future forest management should carefully consider the region-specific conditions of sites and adaptive practices to climate change for maintained or enhanced forest production and carbon sequestration.     

Changed thinning regimes may increase carbon stock under climate change: A case study from a Finnish boreal forest

A physiological growth and yield model was applied for assessing the effects of forest management and climate change on the carbon (C) stocks in a forest management unit located in Finland. The aim was to outline an appropriate management strategy with regard to C stock in the ecosystem (C in trees and C in soil) and C in harvested timber. Simulations covered 100 years using three climate scenarios (current climate, ECHAM4 and HadCM2), five thinning regimes (based on current forest management recommendations for Finland) and one unthinned. Simulations were undertaken with ground true stand inventory data (1451 hectares) representing Scots pine (Pinus sylvestris), Norway spruce (Picea abies) and silver birch (Betula pendula) stands. Regardless of the climate scenario, it was found that shifting from current practices to thinning regimes that allowed higher stocking of trees resulted in an increase of up to 11% in C in the forest ecosystem. It also increased the C in the timber yield by up to 14%. Compared to current climatic conditions, the mean increase over the thinning regimes in the total C stock in the forest ecosystem due to the climate change was a maximum of 1%; but the mean increase in total C in timber yield over thinning regimes was a maximum of 12%.     

Computations on the influence of changing climate on the soil moisture and productivity in scots pine stands in southern and northern Finland

A model computation on the evaporative demand in relation to precipitation indicated that, under a changing climate with elevating temperatures, evapotranspiration could exceed the concurrent precipitation during the growing period in southern Finland (61° N), but not in northern Finland (66° N). This could reduce the supply of soil water to enable tree growth on sites with soil of low water holding capacity. This in turn could reduce the productivity of Scots pine more in southern Finland than in northern Finland. In northern Finland, the reduction in growth due to a limited supply of water was partly compensated by the enhanced growth due to a rise in temperature outside dry periods.     

The probability of wind damage in forestry under a changed wind climate

We (1) estimated how the possible changes in wind climate due to climatic change may affect the probability of exceeding critical wind speeds (CWS) expected to cause significant wind damage within a forest management unit located in southern Sweden, (2) analysed how the probability of exceeding an approximate CWS as observed in the management unit would change in different regions in Sweden if expecting a similar kind of forested area to occur in different geographical locations. The topography of the management unit was relatively gentle and the forests were dominated by Norway spruce (Picea abies (L.) Karst.). Seven regions across Sweden were selected for comparison of possible future probability of damaging wind speed. The model-system WINDA was modified and used for calculations of the probability of wind damage together with regionally downscaled climate change scenario (CCS) data. In total, two climate scenarios downscaled using the RCAO model for the control period 1961–1990 and four for the period 2071–2100 were used. The CCSs represent fairly central projections on a 100-year time scale in terms of global mean warming. Although there is ambiguity between different CCSs, the results indicated that the present pattern of more windy conditions in southern than in northern Sweden will remain. For most sites the probability of exceeding the CWS from westerly to south-westerly directions was indicated to remain comparatively high and the probability of damaging wind from south-westerly to south-easterly directions was indicated to increase in many places. For southernmost Sweden increasing probability of exceeding the CWS from the north-westerly to south-easterly wind directions were indicated for all but one CCS. The results were discussed with respect to spatial planning in forestry under a changing wind climate.     

Investigating the Balance between Timber Harvest and Productivity of Global Coniferous Forests under Global Change

A widely used assumption in forestry is that thedemand for timber will exceed the maximum levelavailable from forests on a sustainable basis. In thisstudy, measurements of extracted timber and modeledforest productivity were used to investigate therelationship between harvested timber and naturalforest productivity for current conditions, and underglobal change scenario. The analysis was confined toconiferous forests and countries that have coniferousforests within their territories. Annual roundwoodproduction from the database of Food and AgricultureOrganization was used as an approximation of annualtimber harvest for each country. Annual stem primaryproductivity of coniferous forests was estimated usingthe BIOME-BGC model. Based on the current rates,annual timber extraction was extrapolated for eachcountry for the next 80 years. Then, on a countrybasis, the timber harvest was related to the modeledforest stem productivity, assuming that the area ofconiferous forest would stay unchanged for the next 80years.The results of this study suggest that globalconiferous forests currently produce more wood thanpeople consume, but that this gap will narrow in thefuture. The results also suggest that wood extractionmay reach forest regrowth by the middle of the nextcentury, even though most coniferous forests arelocated in high latitudes and may have an acceleratedstem growth associated with the joint effect ofclimate change and elevated carbon dioxideconcentration in the atmosphere.     

Climate affects severity and altitudinal distribution of outbreaks in an eruptive bark beetle

Temperature warming and the increased frequency of climatic anomalies are expected to trigger bark beetle outbreaks with potential severe consequences on forest ecosystems. We characterized the combined effects of climatic factors and density-dependent feedbacks on forest damage caused by Ips typographus (L.), one of the most destructive pests of European spruce forests, and tested whether climate modified the interannual variation in the altitudinal outbreak range of the species. We analyzed a 16-year time-series from the European Alps of timber loss in Picea abies Karsten forests due to I. typographus attacks and used a discrete population model and an information theoretic approach to compare multiple competing hypotheses. The occurrence of dry summers combined with warm temperatures appeared as the main abiotic triggers of severity of outbreaks. We also found an endogenous negative feedback with a 2-year lag suggesting a potential important role of natural enemies. Forest damage per hectare averaged 7-fold higher where spruce was planted in sites warmer than those within its historical climatic range. Dry summers, but not temperature, was related to upward shifts in the altitudinal outbreak range. Considering the potential increased susceptibility of spruce forests to insect outbreaks due to climate change, there is growing value in mitigating these effects through sustainable forest management, which includes avoiding the promotion of spruce outside its historical climatic range.     

Impacts of Present and Future Climate Variability on Agriculture and Forestry in the Temperate Regions: Europe

Agriculture and forestry will be particularly sensitive to changes in mean climate and climate variability in the northern and southern regions of Europe. Agriculture may be positively affected by climate change in the northern areas through the introduction of new crop species and varieties, higher crop production and expansion of suitable areas for crop cultivation. The disadvantages may be determined by an increase in need for plant protection, risk of nutrient leaching and accelerated breakdown of soil organic matter. In the southern areas the benefits of the projected climate change will be limited, while the disadvantages will be predominant. The increased water use efficiency caused by increasing CO₂ will compensate for some of the negative effects of increasing water limitation and extreme weather events, but lower harvestable yields, higher yield variability and reduction in suitable areas of traditional crops are expected for these areas. Forestry in the Mediterranean region may be mainly affected by increases in drought and forest fires. In northern Europe, the increased precipitation is expected to be large enough to compensate for the increased evapotranspiration. On the other hand, however, increased precipitation, cloudiness and rain days and the reduced duration of snow cover and soil frost may negatively affect forest work and timber logging determining lower profitability of forest production and a decrease in recreational possibilities. Adaptation management strategies should be introduced, as effective tools, to reduce the negative impacts of climate change on agricultural and forestry sectors.     

Decreasing teleconnections with inter-site distance in monthly climatic data and tree-ring width networks in a mountainous Alpine area

Summary The similarities in time series recorded at sites which are distant from each other are called teleconnections. In this paper, the loss of such correlations with inter-site distance was investigated for both climatic and dendrochronological data sets, with 70 tree-ring chronologies. A dense network of weather stations was studied in the southeastern French Alps, covering complex climatic gradients over three departments. 78 sites with precipitation data (with a total of 48 756 monthly values), and 48 stations that recorded temperature (with 20 722 monthly mean values) were analysed. In the same area, four coniferous species (mountain pine and stone pine, European larch and Norway spruce) provided 37 ring-width chronologies for high elevation sites near the timberline. Both silver fir and Norway spruce provided a second tree-ring chronology network for 33 different sites at lower elevations. The teleconnections between precipitation series were found to be higher than those observed for temperature over short distances, but the maximum threshold distance was lower (193 km) compared to a positive correlation distance that exceeds 500 km for temperature. The maximum temperatures had stronger teleconnections than minimum values (522 km versus 476 km), since the latter are linked more with other site factors, such as slope, exposure and local topography. As expected, the tree-ring chronologies showed weaker teleconnections than the climatic series, with a threshold distance of 374 km obtained for all high elevation forests. The coniferous species with high intra-specific teleconnections over large distances were, in decreasing importance, Pinus uncinata (> 500 km), Picea abies (477 km), Pinus cembra (over 254 km) and Larix decidua (over 189 km only). The two former species showed the highest intra-specific correlations (with mean correlation R=0.625 and 0.666). The dendrochronological teleconnections were found to have a extent lesser for trees species that depend on rainfall (such as larch, and stone pine). They are enhanced, however, for temperature sensitive species such as spruce and mountain pine (a drought resistant tree). Therefore, these two latter conifers appear to be especially suitable for climatic reconstruction over large distances in mountainous areas. However, teleconnections within silver fir (Abies alba) and spruce chronologies were sharply reduced (over 131 km and 135 km) in lower elevation forests, underlining the interest of timberline forests for dendroclimatology. A better knowledge of the spatial correlations in climatic series and ring-width data may enable the optimisation of weather station networks. It may also permit a better choice of weather stations used for dendroclimatology, either for tree-ring and climate relationship calibration or for climate reconstructions. In dendrochronology, wood dating also requires the knowledge of to what extent remote ring-width chronologies can be used. Received September 11, 2000 Revised March 26, 2001     

On predicting the response of forests in eastern North America to future climatic change

Our ability to accurately predict the response of forests in eastern North America to future climatic change is limited by our knowledge of how different tree species respond to climate. When the climatic response of eastern hemlock is modeled across its range, we find that the assumed climatic response used in simulation models is not sufficient to explain how this species is presently responding to climate. This is also the case for red spruce growing in the northern Appalachian Mountains. Consequently, simulations of future change to forests that include eastern hemlock and red spruce may need to be improved. We suspect that similar findings will be made when other tree species are studied in detail using tree-ring analysis. If so, our present understanding of how individual tree species respond to climate may not be adequate for accurately predicting future changes to these forests. Tree-ring analysis can increase our understanding of how climate affects tree growth in eastern North America and, hence, provide the knowledge necessary to produce more accurate predictions.     

全球FLUXNET站点总初级生产力的年际变化及其主导因子解析

基于耶鲁互动生物圈模式（YIBs）,结合FLUXNET网络观测时间超过8 a的站点的观测数据,研究对不同气象因子影响下总初级生产力（Gross Primary Productivity,GPP）的年际变化进行分析,探讨各植被类型GPP年际变化的主要驱动因子。结果表明,光合有效辐射的变化是落叶阔叶林和常绿针叶林站点GPP年际变化的主要驱动因子,贡献了这些森林类型年际变率的80%。相对湿度变化是作物站点GPP年际变化的主要驱动因子,贡献了作物GPP年际变率的65%。温度是灌木丛站点GPP年际变率的重要因子,其贡献率达到58%。草原站点GPP年际变化的不确定性较大,没有统一的主导因子。研究结果表明,气象要素是全球陆地GPP年际变化的主导因素。在未来气候变化背景下,极端气候事件频发可能会进一步增加GPP的年际变率。     

新疆伊犁乌孙山北坡树轮灰度年表的建立及气候意义

利用2006年采自新疆伊犁乌孙山北坡3个采点的树轮样本,建立了树轮宽度年表,采用树轮图像分析法进一步建立了3种宽度年表和5种灰度年表。宽度年表对比发现,树木年轮对气候响应南疆强于北疆,北疆其它地方强于伊犁地区,而乌孙山北坡与伊犁其它地区比较,树轮宽度对气候的响应较敏感。树轮图像分析法建立的8种年表特征进行对比发现,恰依恰孜宽度年表对气候的响应最为强烈;而阿乌里亚乔克标准化灰度年表对气候的响应较好。相关分析表明,全轮灰度、早材灰度和最大灰度与4、5月气温相关较好。4月降水和实测宽度、全轮宽度以及早材宽度相关较好,而6月降水与全轮灰度和早材灰度相关较好。早材宽度对气候的响应强于晚材宽度;全轮灰度和早材灰度对气候的响应较好。利用树轮图像分析所得到的8个树轮参数年表可增加气候重建代用指标,提高气候重建的精度。     

Modelling Regional Climate Change Effects On Potential Natural Ecosystems in Sweden

This study aims to demonstrate the potential of a process-based regional ecosystem model, LPJ-GUESS, driven by climate scenarios generated by a regional climate model system (RCM) to generate predictions useful for assessing effects of climatic and CO₂ change on the key ecosystem services of carbon uptake and storage. Scenarios compatible with the A2 and B2 greenhouse gas emission scenarios of the Special Report on Emission Scenarios (SRES) and with boundary conditions from two general circulation models (GCMs) – HadAM3H and ECHAM4/OPYC3 – were used in simulations to explore changes in tree species distributions, vegetation structure, productivity and ecosystem carbon stocks for the late 21st Century, thus accommodating a proportion of the GCM-based and emissions-based uncertainty in future climate development. The simulations represented in this study were of the potential natural vegetation ignoring direct anthropogenic effects. Results suggest that shifts in climatic zones may lead to changes in species distribution and community composition among seven major tree species of natural Swedish forests. All four climate scenarios were associated with an extension of the boreal forest treeline with respect to altitude and latitude. In the boreal and boreo-nemoral zones, the dominance of Norway spruce and to a lesser extent Scots pine was reduced in favour of deciduous broadleaved tree species. The model also predicted substantial increases in vegetation net primary productivity (NPP), especially in central Sweden. Expansion of forest cover and increased local biomass enhanced the net carbon sink over central and northern Sweden, despite increased carbon release through decomposition processes in the soil. In southern Sweden, reduced growing season soil moisture levels counterbalanced the positive effects of a longer growing season and increased carbon supply on NPP, with the result that many areas were converted from a sink to a source of carbon by the late 21st century. The economy-oriented A2 emission scenario would lead to higher NPP and stronger carbon sinks according to the simulations than the environment-oriented B2 scenario.     

Estimates of monoterpene and isoprene emissions from the forests in Switzerland

Emission rates of biogenic volatile organic compounds emitted by the forests were estimated for five geographical regions as well as for all Switzerland. Monoterpene and isoprene emissions rates were calculated for each main tree species separately using the relevant parameters such as temperature, light intensity and leaf biomass density. Biogenic emissions from the forests were found to be about 23% of the total annual VOC emissions (anthropogenic and biogenic) in Switzerland. The highest emissions are in July and lowest in January. Calculations showed that the coniferous trees are the main sources of the biogenic emissions. The major contribution comes from the Norway spruce (picea abies) forests due to their abundance and high leaf biomass density. Although broad-leaved forests cover 27% of all the forests in Switzerland, their contribution to the biogenic emissions is only 3%. Monoterpenes are the main species emitted, whereas only 3% is released as isoprene. The highest emission rates of biogenic VOC are estimated to be in the region of the Alps which has the largest forest coverage in Switzerland and the major part of these forests consists of Norway spruce. The total annual biogenic VOC emission rate of 87 ktonnes y^–1 coming from the forests is significantly higher than those from other studies where calculations were carried out by classifying the forests as deciduous and coniferous. The difference is attributed to the high leaf biomass densities of Norway spruce and fir (abies alba) trees which have a strong effect on the results when speciation of trees is taken into account. Besides the annual rate, emission rates were calculated for a specific period during July 4–6, 1991 when a photochemical smog episode was investigated in the Swiss field experiment POLLUMET. Emission rates estimated for that period agree well with those calculated for July using the average temperatures over the last 10 years.     

Increasing Aridity is Enhancing Silver Fir Abies Alba Mill.) Water Stress in its South-Western Distribution Limit

Tree populations located at the geographical distribution limit of the species may provide valuable information about the response of tree growth to climate warming across climatic gradients. Dendroclimatic information was extracted from a network of 10 silver-fir (Abies alba) populations in the south-western distribution limit of the species (Pyrenees, NE Iberian Peninsula). Ring-width chronologies were built for five stands sampled in mesic sites from the Main Range in the Pyrenees, and for five forests located in the southern Peripheral Ranges where summer drought is more pronounced. The radial growth of silver-fir in this region is constrained by water stress during the summer previous to growth, as suggested by the negative relationship with previous September temperature and, to a lesser degree, by a positive relationship with previous end of summer precipitation. Climatic data showed a warming trend since the 1970s across the Pyrenees, with more severe summer droughts. The recent warming changed the climate-growth relationships, causing higher growth synchrony among sites, and a higher year-to-year growth variation, especially in the southernmost forests. Moving-interval response functions suggested an increasing water-stress effect on radial growth during the last half of the 20th century. The growth period under water stress has extended from summer up to early autumn. Forests located in the southern Peripheral Ranges experienced a more intense water stress, as seen in a shift of their response to precipitation and temperature. The Main-Range sites mainly showed a response to warming. The intensification of water-stress during the late 20th century might affect the future growth performance of the highly-fragmented A. alba populations in the southwestern distribution limit of the species.     

Integrating climatic change and forests: Economic and ecologic assessments

Effective policies for dealing with anticipated climatic changes must reflect the two-way interactions between climate, forests and society. Considerable analysis has focused on one aspect of forests - timber production - at a local and regional scale, but no fully integrated global studies have been conducted. The appropriate ecological and economic models appear to be available to do so. Nontimber aspects of forests dominate the social values provided by many forests, especially remote or unmanaged lands where the impacts of climatic change are apt to be most significant. Policy questions related to these issues and lands are much less well understood. Policy options related to afforestation are well studied, but other ways the forest sector can help ameliorate climatic change merit more extensive analysis. Promising possibilities include carbon taxes to influence the management of extant forests, and materials policies to lengthen the life of wood products or to encourage the substitution of CO₂-fixing wood products for ones manufactured from less benign materials.     

Impacts of Present and Future Climate Change and Climate Variability on Agriculture in the Temperate Regions: North America

The potential impact of climate variability and climate change on agricultural production in the United States and Canada varies generally by latitude. Largest reductions are projected in southern crop areas due to increased temperatures and reduced water availability. A longer growing season and projected increases in CO₂ may enhance crop yields in northern growing areas. Major factors in these scenarios analyzes are increased drought tendencies and more extreme weather events, both of which are detrimental to agriculture. Increasing competition for water between agriculture and non-agricultural users also focuses attention on water management issues. Agriculture also has impact on the greenhouse gas balance. Forests and soils are natural sinks for CO₂. Removal of forests and changes in land use, associated with the conversion from rural to urban domains, alters these natural sinks. Agricultural livestock and rice cultivation are leading contributors to methane emission into the atmosphere. The application of fertilizers is also a significant contributor to nitrous oxide emission into the atmosphere. Thus, efficient management strategies in agriculture can play an important role in managing the sources and sinks of greenhouse gases. Forest and land management can be effective tools in mitigating the greenhouse effect.     

Possible Impacts of Climate Change on Soil Moisture Availability in the Southeast Anatolia Development Project Region (GAP): An Analysis from an Agricultural Drought Perspective

This paper presents probable effects of climate change on soil moisture availability in the Southeast Anatolia Development Project (GAP) region of Turkey. A series of hypothetical climate change scenarios and GCM-generated IPCC Business-as-Usual scenario estimates of temperature and precipitation changes were used to examine implications of climate change for seasonal changes in actual evapotranspiration, soil moisture deficit, and soil moisture surplus in 13 subregions of the GAP. Of particular importance are predicted patterns of enhancement in summer soil moisture deficit that are consistent across the region in all scenarios. Least effect of the projected warming on the soil moisture deficit enhancement is observed with the IPCC estimates. The projected temperature changes would be responsible for a great portion of the enhancement in summer deficits in the GAP region. The increase in precipitation had less effect on depletion rate of soil moisture when the temperatures increase. Particularly southern and southeastern parts of the region will suffer severe moisture shortages during summer. Winter surplus decreased in scenarios with increased temperature and decreased precipitation in most cases. Even when precipitation was not changed, total annual surplus decreased by 4 percent to 43 percent for a 2°C warming and by 8 percent to 91 percent for a 4°C warming. These hydrologic results may have significant implications for water availability in the GAP as the present project evaluations lack climate change analysis. Adaptation strategies – such as changes in crop varieties, applying more advanced dry farming methods, improved water management, developing more efficient irrigation systems, and changes in planting – will be important in limiting adverse effects and taking advantage of beneficial changes in climate.     

Surface fluxes of heat and water vapour from sites in the European Arctic

Summary  Measurements of the surface fluxes of heat and water vapour were taken at four sites across the European Arctic as part of the EU funded LAPP project. The sites cover a range of latitudinal, altitudinal and climatic conditions. The most northerly site is near Ny-?lesund, Svalbard, a polar semi-desert with continuous permafrost. A second permafrost site is a fen area in the Zackenberg valley, East Greenland. Finally two sites in northern Finland, Skalluvaara and Kaamanen are on the southern boundary of the region affected by permafrost. At all sites measurements were made of the turbulent fluxes of heat and water vapour using eddy correlation equipment for at least one active season. The net radiation totals for July and August are similar at all sites. At the sites with permafrost a substantial proportion (over 20%) of the net radiation goes into soil heat flux, to thaw the soil moisture in the top metre. Of the remaining energy just over half is used for evaporation. At the Finnish sites the vegetation is largely deciduous and this is seen in the record with higher evaporative ratios in July and August, after the vegetation becomes green. The Finnish sites tend to have higher surface resistance to evaporation; however, the evaporative demand is greater leading to slightly higher evaporation rates. The two Finnish sites have a similar seasonal pattern determined by the water table and seasonality of the vegetation. The two northern sites show a pattern that is determined primarily by the variation of water table only. It is concluded that the water balance through the active season is influenced primarily by the history of snow cover. The seasonality of the vegetation, the permafrost and the depth of water table are also important influences. Received November 1, 1999 Revised April 17, 2000     

Phytomass carbon pool of trees and forests in India

The study reports estimates of above ground phytomass carbon pools in Indian forests for 1992 and 2002 using two different methodologies. The first estimate was derived from remote sensing based forest area and crown density estimates, and growing stock data for 1992 and 2002 and the estimated pool size was in the range 2,626–3,071 Tg C (41 to 48 Mg C ha^???1) and 2,660–3,180 Tg C (39 to 47 Mg C ha^???1) for 1992 and 2002, respectively. The second methodology followed IPCC 2006 guidelines and using an initial 1992 pool of carbon, the carbon pool for 2002 was estimated to be in the range of 2,668–3,112 Tg C (39 to 46 Mg C ha^???1), accounting for biomass increment and removals for the period concerned. The estimated total biomass increment was about 458 Tg over the period 1992–2002. Removals from forests include mainly timber and fuel wood, whereby the latter includes large uncertainty as reported extraction is lower than actual consumption. For the purpose of this study, the annual extraction values of 23 million m³ for timber and 126 million m³ for fuel wood were used. Out of the total area, 10 million ha are plantation forests with an average productivity (3.2 Mg ha^???1 year^???1) that is higher than natural forests, a correction of 408 Tg C for the 10 year period was incorporated in total estimated phytomass carbon pool of Indian forests. This results in an estimate for the net sink of 4 Tg C year^???1. Both approaches indicate Indian forests to be sequestering carbon and both the estimates are in agreement with recent studies. A major uncertainty in Indian phytomass carbon pool dynamics is associated with trees outside forests and with soil organic carbon dynamics. Using recent remote-sensing based estimates of tree cover and growing stock outside forests, the estimated phytomass carbon pool for trees outside forests for the year 2002, is 934 Tg C with a national average tree carbon density of 4 Mg C ha^???1 in non-forest area, in contrast to an average density of 43 Mg C ha^???1 in forests. Future studies will have to consider dynamics in both trees outside forests and soil for total terrestrial carbon dynamics.     

东亚季风区夏季陆地生态系统碳循环对东亚夏季风的响应

东亚地区陆地生态系统的时空变率表现出明显的对季风气候的响应特征。使用EOF（经验正交分解）方法分析了AVIM2动态植被陆面模式离线模拟试验模拟的1953～2004年东亚季风区夏季陆地生态系统总初级生产力（GPP）、生态系统净初级生产力（NPP）、净生态系统初级生产力（NEP）、植被呼吸以及土壤呼吸的时空分布特点,探讨了东亚夏季风对陆地生态系统碳循环影响机制。研究发现,在强季风年,江淮地区高温少雨的特点限制了光合作用,造成GPP偏低;而华南地区在强季风年气候温暖湿润,利于植被生长,GPP偏高。季风对于植被呼吸和土壤呼吸影响不明显,使得GPP和植被呼吸之差NPP的变化及NPP和土壤呼吸之差NEP的变化与GPP的变化保持一致。在强季风年江淮流域地区干热的气候条件使得NPP和NEP降低;但是在华南地区温度升高的同时降水增多使得在NPP偏高的基础上NEP也偏高。