Adamantinasuchus navae: A new Gondwanan Crocodylomorpha (Mesoeucrocodylia) from the Late Cretaceous of Brazil

A new Late Cretaceous Mesoeucrocodylian from the Adamantina Formation (Bauru Basin), São Paulo State, Brazil is described. The main features of this new species are the short, high oreinirostral rostrum, the large laterally positioned orbital notches and external nares in the anteriormost portion of the rostrum. The mandible is robust and concave-shaped in relation to the skull. The dentition is highly specialized, with two prominent incisiform teeth, a hypertrophied caniniform, and seven molariform teeth. The molariform teeth are ornamented with denticles in their lingual surface and are smooth on their labial surface. The molariforms are elliptical in cross-section, presenting the largest axis in the labial-lingual direction. Such dental characteristics are unique among the terrestrial crocodylomorphs of the Gondwana.     

Gondwanasuchus scabrosus gen. et sp. nov., a new terrestrial predatory crocodyliform (Mesoeucrocodylia: Baurusuchidae) from the Late Cretaceous Bauru Basin of Brazil

Baurusuchids are among the most common and diverse crocodyliform fossils from the Late Cretaceous Bauru Basin of Brazil. This inland continental basin was the habitat of a rich crocodyliform fauna containing five mesoeucrocodylian families, of which the Baurusuchidae represents highly specialized predatory crocodyliforms of terrestrial habits as indicated by their dental, cranial, and postcranial features. The large size they achieved, together with likely predatory adaptations, would suggest they competed and occupied theropod ecological niches in the Bauru Basin. Here we describe Gondwanasuchus scabrosus gen. et sp. nov., a medium-sized baurusuchid with a strongly laterally compressed skull, bearing unique dentition with deep apicobasal sulci and probably well-developed binocular vision. The cranial and dental features in Gondwanasuchus suggest that this active predator would have fed on small vertebrates and took the role of small theropods in terrestrial guild. Gondwanasuchus is the most distinctive baurusuchid known to date and enriches the knowledge on these important Gondwanan terrestrial predatory crocodyliforms.     

Baurusuchus salgadoensis, a New Crocodylomorpha from the Bauru Basin (Cretaceous), Brazil

Baurusuchus salgadoensis is a new baurusuchid crocodylomorph from Bauru Basin (Cretaceous), Brazil, partially preserved through a complete skull. The fossil comes from a fine sandstone sequence of Adamantina Formation, General Salgado County, São Paulo State. The sedimentary sequence where it was found, located in Fazenda Buriti, is considered Turonian-Santonian in age. The described species _Baurusuchus salgadoensis sp. nov. _ is a baurusuchid with an antorbital fenestra, double external nares with a bony septum, two well-fused supraorbitals, the supratemporal fenestrae larger than the orbits and a quadrangular-shaped laterotemporal fenestra. The position of the external nares, located on anterior and terminal portion of the rostrum together with the theropod-like lateral compression of the snout and teeth are indicators that Baurusuchus salgadoensis was a terrestrial crocodyliform. This was a carnivorous species and the lateral compression of the rostrum could be interpreted as a mechanism to increase the skull resistance forces during biting. The pointed, conical teeth, some with crenulated borders, could be used to perforate and to carve the prey. The geological context of Baurusuchus salgadoensis indicates that it probably lived in a hot and arid climate.     

Uberabasuchus terrificus sp. nov., a New Crocodylomorpha from the Bauru Basin (Upper Cretaceous), Brazil

A new fairly complete and articulated skull of a Peirosauridae crocodylomorph from Bauru Basin (Late Cretaceous), Brazil, is described. The fossil is from a level of clayish sandstone within Serra do Veadinho sequence, Peirópolis, Uberaba County, Minas Gerais State. The sedimentary strata of Serra do Veadinho belong to the Marília Formation (Serra da Galga Member), Bauru Group, considered to be Campanian-Maastrichtian in age. The species -Uberabasuchus terrificus sp. nov. - is a peirosaurid with moderately narrow snout, large round orbits protected by supraorbital bones of triangular shape and an antorbital fenestra bounded posteriorly by a deep groove. This fossil resembles Peirosaurus tormini Price, 1955 in the size pattern of premaxillary teeth and by showing a similar wedge-like maxillary process in the premaxilla. It also shares some morphological features with the other species of the Peirosauridae, namely the crocodylomorph Lomasuchus palpebrosus Gasparini, Chiappe and Fernandez, 1991 from Argentina. Their common features comprise a moderately narrow snout and the deep lateral groove at the premaxilla and maxilla articulation for the reception of a large mandibular tooth. However, the nasal participates in the external nares and does not divide the nasal aperture, producing a “beak-like” structure at the extremity of the snout which is unique among peirosaurids. The stratigraphic setting suggests that the specimen was buried when a flash flood overflowed the shallow channels of a braided fluvial system.

Parsimony analysis of 183 morphological characters is performed for 23 crocodylomorphs. Analysis of the morphological data matrix resulted in three most parsimonious trees (374 steps, CI = 0.679; RI = 0.826). The new species is closely related to Mahajangasuchus and both, in addition to Peirosaurus and Lomasuchus, compose the Peirosauridae.     

Climate-induced sediment-palaeosol cycles in a Late Cretaceous dry aeolian sand sheet: Marília Formation (North-West Bauru Basin, Brazil)

Aeolian sand sheets, which are characterized by low relief surfaces that lack dunes, are common in arid and semi‐arid climatic settings. The surface of an aeolian sand sheet can either be stable and subject to pedogenetic effects, or unstable such that it is affected by deflation or sedimentation. The Marília Formation (Late Cretaceous) may be interpreted as an ancient aeolian sand sheet area, where alternating phases of stability and instability of the accumulation surface have been recorded. Detailed field studies were carried out in several sections of the Marília Formation, where cyclic alternations of palaeosols and aeolian deposits were evident, using palaeopedological and facies analysis methods, supported in the laboratory by the analysis of rock samples, cut and polished in slabs, thin sections, scanning electron microscope images and X‐ray diffraction data from the clay minerals. The deposits comprise three lithofacies that, in order of abundance, are characterized by: (i) translatent wind‐ripple strata; (ii) flood deposits; and (iii) ephemeral river channel deposits. Palaeosols constitute, on average, 65% of the vertical succession. Three types of palaeosols (pedotypes) are recognized: (i) Aridisols; (ii) Entisols; and (iii) Vertisols. Erosional surfaces due to aeolian deflation divide the top of the palaeosol profiles from the overlying aeolian deposits. The palaeoenvironmental interpretation of the deposits and the palaeosols allows the depositional system of the Marília Formation to be defined as a flat area, dominated by aeolian sedimentation, with subordinate ephemeral river sedimentation, and characterized by a dry climatic setting with occasional rainfall. The climate is the main forcing factor controlling the alternation between episodes of active sedimentation and periods of palaeosol development. A climate‐controlled model is proposed in which: (i) the palaeosols are indicative of a stable surface that is developed during the more humid climatic phases; and (ii) the erosional surfaces and the overlying aeolian sediments attest to periods of deflation and subsequent sedimentation, thereby increasing the availability of sediment during the drier climatic phases. The ephemeral fluvial deposits mark the more humid climatic conditions and contribute to the lagged sediment influx caused during the drier periods by the erosion of previously stored sediment.     

Taphonomy of a Baurusuchus (Crocodyliformes,Baurusuchidae) from the Adamantina Formation (Upper Cretaceous,Bauru Basin), Brazil: Implications for preservational modes,time resolution and paleoecology

Upper Cretaceous vertebrate accumulations from the Adamantina Formation are known due to their high taxonomic diversity. On the other hand, taphonomic analyses still are rare, limiting the understanding of processes related to the biostratinomic and fossildiagenetic histories of this lithostratigraphic unit. In 2005, fossils were collected from an outcrop located at Jales municipality, state of São Paulo, southeastern Brazil. From this outcrop, a well-preserved Baurusuchus was recovered, which displays a peculiar set of taphonomic signatures. This paper identifies and interprets taphonomic features of a specimen of Baurusuchus (Crocodyliformes, Baurusuchidae; UFRJ DG 418-R) from the Adamantina Formation (Upper Cretaceous of the Bauru Basin), in Jales municipality, state of São Paulo. Brief taphonomic comparisons with other specimens previously studied (crocodiles and dinosaurs) and a lithofaciological analysis of the outcrop were undertaken in order to enhance the knowledge of the stratigraphy and paleoenvironment and improve the time resolution for the Adamantina Formation in the region of Jales. Furthermore, paleoecological data were interpreted based on the taphonomic analysis.The fossil is composed of an articulated segment of vertebral column, ribs, part of the pelvic girdle and gastralia. There is no hydraulic equivalence between both the specimen of Baurusuchus and the size of quartz grain predominant in the fossiliferous layer, suggesting death in situ or short transport as a “water carcass”. Teeth marks identified on the pubes were assigned to a small/juvenile baurusuchid crocodyliform or a theropod dinosaur. The repositioning of some elements (ribs and dorsal osteoderms) is suggestive of mummification. Desiccation marks were observed and attributed to the stage 1 of weathering. These features suggest subaerial exposure of the carcass prior to burial, however, probably after the mummification. On the other hand, the subaerial exposure was short, because the individual was not fully disarticulated. Furthermore, the degrees of articulation and preservation of the specimen nullify the hypothesis of reworking.Lithofaciological and taphonomic analyses suggest a well-drained floodplain as the burial environment and an arid or semi-arid climate in the region of Jales outcrop. In addition, the taphonomic signatures seem to indicate a time resolution about 10⁰–10⁴ years for the layer where the crocodyliforms were collected, but it seems to have, within this time limit, temporal-mixing among terrestrial crocodiles and dinosaurs collected from the same layer, suggesting time-averaging in this outcrop.     

Tetrapods from the Upper Cretaceous (Turonian–Maastrichtian) Bauru Group of Brazil: a reappraisal

An updated, annotated list of all tetrapods from the Adamantina, Uberaba and Marília formations (Bauru Group), which constitute some of the best studied Upper Cretaceous units in Brazil, is presented. Tetrapod diversity in the Bauru Group is remarkable, including an admixture of typically austral Gondwanan taxa (e.g., abelisaurids, notosuchians) and boreal Gondwanan forms (e.g., carcharodontosaurids). Of note is the absence of Laurasian taxa in the upper portion of the Bauru Group. With the exception of some turtles, an anuran, mesoeucrocodylians and one titanosaur, most taxa from the Bauru Group are based on fragmentary and isolated bones, and as such many specimens can be identified only to a higher taxonomic level. Fishes, turtles, anurans, mesoeucrocodylians, dinosaurs, birds and mammals from the Adamantina and Marília formations resemble the latest Late Cretaceous vertebrate faunas from southern South America, except for the absence of ornithischian dinosaurs.     

A new species of Leptotarsus (Diptera: Tipulidae) from the Lower Cretaceous Crato Formation of Brazil

Leptotarsus (sensu lato) lukashevichae sp. nov. is described and illustrated, based on a single but very well preserved female specimen from the Crato Formation of Brazil (Aptian, ca. 112 Ma). Along with other Leptotarsus species recently described from Lower Cretaceous beds of Brazil, Spain, Russia and China, this new species is among the oldest known records of the genus Leptotarsus and the family Tipulidae.     

A New Oviraptorid Dinosaur (Theropoda: Oviraptorosauria) from the Late Cretaceous of Jiangxi Province, Southern China

A new oviraptorid dinosaur Jiangxisaurus ganzhouensis gen. et sp. nov., is erected based on a partial skeleton from the Upper Cretaceous Nanxiong Formation of Ganzhou City, Jiangxi Province. The new taxon differs from other oviraptorids in the weakly downturned rostrum of the lower jaw, much-elongated mandible with a height-to-length ratio being about 20% and the length ratio of radius to humerus of about 0.70. This species not only adds a new member to oviraptorid dinosaurs, but also provides more information about oviraptorid paleogeographical distribution in southern China.     

A new pterodactyloid pterosaur from the Santana Formation (Cretaceous) of Brazil

A partial skull comprising fused maxilla/premaxilla and palate of a ctenochasmatoid pterosaur from the Santana Formation of the Araripe Basin in NE Brazil is named as the new genus and species Unwindia trigonus gen. et sp. nov. on account of its long slender rostrum, isodonty with raised dental alveoli and dentition of seven tooth pairs restricted to the portion of the rostrum anterior to the nasoantorbital fenestra. Unwindia is assigned to the Ctenochasmatoidea, and is probably basal within the clade.     

A new titanosaur from the Lower Cretaceous of Brazil

Here we describe a new record of a sauropod dinosaur from the Lower Cretaceous (Hauterivian–Barremian) Rio Piranhas Formation, Sousa Basin, NE Brazil. Dinosaur fossil bones from this deposit were unknown until now. Thus, the discovery of a sauropod fibula from this locality is highly significant. Our discovery represents an indeterminate titanosaur and the earliest stratigraphic occurrence of this group in central Gondwana. When compared to chronocorrelate titanosaur trackmakers of this geological unit, this fossil specimen appears substantially smaller. Histological analysis of the fibula suggests that this is a relatively young individual (approximately 40–50% adult body size) that had passed its most rapid phase of early juvenile growth, but had not yet attained somatic maturity. Thus, the fibula recovered is from a young individual rather than from a small-bodied adult titanosaur.     

Late Cretaceous lithoevents in the Bohemian Cretaceous Basin, Czechoslovakia

Facies evolution and vertical changes within the Late Cretaceous sequence of the Bohemian Cretaceous Basin reveal fluctuations of intra- and extrabasinal circumstances. Evidence of periodic oscillations is recognized and two categories of aperiodic event indications are distinguished according to their significance and lateral persistence. Several lithoevents may be related to eustatic changes, while others indicate the independent evolution of the basin, influenced by epeirogenetic movements of segments of the Bohemian Massif.     

A new tree fern stem,Tempskya zhangii sp. nov. (Tempskyaceae) from the Cretaceous of Northeast China

A new Cretaceous tree fern species from the family Tempskyaceae, Tempskya zhangii sp. nov., from Keshan County, Heilongjiang Province, Northeast China, is described based on silicified fragments. The species is characterized by dichotomizing stems embedded in a matrix of adventitious roots, together forming a solid and compact false trunk. The dorsiventral stems are composed of solenosteles, and their internodes are short with 2–3 (rarely 4) leaf traces. A three-layered cortex includes a sclerenchymatous outer zone, parenchymatous middle zone, and an inner zone composed of sclerenchyma band and parenchyma cells. The pith consists of a parenchymatous outer zone and sclerenchymatous inner zone. The petiole has a single C- or U-shaped vascular strand. The adventitious root has diarch xylem surrounded by an inner sclerenchymatous and an outer parenchymatous cortex. In possessing these characters, T. zhangii is more similar to T. wyomingensis from the Cretaceous of North America than to other species of Tempskya. However, the Chinese species distinctly differs from T. wyomingensis in having a sclerenchymatous outer cortex, parenchymatous middle cortex, and sparse parenchyma in the xylem. It is the first record of Tempskya from China and the fourth from Asia. This species, together with Cyatheaceae, Osmundaceae, Cycadeoidaceae, and conifers found in the same region constituted the plant community at the time.     

A new species of Cyrtophyllitinae (Insecta: Ensifera) from the Cretaceous China

A new fossil species of Cyrtophyllitinae, Vitimoilus ovatus sp. nov., is described from the Dabeigou Formation and Yixian Formation of the Lower Cretaceous of China. It is clearly placed in Vitimoilus Gorochov, 1996 according to the following characters: R forking into RA and RP distally, M forking into MA and MP distally; CuA fused with CuPaα distal of the middle of the wing length; cross-veins in basal part of CuPb–CuPaβ area non-strongly curved.     

山东晚白垩世一新的巨龙类恐龙

记述了产自山东诸城上白垩统王氏群中一个较完整的蜥脚类恐龙肱骨(ZJZ-57)。它具有一些进步特征,如肱骨三角胸嵴向远端扩展,肱骨近端外侧1/3处发育隆凸,表明其属于较进步的巨龙类成员。另外,它具有一些独特的鉴定特征组合区别于其他任何巨龙类:近端横向强烈扩展,近端最大宽度与骨干长度之比为0.55;骨干粗壮,粗壮指数为0.39。综合研究表明,ZJZ-57所属个体代表了一种新的巨龙类恐龙:臧家庄诸城巨龙(Zhuchengtitan zangjiazhuangensis gen.et sp.nov.),它可能与Opisthocoelicaudia的亲缘关系较接近。这是山东晚白垩世地层中首次发现的巨龙类恐龙。     

A remarkable brachyceran fly (Diptera: Tabanomorpha) from Late Cretaceous Burmese amber

A new brachyceran fly (Diptera: Tabanomorpha), Pseudorhagio zhangi gen. et sp. nov., is described from Late Cretaceous Burmese amber. It is tentatively placed as Family incertae sedis in Tabanomorpha and distinguished from other Tabanomorpha by the following unusual combination of characters: head wider than thorax; body densely covered with fine and short setae, devoid of macrosetae; scutum strongly convex, nearly spherical; scutellum rather small, convex; antennal flagellum elongated, tapering, unsegmented; vein R4 perpendicular to R5, and strongly curved; crossvein m-m very long, strongly sinuated; tibial spur formula 0, 2, 0. This discovery further confirms the high diversity of Tabanomorpha in Late Cretaceous Burmese amber. An updated list of brachyceran species in Burmese amber is given.     

A new genus of Tanyderidae (Insecta: Diptera) from Myanmar amber,Upper Cretaceous

A new genus and species, Similinannotanyderus lii gen. et sp. nov., is described from Myanmar amber. It differs from other genera of Tanyderidae based on special characters of wing venation and male genitalia. This new taxon broadens the diversity of Tanyderidae in the Cretaceous, and its morphological characters enhance our understanding of the development and evolution of the primitive crane flies.