Estimation of soil reinforcement by the roots of four post-dam prevailing grass species in the riparian zone of Three Gorges Reservoir,China

Soil erosion and bank degradation is a major post-dam concern regarding the riparian zone of the Three Gorges Reservoir. The development and succession of vegetation is a main countermeasure,especially to enhance bank stability and mitigate soil erosion by the root system. In this study, the roots of four prevailing grass species, namely, Cynodon dactylon, Hemarthria altissima, Hemarthria compressa, and Paspalum paspaloides, in the riparian zone were investigated in relation to additional soil cohesion. Roots were sampled using a single root auger. Root length density(RLD) and root area ratio(RAR) were measured by using the Win RHIZO image analysis system. Root tensile strength(TR) was performed using a manualdynamometer, and the soil reinforcement caused by the roots was estimated using the simple Wu's perpendicular model. Results showed that RLD values of the studied species ranged from 0.24 cm/cm3 to20.89 cm/cm3 at different soil layers, and RLD were significantly greater at 0–10 cm depth in comparison to the deeper soil layers(10 cm). RAR measurements revealed that on average 0.21% of the reference soil area was occupied by grass roots for all the investigated species. The measured root tensile strength was the highest for P. paspaloides(62.26MPa) followed by C. dactylon(51.49 MPa), H.compressa(50.66 MPa), and H. altissima(48.81MPa). Nevertheless, the estimated maximum root reinforcement in this investigation was 22.5 k Pa for H.altissima followed by H. compressa(21.1 k Pa), P.paspaloides(19.5 k Pa), and C. dactylon(15.4 k Pa) at0–5 cm depth soil layer. The root cohesion values estimated for all species were generally distributed at the 0–10 cm depth and decreased with the increment of soil depth. The higher root cohesion associated with H. altissima and H. compressa implies their suitability for revegetation purposes to strengthen the shallow soil in the riparian zone of the Three Gorges Reservoir. Although the soil reinforcement induced by roots is only assessed from indirect indicators, the present results still useful for species selection in the framework of implementing and future vegetation recovery actions in the riparian zone of the Three Gorges Reservoir and similar areas in the Yangtze River Basin.     

Soil anti-scourability enhanced by herbaceous species roots in a reservoir water level fluctuation zone

Revegetation is one of the successful approaches to soil consolidation and streambank protection in reservoir water level fluctuation zones(WLFZs).However,little research has been conducted to explore the impact of herbaceous species roots on soil anti-scourability during different growth stages and under different degrees of inundation in this zone.This study sampled two typical grasslands(Hemarthria compressa grassland and Xanthium sibiricum grassland)at two elevations(172 and 165 m a.s.l.)in the water level fluctuation zone(WLFZ)in the Three Gorges Reservoir(TGR)of China to quantify the changes in soil and root properties and their effects on soil anti-scourability.A simulated scouring experiment was conducted to test the soil anti-scourability in April and August of 2018.The results showed that the discrepancy in inundation duration and predominant herbaceous species was associated with a difference in root biomass between the two grasslands.The root weight density(RWD)values in the topsoil(0-10 cm)ranged from 7.31 to 13 mg cm^-3 for the Hemarthria compressa grassland,while smaller values ranging from 0.48 to 8.61 mg cm^-3 were observed for the Xanthium sibiricum grassland.In addition,the root biomass of the two herbs was significantly greater at 172 m a.s.l.than that at 165 m a.s.l.in the early recovery growth period(April).Both herbs can effectively improve the soil properties;the organic matter contents of the grasslands were 128.06%to 191.99%higher than that in the bare land(CK),while the increase in the water-stable aggregate ranged from 8.21%to 18.56%.Similarly,the topsoil antiscourability indices in both the herbaceous grasslands were larger than those in the CK.The correlation coefficients between the root length density(RLD),root surface area density(RSAD)and root volume density(RVD)of fine roots and the soil antiscourability index were 0.501,0.776 and 0.936,respectively.Moreover,the change in the soil antiscourability index was more sensitive to alternations in the RLD with diameters less than 0.5 mm.Overall,the present study showed that the perennial herbaceous(H.compressa)has great potential as a countermeasure to reduce or mitigate the impact of erosion in the WLFZ of the Three Gorges Reservoir.     

An exploratory analysis of vegetation strategies to reduce shallow landslide activity on loess hillslopes,Northeast Qinghai-Tibet Plateau,China

Heavy summer rainfall induces significant soil erosion and shallow landslide activity on the loess hillslopes of the Xining Basin at the northeast margin of the Qinghai-Tibet Plateau. This study examines the mechanical effects of five native shrubs that can be used to reduce shallow landslide activity. We measured single root tensile resistance and shear resistance, root anatomical structure and direct shear and triaxial shear for soil without roots and five root- soil composite systems. Results show that Atriplex canescens (Pursh) Nutt. possessed the strongest roots, followed by Caragana korshinskii Kom., Zygophyllum xanthoxylon (Bunge) Maxim., Nitraria tangutorum Bobr. and Lycium chinense Mill. Single root strength and shear resistance relationships with root diameter are characterized by power or exponential relations, consistent with the Mohr- Coulomb law. Root mechanical strength reflects their anatomical structure, especially the percentage of phloem and xylem cells, and the degree and speed of periderm lignifications. The cohesion force of root- soil composite systems is notably higher than that of soil without roots, with increasing amplitudes of cohesion force for A. canescens, C. korshinskii, Z. xanthoxylon, N. tangutorum and L. chinense of 75.9%, 75.1%, 36.2%, 24.6% and 17.0 % respectively. When subjected to shear forces, the soil without root samples show much greater lateral deformation thanthe root-soil composite systems, reflecting the restraining effects of roots. Findings from this paper indicate that efforts to reduce shallow landslides in this region by enhancing root reinforcement will be achieved most effectively using A. canescens and C. korshinskii.     

Mechanisms of root-soil reinforcement in bioembankments of sloping farmland in the purple hilly area,China

Bio-embankment is an important soil and water conservation measure in the purple hilly area in China,which can effectively improve the ability of cultivated soil layers to resist rainfall erosion and runoff scour.In contrast,the ecological effect of bioembankment depends on the stability of the earth bank.Taking the natural grass bank as a control(CK),the root distribution,root tensile properties and shear resistance of root-soil composites for 3 typical soil and water conservation bio-embankments,namely,Morus alba Linn(Morns),Zanthoxylum bungeanum Maxim(Zanthoxylum)and Medicago sativa(Medicago)were analysed.The results included the following:(1)The root system of the bio-embankments generally decreased in extent with the soil depth;fine roots in the o-io cm depth were most prevalent and significantly higher than those at the other depths,and coarse roots were mainly distributed in the o-30cm layer.(2)The stress-strain curves of the roots of each bio-embankment were single-peak curves without clear strain softening phenomena.The smaller the root diameters were,the smoother the stress-strain curves,and the lower the capability of the earth bank to resist collapse.The larger the root diameters were,the lower the tensile strength.The average root tensile force was highest for Zanthoxylum(73.91 N),followed by Medicago(68.07N)and Morus(61.88 N),and the average root tensile strength showed the same trend,16.52 MPa for Zanthoxylum,16.08 MPa for Medicago and 13.02MPa for Morus.(3)The bio-embankment measures significantly improved the soil shear resistance,especially under vertical loads of 1oo kPa and 200kPa.The soil internal friction angle showed a significant log-positive correlation with root morphological parameters of root length density(RLD),root surface area density(RSAD) and rootweight density(RWD),while the soil cohesion force showed a positive linear correlation with these parameters.The results provide effective parameters supporting for the design of bio-embankments and promoting the use of soil reinforcement with suitable species selection in protective earth banks for stability in the purple hilly area.     

Impacts of warming on root biomass allocation in alpine steppe on the north Tibetan Plateau

Biomass is an important component of global carbon cycling and is vulnerable to climate change. Previous studies have mainly focused on the responses of aboveground biomass and phenology to warming, while studies of root architecture and of root biomass allocation between coarse and fine roots have been scarcely reported in grassland ecosystems. We conducted an open-top-chamber warming experiment to investigate the effect of potential warming on root biomass and root allocation in alpine steppe on the north Tibetan Plateau. The results showed that Stipa purpurea had significantly higher total root length, root surface area and tips than Carex moocroftii. However,there were no differences in total root volume, mean diameter and forks for the two species. Warming significantly increased total root biomass(27.60%), root biomass at 0–10 cm depth(27.84%) and coarse root biomass(diameter 0.20 mm, 57.68%) in the growing season(August). However, warming had no significant influence on root biomass in the non-growing season(April). Root biomass showed clear seasonalvariations: total root biomass, root biomass at 0–10 cm depth and coarse root biomass significantly increased in the growing season. The increase in total root biomass was due to the enhancement of root biomass at 0–10 cm depth, to which the increase of coarse root biomass made a great contribution. This research is of significance for understanding biomass allocation, carbon cycling and biological adaptability in alpine grassland ecosystems under future climate change.     

The influence of plant root system architectural properties upon the stability of loess hillslopes,Northeast Qinghai,China

To investigate the influence of root system architectural properties of three indigenous (cold-adapted) shrubs on the hillslope stability of loess deposits in the Xining Basin, northeast part of Qinghai-Tibet Plateau (QTP), indoor direct shear tests have been conducted on the remolded rooted soil of three shrubs. Test results show that root system architectural indices (root area ratio (RAR), root length density (RLD) and root density (RD)) of the shrubs decline with depth and the relationship between RAR, RD and depth is exponential, while a power relationship describes the relationship between RLD and depth. The cohesion force of remolded rooted soil for the shrubs initially increases with depth, but it then demonstrates a slightly decreasing trend, which can be described with a power relationship. Power relationships also describe relationships between cohesion force and RAR, RLD and RD for the shrubs. As the growth period increases from 10 to 17 months, the incremental increase in RAR is 48.32% ~ 210.25% for Caragana korshinskii Kom and 0.56% ~ 166.85% for Zygophyllum xanthoxylon (Bunge) Maxim. This proportional increase is notably larger than that for RLD and RD. The increment in RAR is marginally greater for C. korshinskii than it is for Z. xanthoxylon. Correspondingly, the cohesion force incremental rates of remolded rooted soil for C. korshinskii and Z. xanthoxylon are 12.41% ~ 25.22% and 3.45% ~ 17.33% respectively. Meanwhile, as root content increases, the contribution by roots to cohesion force increases markedly until a threshold condition is reached.     

Influence of root suction on tensile strength of Chrysopogon zizanioides roots and its implication on bioslope stabilization

Root tensile strength is an important factor controlling the performance of bio-slope stabilization works. Due to evapotranspiration and climate factors, the root moisture content and its suction can vary seasonally in practice and may not equal soil suction. The influences of suction and root moisture contents were investigated on Chrysopogon zizanioides(vetiver grass) root tensile strength. The root specimens were equilibrated with moist air in different suction conditions(0, 10, 20, and 50 kPa), prior to root tension tests. The root-water characteristic curve or relationship between root moisture and suction, was determined. The increase in suction resulted in decreased tensile strengths of the grass roots, particularly those with diameter of about 0.2 mm, which constituted 50.7% of all roots. For 1 mm roots, the tensile strength appeared to be unaffected by suction increase. The average root tensile strengths were used to estimate the root cohesion in slope stability analysis to find variation of safety factors of a bioengineered slope in different suction conditions. The analysis showed that the critical condition of slope with the lowest factor of safety would happen when the soil suction was zero and the root suction was high. Such condition may occur during a heavy rain period after a prolonged drought.     

Comparison of artificial neural networks,geographically weighted regression and Cokriging methods for predicting the spatial distribution of soil macronutrients (N,P, and K)

Soil macronutrients(i.e. nitrogen(N), phosphorus(P), and potassium(K)) are important soils components and knowing the spatial distribution of these parameters are necessary at precision agriculture. The purpose of this study was to evaluate the feasibility of different methods such as artificial neural networks(ANN) and two geostatistical methods(geographically weighted regression(GWR) and cokriging(CK)) to estimate N, P and K contents. For this purpose, soil samples were taken from topsoil(0–30 cm) at 106 points and analyzed for their chemical and physical parameters. These data were divided into calibration(n = 84) and validation(n = 22). Chemical and physical variables including clay, p H and organic carbon(OC) were used as auxiliary soil variables to estimate the N, P and K contents. Results showed that the ANN model(with coefficient of determination R~2 = 0.922 and root mean square error RMSE = 0.0079%) was more accurate compared to the CK model(with R~2 = 0.612 and RMSE = 0.0094%), and the GWR model(with R~2 = 0.872 and RMSE = 0.0089%) to estimate the N variable. The ANN model estimated the P with the RMSE of 3.630 ppm, which was respectively 28.93% and 20.00% less than the RMSE of 4.680 ppm and 4.357 ppm from the CK and GWR models. The estimated K by CK, GWR and ANN models have the RMSE of 76.794 ppm, 75.790 ppm and 52.484 ppm. Results indicated that the performance of the CK model for estimation of macro nutrients(N, P and K) was slightly lower than the GWR model. Also, the accuracy of the ANN model was higher than CK and GWR models, which proved to be more effective and reliable methods for estimating macro nutrients.     

Soil Organic Carbon Contents and Stocks in Coastal Salt Marshes with <Emphasis Type="Italic">Spartina alterniflora</Emphasis> Following an Invasion Chronosequence in the Yellow River Delta,China

Plant invasion alters the fundamental structure and function of native ecosystems by affecting the biogeochemical pools and fluxes of materials and energy. Native (Suaeda salsa) and invasive (Spartina alterniflora) salt marshes were selected to study the effects of Spartina alterniflora invasion on soil organic carbon (SOC) contents and stocks in the Yellow River Delta. Results showed that the SOC contents (g/kg) and stocks (kg/m²) were significantly increased (P < 0.05) after Spartina alterniflora invasion of seven years, especially for the surface soil layer (0–20 cm). The SOC contents exhibited an even distribution along the soil profiles in native salt marshes, while the SOC contents were gradually decreased with depth after Spartina alterniflora invasion of seven years. The natural ln response ratios (LnRR) were applied to identify the effects of short-term Spartina alterniflora invasion on the SOC stocks. We also found that Spartina alterniflora invasion might cause soil organic carbon losses in a short-term phase (2–4 years in this study) due to the negative LnRR values, especially for 20–60 cm depth. And the SOCD in surface layer (0–20 cm) do not increase linearly with the invasive age. Spearman correlation analysis revealed that silt + clay content was exponentially related with SOC in surface layer (Adjusted R2 = 0.43, P < 0.001), suggesting that soil texture could play a key role in SOC sequestration of coastal salt marshes.     

Age-related changes of leaf traits and stoichiometry in an alpine shrub (<Emphasis Type="Italic">Rhododendron agglutinatum</Emphasis>) along altitudinal gradient

Leaf morphological and stoichiometric characteristics are considered to represent both the interior inheritable characters in the plant and its adaptations to specific exterior environments. Rhododendron agglutinatum, an evergreen alpine shrub species, occupies a wide range of habitats above timberline in the Miyaluo Natural Reserve, southwestern China. Along an altitudinal gradient ranging from 3700 to 4150 m, we measured leaf morphological characters including leaf dry matter content (LDMC), leaf dry mass per unit area (LMA), and one leaf area (OLA), as well as carbon (C) and nutrient (N, P) contents in leaves of three different age groups (juvenile leaves, mature leaves and senescent leaves). We also calculated the stoichiometric relationships among carbon and nutrients (C/N, C/P and N/P). Results showed that both age and altitude affected the leaf morphological and stoichiometric properties of R. agglutinatum. Mature leaves possessed the highest LDMC, LMA and C contents both on a dry mass basis and on a unit area basis. Younger leaves possessed higher contents of nutrients. OLA as well as ratios between carbon and nutrients (C/N, C/P) increased with ages. Juvenile leaves possessed lowest ratio between nitrogen and phosphorus. In juvenile leaves, nutrients increased with altitudinal elevation, whereas other traits decreased. In mature leaves, nutrients and their ratios with carbon showed consistent trends with juvenile leaves along increasing altitude, whereas LMA and carbon on a unit area basis showed opposite trends with juvenile leaves along increasing altitude. In senescent leaves, only content of phosphorus on a unit area basis and N/P were found linearly correlated with altitude. Our results demonstrated a clear pattern of nutrient distribution with aging process in leaves and indicated that a high possibility of N limitation in this region. We also concluded that younger leaves could be more sensitive to climate changes due to a greater altitudinal influence on the leaf traits in younger leaves than those in elder leaves.     

Distribution of sediment chloroplastic pigments in the Southern Yellow Sea,China

The distribution of sediment chloroplastic pigments (Chl-a, i.e. chlorophyll a and Pha-a, i.e. phaeophorbide a) in the Southern Yellow Sea of China was studied. Samples were collected from four cruises in January and June 2003, and January and June 2004. The results show that the vertical distribution of Chl-a and Pha-a in the sediment layers 0-2cm, 2-5cm and 5-8cm, follows a stable ratio, 5:3:2. The average ratio of Pha-a to Chl-a in sediment is 2.83. Spearman 2-tailed rank correlation analysis shows that Chl-a and Pha-a contents in each sediment layer have a highly significant correlation. The average contents of Chl-a and Pha-a in the sediment of the 0-8cm layer in the investigated area are 0.31 -0.47μgg-1 and 1.28-1.40 μgg-1 sediment (dry weight), respectively. The average Chl-a and Pha-a contents in sediment are higher in summer than in winter. ANOVA analysis shows that there is a highly significant variation among the Chl-a contents (P = 0.002 <0.01) of the four cruies, but this is not true for the case of Pha-a content (P = 0.766>0.05). The average Chl-a and Pha-a contents in the 2 sediment layers (0-2cm and 2-5cm) have significant or highly significant correlations with organic matter (OM), median diameter (Mdφ), silt plus clay percentage in the January 2003 cruise. In the June 2003 cruise, the average Chl-a content in the 3 sediment layers (0-2cm, 2-5cm, and 5-8cm) has a significant correlation with meiofauna biomass, and Pha-a content has highly significant correlations with water depth, bottom water temperature, OM and Mdφ The contents of Chl-a and Pha-a are lower than those in estuaries and intertidal areas, but close to those in the same area studied previously.     

章丘大葱特征元素农业地球化学研究

以章丘大葱产区生态地球化学调查为基础,采集了表层土壤、大葱、根系土及出露的代表性岩石样品,实测了大量和微量营养元素的含量,查明了研究区土壤地球化学背景。通过计算大葱对土壤营养元素的吸收率及大葱品质与营养元素的相关性分析,查明了大葱生长的特征营养元素为N,K2O,Zn,B,P,Cu,Mn。在岩石-土壤-大葱这一生态体系中,营养元素在土壤中保持一定的储备和积累,为大葱的生长提供了有利的农业地球化学环境。     

Influence of livestock soil eutrophication on floral composition in the Pyrenees mountains

Livestock behaviour in the Pyrenees includes free grazing and a long resting period that provokes the accumulation of dung and urine in some places,so-called camping areas. The aims of this study were （i） to analyze any change in floral composition,and in nutritional and chemical contents of plants in a livestock camping area; and （ii） to relate the floral composition with soil chemical properties. In a linear transect,five sampling zones were established,from the centre of the camping area to the surrounding Nardus stricta-dominant pasture. The above ground plant biomass and the topsoil were sampled in each zone with 6 replicates per zone. Plant species were classified and weighed to calculate above ground biomass,nutritional and chemical contents,and Shannon diversity and evenness indices. Additionally,soils were sampled in two periods,at the beginning and at the end of grazing period. Soil available nutrients （nitrate,ammonium,phosphorus,potassium,calcium and magnesium）,total nitrogen,organic carbon and pH were measured.
Plant chemical contents （protein,lignin and others） were significantly related to the proportions of grasses,legumes and other plants; so,the protein content is positively correlated with legumes plant biomass while lignin content is negatively correlated with grasses. Both plant and soil nutrients increased linearly towards the centre of the camping area. However,the relationship among plant species richness,diversity and evenness relative to its position along the studied transect was bell-shaped. From the outskirts to the centre of the camping area,plants with low nutrient demand were progressively replaced by those with medium and high nutrients demand and by pioneers.
Nardus stricta-dominant pasture has low plant diversity and plant nutrient content as well as a poor soil nutrient availability. The presence of the camping area introduced patches with more soil nutrients and new species in the large spatial scale. However at a small spatial scale,the strong soil nutrien     

Responses of <Emphasis Type="Italic">Dodonaea viscosa</Emphasis> growth and soil biological properties to nitrogen and phosphorus additions in Yuanmou dry-hot valley

Nitrogen (N) and phosphorus (P) are limited nutrients in terrestrial ecosystems, and their limitation patterns are being changed by the increase in N deposition. However, little information concerns the plant growth and the soil biological responses to N and P additions among different soils simultaneously, and these responses may contribute to understand plant-soil interaction and predict plant performance under global change. Thus, this study aimed to explore how N and P limitation changes in different soil types, and reveal the relationship between plant and soil biological responses to nutrient additions. We planted Dodonaea viscosa, a globally distributed species in three soil types (Lixisols, Regosols and Luvisols) in Yuanmou dry-hot valley in Southwest China and fertilized them factorially with N and P. The growth and biomass characters of D. viscosa, soil organic matter, available N, P contents and soil carbon (C), N, P-related enzyme activities were quantified. N addition promoted the growth and leaf N concentration of D. viscosa in Lixisols; N limitation in Lixisols was demonstrated by lower soil available N with higher urease activity. P addition promoted the growth and leaf P concentration of D. viscosa in Luvisols; severe P limitation in Luvisols was demonstrated by a higher soil available N: P ratio with higher phosphatase activity. Urease activity was negatively correlated with soil available N in Nlimited Lixisols, and phosphatase activity was negatively correlated with soil available P in P-limited Luvisols. Besides, the aboveground biomass and leaf N concentration of D. viscosa were positively correlated with soil available N in Lixisols, but the aboveground biomass was negatively correlated with soil available P. Our results show similar nutrient limitation patterns between plant and soil microorganism in the condition of enough C, and the nutrient limitations differ across soil types. With the continued N deposition, N limitation of the Lixisols in dry hot valleys is expected to be alleviated, while P limitation of the Luvisols in the mountaintop may be worse in the future, which should be considered when restoring vegetation.     

Ectomycorrhiza of <Emphasis Type="Italic">Larix sibirica</Emphasis> Ledeb. along the gradients of main ecological factors and elevation at the Northern and Subpolar Urals

Siberian larch (Larix sibirica Ledeb.) forests cover the largest areas in the Eurasian boreal zone, but there are insufficient data on its root system including the structure and functional traits of ectomycorrhizas (EM). The aim of this research is to find out if the morphological parameters of Larix sibirica EMs responded to the changes in elevation and main ecological factors (soil humidity, soil richness, soil acidity and habitat illumination). Using light microscopy, we studied EM diameter, root diameter, mantle width, and mantle volume share, share of tannin cells layers, EM density and EM length of Larix sibirica in two main types of plant communities along the elevation gradient at the Northern and Subpolar Urals. Differences in the environment were traced using phytoindication approach and the Ellenberg ecological scales. All the studied traits depend on the elevation and studied ecological factors. The diversity of fungal mantles is low, and the proportion of unstructured and pseudoparenchymatous mantles is high in response to the deterioration of the humidity, soil nitrogen content and acidity at higher-altitude habitats. Results of EM quantitative parameters measurements confirmed this pattern. We found a decline in the EM linear dimensions accompanied by a compensatory growth of the EM density with the raised elevation and the deterioration of environmental conditions.     

Relationships between river water quality and landscape factors in Haihe River Basin,China: Implications for environmental management

River water plays a key role in human health, and in social and economic development, and is often affected by both natural factors and human activities. An in-depth understanding of the role of these factors can help in developing an effective catchment management strategy to protect precious water resources. This study analyzed river water quality, patterns of terrestrial and riparian ecosystems, intensity of agricultural activities, industrial structure, and spatial distribution of pollutant emissions in the Haihe River Basin in China for the year of 2010, identifying the variables that have the greatest impact on river water quality. The area percentage of farmland in study area, the percentage of natural vegetation cover in the 1000-m riparian zone, rural population density, industrial Gross Domestic Product (GDP)/km², and industrial amino nitrogen emissions were all significantly correlated with river water quality (P < 0.05). Farming had the largest impact on river water quality, explaining 43.0% of the water quality variance, followed by the coverage of natural vegetation in the 1000-m riparian zone, which explained 36.2% of the water quality variance. Industrial amino nitrogen emissions intensity and rural population density explained 31.6% and 31.4% of the water quality variance, respectively, while industrial GDP/km² explained 26.6%. Together, these five indicators explained 67.3% of the total variance in water quality. Consequently, water environmental management of the Haihe River Basin should focus on adjusting agricultural activities, conserving riparian vegetation, and reducing industrial pollutant emissions by optimizing industrial structure. The results demonstrate how human activities drive the spatial pattern changes of river water quality, and they can provide reference for developing land use guidelines and for prioritizing management practices to maintain stream water quality in a large river basin.     

Lowland pasture in Himalayan highland: edaphic properties and species composition

This study was carried out to investigate if on-farm cut vs. grazed pastures differed in soil fertility and species density in the temperate Himalayan highlands. Soil fertility and species relative density were measured from 30 cut and 30 grazed dairy pastures. In both types of pasture, soil pH, available phosphorus (P) and exchangeable potassium (K) were negatively correlated with pasture age and slope. In cut pasture, cocksfoot (Dactylis glomerata) and pasture age were positively correlated, whereas in grazed pasture, they were negatively correlated. In grazed pasture, unsown species and pasture age were positively correlated. Soil available P was significantly greater in cut pastures whereas soil exchangeable K was significantly greater in grazed pastures. In terms of species density, cut pasture had greater densities of cocksfoot and Italian ryegrass (Lolium multiflorum), whereas grazed pasture showed greater densities of white clover, sedge and local grass. Our study suggests that, if there are no improvements in the current method of pasture management, the cut pastures in the future are likely to have a simple pasture mixture constituting only cocksfoot and white clover (Trifolium repens). Whereas in grazed pasture, the pasture mixture is likely to be comprised of white clover and unsown species such as sedge, local grass and broadleaf weeds.     

Variable hydrological effects of herbs and shrubs in the arid northeastern Qinghai-Tibet Plateau,China

This study aims to assess the hydrological effects of four herbs and four shrubs planted in a selfestablished test area in Xining Basin of northeastern Qinghai-Tibet Plateau, China. The RainfallIntercepting Capability(RIC) of the herbs and shrubs was evaluated in rainfall interception experiment at the end of the third, fourth and fifth month of the growth period in 2007. The leaf transpiration rate and the effects of roots on promoting soil moisture evaporation in these plants were also assessed in transpiration experiment and root-soil composite system evaporation experiment in the five month's growth period. It is found that the RIC of the fourstudied herbs follows the order of E. repens, E. dahuricus, A. trachycaulum and L. secalinus; the RIC of the four shrubs follows the order of A. canescens, Z. xanthoxylon, C. korshinskii and N. tangutorum. The RIC of all the herbs is related linearly to their mean height and canopy area(R~2 ≥ 0.9160). The RIC of all the shrubs bears a logarithmic relationship with their mean height(R~2 ≥ 0.9164), but a linear one with their canopy area(R~2 ≥ 0.9356). Moreover, different species show different transpiration rates. Of the four herbs, E. repens has the highest transpiration rate of 1.07 mg/(m~2·s), and of the four shrubs, A. canescens has the highest transpiration rate(0.74 mg/(m~2·s)). The roots of all the herbs and shrubs can promote soil moisture evaporation. Of the four herbs, the evaporation rate of E. repens root-soil composite system is the highest(2.14%), and of the four shrubs,the root-soil composite system of A. canescens has the highest evaporation rate(1.41%). The evaporation rate of the root-soil composite system of E. dahuricus and Z. xanthoxylon bears a second-power linear relationship with evaporation time(R~2 ≥ 0.9924). The moisture content of all the eight root-soil composite systems decreases exponentially with evaporation time(R~2 ≥ 0.8434). The evaporation rate and moisture content of all the plants' root-soil composite systems increases logarithmically(R~2 ≥ 0.9606) and linearly(R~2 ≥ 0.9777) with root volume density. The findings of this study indicate that among the four herbs and four shrubs, E. repens and A. canescens possess the most effective hydrological effects in reducing the soil erosion and shallow landslide in this region.     

Regional variation in carbon sequestration potential of forest ecosystems in China

Enhancing forest carbon(C) storage is recognized as one of the most economic and green approaches to offsetting anthropogenic CO_2 emissions. However, experimental evidence for C sequestration potential(C_(sp)) in China's forest ecosystems and its spatial patterns remain unclear, although a deep understanding is essential for policy-makers making decisions on reforestation. Here, we surveyed the literature from 2004 to 2014 to obtain C density data on forest ecosystems in China and used mature forests as a reference to explore C_(sp). The results showed that the C densities of vegetation and soil(0–100 cm) in China's forest ecosystems were about 69.23 Mg C/ha and 116.52 Mg C/ha, respectively. In mature forests, the C_(sp) of vegetation and soil are expected to increase to 129.26 Mg C/ha(87.1%) and 154.39 Mg C/ha(32.4%) in the coming decades, respectively. Moreover, the potential increase of C storage in vegetation(10.81 Pg C) is estimated at approximately twice that of soil(5.01 Pg C). Higher C_(sp) may occur in the subtropical humid regions and policy-makers should pay particular attention to the development of new reforestation strategies for these areas. In addition to soil nutrients and environment, climate was an important factor influencing the spatial patterns of C density in forest ecosystems in China. Interestingly, climate influenced the spatial patterns of vegetation and soil C density via different routes, having a positive effect on vegetation C density and a negative effect on soil C density. This estimation of the potential for increasing forest C storage provided new insights into the vital roles of China's forest ecosystems in future C sequestration. More importantly, our findings emphasize that climate constraints on forest C sequestration should be considered in reforestation strategies in China because the effects of climate were the opposite for spatial patterns of C density in vegetation and soil.Enhancing forest carbon(C) storage is recognized as one of the most economic and green approaches to offsetting anthropogenic CO2 emissions. However, experimental evidence for C sequestration potential(Csp) in China's forest ecosystems and its spatial patterns remain unclear, although a deep understanding is essential for policy-makers making decisions on reforestation. Here, we surveyed the literature from 2004 to 2014 to obtain C density data on forest ecosystems in China and used mature forests as a reference to explore Csp. The results showed that the C densities of vegetation and soil(0–100 cm) in China's forest ecosystems were about 69.23 Mg C/ha and 116.52 Mg C/ha, respectively. In mature forests, the Csp of vegetation and soil are expected to increase to 129.26 Mg C/ha(87.1%) and 154.39 Mg C/ha(32.4%) in the coming decades, respectively. Moreover, the potential increase of C storage in vegetation(10.81 Pg C) is estimated at approximately twice that of soil(5.01 Pg C). Higher Csp may occur in the subtropical humid regions and policy-makers should pay particular attention to the development of new reforestation strategies for these areas. In addition to soil nutrients and environment, climate was an important factor influencing the spatial patterns of C density in forest ecosystems in China. Interestingly, climate influenced the spatial patterns of vegetation and soil C density via different routes, having a positive effect on vegetation C density and a negative effect on soil C density. This estimation of the potential for increasing forest C storage provided new insights into the vital roles of China's forest ecosystems in future C sequestration. More importantly, our findings emphasize that climate constraints on forest C sequestration should be considered in reforestation strategies in China because the effects of climate were the opposite for spatial patterns of C density in vegetation and soil.     

Soil organic carbon pool along different altitudinal level in the Sygera Mountains,Tibetan Plateau

Labile organic carbon (LOC) is one of the most important indicators of soil organic matter quality and dynamics elevation and plays important function in the Tibetan Plateau climate. However, it is unknown what the sources and causes of LOC contamination are. In this study, soil organic carbon (SOC), total nitrogen (TN), microbial biomass carbon (MBC), microbial biomass nitrogen (MBN) and LOC were analyzed based on different soil horizons and elevations using turnover time in an experimental site (3700 m to 4300 m area) in Sygera. SOC and LOC in higher-elevation vegetation types were higher than that of in lower-elevation vegetation types. Our results presented that the soil microbial biomass carbon (SMBC) and soil microbial biomass nitrogen (SMBN) were positively correlated with SOC. The content of easily oxidized carbon (EOC), particulate organic carbon (POC) and light fraction organic carbon (LFOC) decreased with depth increasing and the content were the lowest in the 60 cm to 100 cm depth. The total SOC, ROC and POC contents decreased with increasing soil horizons. The SOC, TN, MBC and MBN contents increased with increasing altitude in the Sygera Mountains. The MBC and MBN contents were different with the changes of SOC (p<0.05), meanwhile, both LFOC and POC were related to total SOC (p<0.05). The physical and chemical properties of soil, including temperature, humidity, and altitude, were involved in the regulation of SOC, TN, MBC, MBN and LFOC contents in the Sygera Mountains, Tibetan Plateau.