Modeling the Impact of Black Spruce on the Fire Regime of Alaskan Boreal Forest

In the boreal biome, fire is the major disturbance agent affecting ecosystem change, and fire dynamics will likely change in response to climatic warming. We modified a spatially explicit model of Alaskan subarctic treeline dynamics (ALFRESCO) to simulate boreal vegetation dynamics in interior Alaska. The model is used to investigate the role of black spruce ecosystems in the fire regime of interior Alaska boreal forest. Model simulations revealed that vegetation shifts caused substantial changes to the fire regime. The number of fires and the total area burned increased as black spruce forest became an increasingly dominant component of the landscape. The most significant impact of adding black spruce to the model was an increase in the frequency and magnitude of large-scale burning events (i.e., time steps in which total area burned far exceeded the normal distribution of area burned). Early successional deciduous forest vegetation burned more frequently when black spruce was added to the model, considerably decreasing the fire return interval of deciduous vegetation. Ecosystem flammability accounted for the majority of the differences in the distribution of the average area burned. These simulated vegetation effects and fire regime dynamics have important implications for global models of vegetation dynamics and potential biotic feedbacks to regional climate.     

Modeling the Influence of Topographic Barriers on Treeline Advance at the Forest-Tundra Ecotone in Northwestern Alaska

The response of terrestrial ecosystems to climate warming has important implications to potential feedbacks to climate. The interactions between topography, climate, and disturbance could alter recruitment patterns to reduce or offset current predicted positive feedbacks to warming at high latitudes. In northern Alaska the Brooks Range poses a complex environmental and ecological barrier to species migration. We use a spatially explicit model (ALFRESCO) to simulate the transient response of subarctic vegetation to climatic warming in the Kobuk/Noatak River Valley (200 × 400 km) in northwest Alaska. The model simulations showed that a significantly warmer (+6 °C) summer climate would cause expansion of forest through the Brooks Range onto the currently treeless North Slope only after a period of 3000–4000 yr. Substantial forest establishment on the North Slope didnot occur until temperatures warmed 9 °C, and only following a 2000 yr time lag. The long time lags between change in climate and change in vegetation indicate current global change predictions greatly over-estimate the response of vegetation to a warming climate in Alaska. In all the simulations warming caused a steady increase in the proportion of early successional deciduous forest. This would reduce the magnitude of the predicted decrease in regional albedo and the positive feedback to climate warming. Simulation of spruce forest refugia on the North Slope showed forest could survive with only a 4 °C warming and would greatly reduce the time lag of forest expansion under warmer climates. Planting of spruce on the North Slope by humans could increase the likelihood of large-scale colonization of currently treeless tundra. Together, the long time lag and deciduous forest dominance would delay the predicted positive regional feedback of vegetation change to climatic warming. These simulated changes indicate the Brooks Range would significantly constrain regional forest expansion under a warming climate, with similar implications for other regions possessing major east-west oriented mountain ranges.     

Modeling interactive effects of climate change,air pollution,and fire on a California Shrubland

A computer simulation model (FINICS) was used to project the interactive effects of CO₂-induced climate change on a drought-deciduous shrubland. FINICS simulates the competitive relations of five dominant shrub species of Californian coastal sage scrub, based on their aboveground growth and reproductive behavior. The model was used to simulate the separate and combined effects of altered precipitation, temperature, ambient ozone levels, and fuel loads and fire intensity, on species composition. Both growth chamber and field data were used to parameterize the model. Projections show that changes attributed to climate variation alone were markedly accentuated when the indirect effects of climate change on fire intensity and ozone pollution were considered. Model results emphasized that change in community composition will result from shifting competitive abilities of individual species under the changed environmental conditions. While neither all of the secondary effects of climate change nor all possible species were included in the model, the projections suggest that inclusion of secondary interactions and species competition will be important in predicting vegetation change realistically.     

Simulating forest dynamics in a complex topography using gridded climatic data

The forest model ForClim was used to evaluate the applicability of gap models in complex topography when the climatic input data is provided by a global database of 0.5° resolution. The analysis was based on 12 grid cells along an altitudinal gradient in the European Alps. Forest dynamics were studied both under current climate as well as under four prescribed 2 × CO₂ scenarios of climatic change obtained from General Circulation Models, which allowed to assess the sensitivity of mountainous forests to climatic change.Under current climate, ForClim produces plausible patterns of species composition in space and time, although the results for single grid cells sometimes are not representative of reality due to the limited precision of the climatic input data.Under the scenarios of climatic change, three responses of the vegetation are observed, i.e., afforestation, gradual changes of the species composition, and dieback of today's forest. In some cases widely differing species compositions are obtained depending on the climate scenario used, suggesting that mountainous forests are quite sensitive to climatic change. Some of the new forests have analogs on the modern landscape, but in other cases non-analog communities are formed, pointing at the importance of the individualistic response of species to climate.The applicability of gap models on a regular grid in a complex topography is discussed. It is concluded that for their application on a continental scale, it would be desirable to replace the species in the models by plant functional types. It is suggested that simulation studies like the present one must not be interpreted as predictions of the future fate of forests, but as means to assess their sensitivity to climatic change.     

Taking the Pulse of Mountains: Ecosystem Responses to Climatic Variability

An integrated program of ecosystem modeling and field studies in the mountains of the Pacific Northwest (U.S.A.) has quantified many of the ecological processes affected by climatic variability. Paleoecological and contemporary ecological data in forest ecosystems provided model parameterization and validation at broad spatial and temporal scales for tree growth, tree regeneration and treeline movement. For subalpine tree species, winter precipitation has a strong negative correlation with growth; this relationship is stronger at higher elevations and west-side sites (which have more precipitation). Temperature affects tree growth at some locations with respect to length of growing season (spring) and severity of drought at drier sites (summer). Furthermore, variable but predictable climate-growth relationships across elevation gradients suggest that tree species respond differently to climate at different locations, making a uniform response of these species to future climatic change unlikely. Multi-decadal variability in climate also affects ecosystem processes. Mountain hemlock growth at high-elevation sites is negatively correlated with winter snow depth and positively correlated with the winter Pacific Decadal Oscillation (PDO) index. At low elevations, the reverse is true. Glacier mass balance and fire severity are also linked to PDO. Rapid establishment of trees in subalpine ecosystems during this century is increasing forest cover and reducing meadow cover at many subalpine locations in the western U.S.A. and precipitation (snow depth) is a critical variable regulating conifer expansion. Lastly, modeling potential future ecosystem conditions suggests that increased climatic variability will result in increasing forest fire size and frequency, and reduced net primary productivity in drier, east-side forest ecosystems. As additional empirical data and modeling output become available, we will improve our ability to predict the effects of climatic change across a broad range of climates and mountain ecosystems in the northwestern U.S.A.     

How Much Physiology is Needed in Forest Gap Models for Simulating Long-Term Vegetation Response to Global Change? Challenges, Limitations, and Potentials

Gap models were introduced over 30 years ago to examine the dynamics of forestvegetation structure and species composition based on plant populationdynamics. While there have been many advances, gap models remain heavilycriticized for their lack of attention to physiology, particularly as itaffects their ability to simulate forest response to elevated CO₂concentration and climatic change. In this paper we provide a summary andsynthesis of the results from the Gap Model Comparison Workshop that was heldin July, 1999 at Pingree Park, Colorado (U.S.A.). We identify some grandchallenges for the future development of forest gap models and discusslimitations as well as potentials of this modeling approach.     

Summer maximum temperature in northern France over the past century: instrumental data versus multiple proxies (tree-ring isotopes, grape harvest dates and forest fires)

Changes in maximum spring and summer temperature are expected to have impacts on plant phenology and the occurrence of forest fires. Homogenised instrumental records of maximum spring and summer temperature are available in northern France for the past century, as well as documentary records of grape harvest dates and forest fire frequencies. Here we provide a new proxy of seasonal climate obtained by the analysis of latewood tree ring cellulose isotopic composition (δ¹⁸O, δ¹³C and δD), from 15 living oak trees (Quercus petraea) sampled in the Fontainebleau forest, near Paris. For the past 30 years, we have conducted a study on the inter-tree (for oxygen isotopes) and inter-station (for oxygen and hydrogen) isotopic variability. Multiple linear regression statistical analyses are used to assess the response function of documentary and tree-ring isotopic records to a variety of climatic and hydrological parameters. This calibration study highlights the correlation between latewood tree-ring δ¹⁸O and δ¹³C, grape harvest dates and numbers of forest fire starts with maximum growing season (April to September) temperature, showing the potential of multiple proxy reconstructions to assess the past fluctuations of this parameter prior to the instrumental period.     

Recent climate change: Rethinking drought in the context of Forest Fire Research in Ticino, South of Switzerland

Summary We analyse trends over the past 32 years in drought variables in the context of forest fire research and climate change in Ticino, Southern Switzerland. January to April and in particular the month of March have become drier. This is the time of the year when forest fires are most frequent. Our results show an increasing trend in all climatic variables favourable to drought and forest fires, including the length of episodes without precipitation, sunshine duration and temperature, and a decrease in relative humidity. The only exception is the number of “foehn” days, which have not significantly increased.     

Study on forest fire danger over Northern China during the recent 50?years

Daily meteorological data at 263 stations in northern China from 1956 to 2005 were used to calculate various forest fire danger weather (FFDW) indices, such as Nesterov Index (NI), Modified Nesterov Index (MNI), Keetch-Byram Drought Index (KBDI), and Forest Fire Danger Index (FFDI), at different time scales. The relationship between each index and forest fire was analyzed. MNI and FFDI were then selected to study the impact on forest fire danger due to climatic change in northern China in the recent 50?years. Results show that forest fire danger has significantly increased in Northeast China where there is the richest forest resource in China, and also increased in North China. However, it has not changed much in eastern part of Northwest China, and the forest fire danger has even significantly decreased in northern Xinjiang. Significant rise of forest fire danger in Northeast China mainly results from the co-effect of increase of temperature, and decreases of humidity and precipitation. Relative humidity change dominants forest fire danger trends in the four forest regions in northern China.     

Potential response of pacific northwestern forests to climatic change,effects of stand age and initial composition

We used an individual-based forest simulator (a gap model) to assess the potential effects of anthropogenic climatic change on conifer forests of the Pacific Northwestern United States. Steady-state simulations suggested that forest zones could be shifted on the order of 500–1000 m in elevation, which could lead to the local extirpation of some high-altitude species. For low-elevation sites, species which currently are more abundant hundreds of kilometers to the south would be favored under greenhouse scenarios. Simulations of transient responses suggested that forest stands could show complex responses depending on initial species composition, stand age and canopy development, and the magnitude and duration of climatic warming. Assumptions about species response to temperature, which are crucial to the model's behaviors, were evaluated using data on species temperature limits inferred from regional distributions. The high level of within-species variability in these data, and other confounding factors influencing species distributions, argue against over-interpreting simulations. We suggest how we might resolve critical uncertainties with further research.     

Social and biophysical determinants of future forest conditions in New England: Effects of a modern land-use regime

The future forests of eastern North America will be shaped by at least three broad drivers: (i) vegetation change and natural disturbance patterns associated with the protracted recovery following colonial era land use, (ii) a changing climate, and (iii) a land-use regime that consists of geographically variable rates and intensities of forest harvesting, clearing for development, and land protection. We evaluated the aggregate and relative importance of these factors for the future forests of New England, USA by simulating a continuation of the recent trends in these drivers for fifty-years, nominally spanning 2010 to 2060. The models explicitly incorporate the modern distribution of tree species and the geographical variation in climate and land-use change. Using a cellular land-cover change model in combination with a physiologically-based forest landscape model, we conducted a factorial simulation experiment to assess changes in aboveground carbon (AGC) and forest composition. In the control scenario that simulates a hypothetical absence of any future land use or future climate change, the simulated landscape experienced large increases in average AGC—an increase of 53% from 2010 to 2060 (from 4.2 to 6.3 kg m⁻²). By 2060, climate change increased AGC stores by 8% relative to the control while the land-use regime reduced AGC by 16%. Among land uses, timber harvesting had a larger effect on AGC storage and changes in tree composition than did forest conversion to non-forest uses, with the most pronounced impacts observed on private corporate-owned land in northern New England. Our results demonstrate a large difference between the landscape’s potential to store carbon and the landscape’s current trajectory, assuming a continuation of the modern land-use regime. They also reveal aspects of the land-use regime that will have a disproportionate impact on the ability of the landscape to store carbon in the future, such as harvest regimes on corporate-owned lands. This information will help policy-makers and land managers evaluate trade-offs between commodity production and mitigating climate change through forest carbon storage.     

Forest ecosystems, disturbance, and climatic change in Washington State, USA

Climatic change is likely to affect Pacific Northwest (PNW) forests in several important ways. In this paper, we address the role of climate in four forest ecosystem processes and project the effects of future climatic change on these processes across Washington State. First, we relate Douglas-fir growth to climatic limitation and suggest that where Douglas-fir is currently water-limited, growth is likely to decline due to increased summer water deficit. Second, we use existing analyses of climatic controls on tree species biogeography to demonstrate that by the mid twenty-first century, climate will be less suitable for key species in some areas of Washington. Third, we examine the relationships between climate and the area burned by fire and project climatically driven regional and sub-regional increases in area burned. Fourth, we suggest that climatic change influences mountain pine beetle (MPB) outbreaks by increasing host-tree vulnerability and by shifting the region of climate suitability upward in elevation. The increased rates of disturbance by fire and mountain pine beetle are likely to be more significant agents of changes in forests in the twenty-first century than species turnover or declines in productivity, suggesting that understanding future disturbance regimes is critical for successful adaptation to climate change.     

Modeling the Effects of Global Climatic Change at the Ecotone of Boreal Larch Forest and Temperate Forest in Northeast China

The dynamics of the forest at the ecotone of the boreal forest and temperate forest in Northeast China were simulated using the adapted gap model BKPF under global climatic change (GFDL scenario) and doubled CO₂ concentrations at 50 years in the future. The response of tree species and species with similar biological characteristics under global climate change and double CO₂ concentrations were based on biophysical limits of the tree species in the area and their biological competition. The results showed that after 50 years the stand density and LAI (leaf area index) of the forest growing from a clear-cut would not be significantly different from those under current conditions. Stand productivity would increase about 7%, and stand aboveground biomass would increase 15%. However, the stand density of the current mature forest would be reduced by more than 20%. The stand would be dominated by Quercus mongolica Fisch., Populus davidiana Dode., Betula spp. and other broadleaved tree species, and Quercus mongolica would account for about 50% of the total density. The stand biomass would be reduced by more than 90%. Quercus mongolica would comprise about 57% of the total stand biomass. The stand productivity would not change significantly, but it would be comprised mainly of Quercus mongolica, Populus davidiana, Betula spp. The current stand height would decrease slightly. The stand LAI would decline dramatically, moreover, Quercus mongolica would comprise about 50% of the stand LAI.     

Estimating australian forest fire danger under conditions of doubled carbon dioxide concentrations

The most appropriate indices with which to quantify Australian bushfire danger are the McArthur fire danger meters. These meters use meteorological information to produce a fire danger index that is directly related to the chance of a fire starting - and to the severity of a fire once it has started. The Mark 5 forest-fire danger meter uses air temperature, relative humidity and wind speed, plus a drought factor that is calculated using daily rainfall and temperature information.Three years of daily data generated from the CSIRO four-level general circulation model, and thirty years of daily data generated from the CSIRO nine-level model were used to estimate the daily McArthur forest fire danger index for simulations corresponding to present conditions, and to those corresponding to doubled atmospheric CO₂. The performance of these models with respect to fire danger was tested by comparing the fire danger index for Sale (in the Eastern part of Victoria, South-eastern Australia) calculated from analysis of daily climatological data with the modelled annual cumulative forest fire danger index for the grid point that was representative of Sale. Data from both models for all Australian grid points were also examined. Both models predict an increase in fire danger over much of Australia for their doubled CO₂ scenarios.The results from the models confirm that annually averaged daily relative humidity is the single most important variable in the estimation of forest fire danger on an annual basis, yet the models tend to produce relative humidities that are slightly too low so that the fire danger is overestimated. A simple one-box model of evaporation indicates that the value of relative humidity to be expected under an altered climatic regime depends on the modelled relation between actual and potential evaporation, the present values of relative humidity and evaporation rate, as well as on the expected changes in wind speed.     

Fire activity in Borneo driven by industrial land conversion and drought during El Niño periods, 1982–2010

Tropical rainforests, naturally resistant to fire when intact, are increasingly vulnerable to burning due to ongoing forest perturbation and, possibly, climatic changes. Industrial-scale forest degradation and conversion are increasing fire occurrence, and interactions with climate anomalies such as El Niño induced droughts can magnify the extent and severity of fire activity. The influences of these factors on fire frequency in tropical forests has not been widely studied at large spatio-temporal scales at which feedbacks between fire reoccurrence and forest degradation may develop. Linkages between fire activity, industrial land use, and El Niño rainfall deficits are acute in Borneo, where the greatest tropical fire events in recorded history have apparently occurred in recent decades. Here we investigate how fire frequency in Borneo has been influenced by industrial-scale agricultural development and logging during El Niño periods by integrating long-term satellite observations between 1982 and 2010 – a period encompassing the onset, development, and consolidation of its Borneo’s industrial forestry and agricultural operations as well as the full diversity of El Niño events. We record changes in fire frequency over this period by deriving the longest and most comprehensive spatio-temporal record of fire activity across Borneo using AVHRR Global Area Coverage (GAC) satellite data. Monthly fire frequency was derived from these data and modelled at 0.04° resolution via a random-forest model, which explained 56% of the monthly variation as a function of oil palm and timber plantation extent and proximity, logging intensity and proximity, human settlement, climate, forest and peatland condition, and time, observed using Landsat and similar satellite data. Oil-palm extent increased fire frequency until covering 20% of a grid cell, signalling the significant influence of early stages of plantation establishment. Heighted fire frequency was particularly acute within 10 km of oil palm, where both expanding plantation and smallholder agriculture are believed to be contributing factors. Fire frequency increased abruptly and dramatically when rainfall fell below 200 mm month⁻¹, especially as landscape perturbation increased (indicated by vegetation index data). Logging intensity had a negligible influence on fire frequency, including on peatlands, suggesting a more complex response of logged forest to burning than appreciated. Over time, the epicentres of high-frequency fires expanded from East Kalimantan (1980’s) to Central and West Kalimantan (1990’s), coincidentally but apparently slightly preceding oil-palm expansion, and high-frequency fires then waned in East Kalimantan and occurred only in Central and West Kalimantan (2000’s). After accounting for land-cover changes and climate, our model under-estimates observed fire frequency during ca. 1990–2002 and over-estimates it thereafter, suggesting that a multi-decadal shift to industrial forest conversion and forest landscapes may have diminished the propensity for high-frequency fires in much of this globally significant tropical region since ca. 2000.     

Aboveground Growth and Competition in Forest Gap Models: An Analysis for Studies of Climatic Change

Gap models have been used extensively in ecological studies of forest structure and succession, and they should be useful tools for studying potential responses of forests to climatic change. There is a wide variety of gap models with different degrees of physiological detail, and the manner in which the effects of climatic factors are analyzed varies across that range of detail. Here we consider how well the current suite of gap models can accommodate climatic-change issues, and we suggest what physiological attributes and responses should be added to better represent responses of aboveground growth and competition. Whether a gap model is based on highly empirical, aggregated growth functions or more mechanistic expressions of carbon uptake and allocation, the greatest challenge will be to express allocation correctly. For example, incorporating effects of elevated CO₂ requires that the fixed allometry between stem volume and leaf area be made flexible. Simulation of the effects of climatic warming should incorporate the possibility of a longer growing season and acclimation of growth processes to changing temperature. To accommodate climatic-change factors, some of the simplicity of gap models must be sacrificed by increasing the amount of physiological detail, but it is important that the capability of the models to predict competition and successional dynamics not be sacrificed.     

Relationship between projected changes in future climatic suitability and demographic and functional traits of forest tree species in Spain

The response of plant species to future climate conditions is probably dependent on their ecological characteristics, including climatic niche, demographic rates and functional traits. Using forest inventory data from 27 dominant woody species in Spanish forests, we explore the relationships between species characteristics and projected changes in their average climatic suitability (occurrence of suitable climatic conditions for a species in a given territory) obtained by empirical niche-based models, under a business-as-usual climate change scenario (A1, HadCM3, 2001–2100). We hypothesize that most species will suffer a decline in climatic suitability, with a less severe for species (i) currently living in more arid climates or exhibiting a broader current climatic niche; (ii) with higher current growth rates; (iii) with functional traits related to resistance to water deficits. The analysis confirm our hypothesis since apart from a few Mediterranean species, most species decrease their climatic suitability in the region under future climate, characterized by increased aridity. Also, species living in warmer locations or under a wider range of climatic conditions tend to experience less decrease in climatic suitability. As hypothesized, a positive relationship was detected between current relative growth rates and increase in future climatic suitability. Nevertheless, current tree mortality did not correlate with changes in future climatic suitability. In contrast with our hypothesis, functional traits did not show a clear relationship with changes in climate suitability; instead species often presented idiosyncratic responses that, in some cases, could reflect past management. These results suggest that the extrapolation of species performance to future climatic scenarios based on current patterns of dominance is constrained by factors other than species autoecology, particularly human activity.     

内蒙古大兴安岭林区雷击火灾气候成因分析

利用常规天气图、数值预报产品、卫星云图以及溃变理论的预报工具V-3θ图,对2005年6月30日至7月2日发生在青藏高原东北侧甘肃省区域性持续暴雨天气过程进行了诊断应用综合分析.结果表明:副热带高压西伸北抬外围西南风气流控制青藏高原东北侧,当东北低涡、西风带的冷空气与西南风交汇时,触发强对流;850～200hPa有深厚的水汽层;700 hPa稳定的低涡切变为暴雨提供了强烈持续的辐合上升运动;卫星云图表明持续性暴雨由多个相继生消的中尺度对流系统影响造成的.基于溃变理论的预报方法在西北区域性持续暴雨的起报、结束及落区有很好的预测能力.     

Faster growth in warmer winters for large trees in a Mediterranean-climate ecosystem

Large trees (>76 cm breast-height diameter) are vital components of Sierra Nevada/Cascades mixed-conifer ecosystems because of their fire resistance, ability to sequester large amounts of carbon, and role as preferred habitat for sensitive species such as the California spotted owl. To investigate the likely performance of large trees in a rapidly changing climate, we analyzed growth rings of five conifer species against 20th century climate trends from local weather stations. Over the local station period of record, there were no temporal trends in precipitation, but maximum temperatures increased by 0.10 to 0.13 °C/decade (summer and autumn), and minimum temperatures increased by 0.11 to 0.19 °C/decade in all seasons. All species responded positively to precipitation, but more variation was explained by a significant positive response to minimum winter temperatures. High maximum summer temperature adversely affected growth of two species, and maximum spring temperatures in the year prior to ring formation were negatively associated with growth of one species. The strong coherent response to increasing minimum temperatures bodes well for growth of large trees in Sierra/Cascades region mixed conifer forest under continued climatic warming, but these trees will still be under threat by the increased fire intensity that is a indirect effect of warming.     

Climate change and the incidence of a forest pest in Mediterranean ecosystems: can the North Atlantic Oscillation be used as a predictor?

Many forest pest species strongly depend on temperature in their population dynamics, so that rising temperatures worldwide as a consequence of climatic change are leading to increased frequencies and intensities of insect-pest outbreaks. In the Mediterranean area, the climatic conditions are strongly linked to the effects of the North Atlantic Oscillation (NAO). The aim of this work is to analyze the dynamics of the pine processionary moth (Thaumetopoea pityocampa), a severe pest of Pinus species in the Circunmediterranean, throughout a region of southern Spain, in relation to NAO indices. We related the percentage of forest plots with high defoliation by pine processionary moth each year with NAO values for the present and the three previous winters, using generalized linear models with a binomial error distribution. The time series is 16-year long, and we performed analyses for the whole database and for the five main pine species separately. We found a consistent relationship between the response variable and the NAO index. The relationship is stronger with pine species living at medium-high altitudes, such as Aleppo (P. halepensis), black (P. nigra), and Scots (Pinus sylvestris) pine, which show the higher defoliation intensities up to 3?years after a negative NAO phase. The results highlight, for the first time, the usefulness of using global drivers in order to understand the dynamics of pest outbreaks at a regional scale, and they open the window to the development of NAO-based predictive models as an early-warning signal of severe pest outbreaks.