Ichnology of the Goradongar Formation,Goradongar hill range,Patcham island,Kachchh, Western India

The middle Jurassic Goradongar Formation exposed in Goradongar hill represents a mixed siliciclasticcarbonate succession with shales and limestones. They contain a large number of well preserved trace fossils. Total 44 ichnospecies of 31 ichnogenera; representing diverse ethology, were grouped in five ichnoassemblages (Planolites, Palaeophycus, Gyrochorte, Rhizocorallium and Arenicolites assemblage). These recurring ichnoassemblages represent the Cruziana ichnofacies and occasionally a mixed Skolithos-Cruziana ichnofacies. Patterns of diversity and density of the trace fossils reveal changes in bathymetry, oxygen level, trophic level and the sub-strate conditions at the time of deposition. These paleoenvironment and palaeo-oceanography changes are co-relatable to world-wide Bathonian-Callovian (middle Jurassic) deposits.     

Cambrian trace fossils of the Cruziana ichnofacies from the Bikaner-Nagaur Basin,north western Indian Craton

The Marwar Supergroup of the Bikaner-Nagaur Basin is composed of sediments deposited from the late Neoproterozoic (Ediacaran) to Upper Cambrian. The Nagaur Sandstone Formation of the Nagaur Group (uppermost division of the Marwar Supergroup) preserves trace fossils significant for establishing Early Cambrian biostratigraphic zones and depositional facies. Fifteen ichnospecies (and eight ichnogenera) identified in the Nagaur Sandstone Formation include “Treptichnus” pedum, Cruziana cf. tenella, Cruziana isp., Diplichnites ispp. A, B, and C, Gyrophyllites isp., Lockeia isp., Merostomichnites isp., Monomorphichnus gregarius isp. nov., Monomorphichnus isp., Planolites isp., Psammichnites isp., Rusophycus bikanerus isp. nov., Rusophycus cf. carbonarius, Rusophycus isp. and radial trace fossils.These trace fossils belong to ethological categories pascichnia, repichnia, cubichnia, and fodinichnia and represent arthropod and worm-like burrowing biota. The assemblage and a regional comparison with contemporaneous trace fossils in the eastern Gondwanan realm suggest that the sequence in the study area belongs to the Cruziana tenella Ichnozone and to Stage 2 (upper part of Terreneuvian), however the Middle Cambrian is not excluded. The trace fossil assemblage belongs to the archetypal Cruziana ichnofacies. Cross bedded sandstone, mud cracks and rainprints in the ichniferous strata of the Nagaur Sandstone Formation indicate deposition in an intertidal sand flat with channels that was exposed episodically.     

Balanoglossites ichnofabrics from the Middle Ordovician Volkhov formation (St. Petersburg Region, Russia)

The limestone succession of the Middle Ordovician Volkhov Formation in the St. Petersburg region (Russia) exposes numerous horizons with firm- and hardgrounds below omission surfaces, which contain trace fossils attributable to the ichnogenus Balanoglossites Mägdefrau, 1932. Although known from the literature since 75 years, such trace fossils were previously only informally described as “Korrosionsgruben”, “Karandashi”, or ascribed to different ichnotaxa such as Trypanites, Arenicolites and Pseudopolydorites. Owing to their complexity and often high bioturbation density, the morphology of these trace fossils is difficult to capture. Ichnofabrics containing Balanoglossites triadicus were studied in detail from sawn rock faces, broken rock blocks and sectioned slabs, including those from historical buildings in St. Petersburg. Accordingly, different trace-fossil elements can be revealed in dependence on the original substrate consistency, reflecting various stages of lithification: Mineral-stained and Trypanites-perforated hardground surfaces are bioeroded with long elongated grooves which are assigned to the ichnogenus Sulcichnus. Subtle openings lead into the partly lithified limestone where they branch into complex galleries of B. triadicus. They are characterized by J-, U- and Y-shaped shafts and multiply branched tunnels, which gradually continue into the underlying firmground. Other portions of the ichnofabric only exhibit biodeformational structures or the strongly compacted and branched burrow networks Labyrintichnus, which is due to the original soft sediment consistency. Balanoglossites ichnofabrics demarcate certain omission surfaces within the Dikari Limestone and can be traced for more than 300 km, supporting the regional lithostratigraphical correlation. The trace maker of B. triadicus and related trace fossils is interpreted to be a eunicid polychate with the ability to bioerode and burrow the sediment. The studied material from the Ordovician is similar to the Balanoglossites from the type area, the Triassic of Germany, in many respects. B. triadicus is a very common trace fossil in Ordovician and other Palaeozoic rocks of Baltoscandia and North America but so far has been seldom identified as such but instead is commonly confused with Thalassinoides, from which, however, it differs in several aspects.     

Oxfordian plug-shaped trace fossils from northwestern Algeria

Plug-shaped trace fossils B. perata, B. hemispherica, Bergaueria isp., Calycraterion samsonowiczi, Conostichus ornatus and Conostichus isp. are found in the middle-upper Oxfordian fine-grained sandstone of the Argiles de Saïda Formation (northwestern Algeria). Except for the trace Calycraterion which is interpreted as an annelid trace, these traces correspond to distinct behaviours of sea anemones, and their occurrence within tempestite deposits is influenced by a stress factors in a storm-dominated environment. The abundance of these plug-shaped trace fossils suggests that the upper Jurassic siliciclastic shelves with fine-grained soft clastic substrate were colonised by actinarian sessile benthos.     

Zoophycos and related trace fossils from the Chefar El Ahmar Formation,Upper Emsian-Frasnian Ia-Ib (Ougarta,SW Algeria)

The Upper Emsian to Frasnian Ia-Ib strata of the Marhouma area (or “km 30” outcrop), exposed in the Ougarta Range (SW Algeria) belong to the Chefar El Ahmar Formation. On the basis of distinct lithological and palaeontological features, this formation is subdivided into three members (Lower Marly Limestones Member, Middle Marly Limestones Member, and Upper Marly Limestones Member). The studied beds show low to moderate diversity of trace fossil assemblage which contains thirteen ichnotaxa: Chondrites intricatus, Chondrites isp., Chondrites cf. targionii, Circulichnis cf. montanus, Cochlichnus isp., Neonereites biserialis, Neonereites multiserialis, Nereites isp., Palaeophycus isp., Planolites isp., Thalassinoides isp., Zoophycos aff. cauda-galli, and Zoophycos isp. A. The two latter ichnotaxa are the most common trace fossils in the assemblage and occur at three different levels showing different bioturbation intensities. The first Zoophycos-bearing level (Zl 1) is characterised by an overall high bioturbation intensity reflecting a very high oxygenation rate and nutrient supply, allowing the development of large and dense Zoophycos specimens. The second Zoophycos-bearing level (Zl 2) has a considerable reduction of bioturbation intensity as compared to the previous level, with an abundance of Chondrites, which is probably due to radical palaeoecological changes that suggests dysoxic and stressful conditions. The third Zoophycos-bearing level (Zl 3) is characterised by an overall moderate bioturbation intensity. The distribution of trace fossils was influenced by lithology, sedimentation rate, energy level (storm events), bottom oxygenation, and nutrient supply. The lithofacies and trace fossils of the Chefar El Ahmar Formation both indicate a depositional environment fluctuating from the lower shoreface to lower offshore zone.     

重庆綦江白垩系夹关组遗迹化石及其沉积环境

遗迹化石大都是原地埋藏,研究遗迹化石在古环境、古气候、古海洋和古生态等方面的重建具有重要的意义。綦江国家地质公园老瀛山园区夹关组地层中已发现的无脊椎动物遗迹化石经鉴定包括1大类3属3种,分别为觅食迹Scoyenia gracilis(纤细斯柯茵迹)、Beconites antarcticus(南极似塔形迹)和Planolites beverleyensis(贝弗利漫游迹)。这些遗迹化石都属于Scoyenia遗迹相,代表了研究区在白垩纪夹关期为间断性暴露地表的河漫滩或者滨浅湖环境。     

Preservation of borings: contrasting examples from the type Maastrichtian (Upper Cretaceous), southern Limburg,the Netherlands

The taphonomy of trace fossils and their substrates remains an understudied facet of sedimentary geology. Contrary to common prejudice, trace fossils are not invariably preserved in situ, but may be exhumed and reworked following lithification. The trace fossils most commonly found ex situ are borings in mobile shelly substrates. Two notable, but contrasting, examples of post-mortem transport of borings are described from the Maastrichtian (Upper Cretaceous) of southern Limburg, the Netherlands. A long, unusually straight and complete calcareous tube assigned to Teredolites longissimus Kelly and Bromley is an organically secreted internal mould, produced by a teredinid or pholadid bivalve boring in wood and lining their tube. Strictly, this is part of the body fossil of the producing bivalve, but it is also an organically generated internal mould of the boring. A flint steinkern of a right valve of Crassatella bosquetiana d’Orbigny preserves a suite of silicified borings. Caulostrepsis taeniola Clarke is a U-shaped boring with a vane connecting the parallel limbs. Talpina isp. is a slender, simple, branched tunnel. Most unexpected, Spirichnus spiralis Fürsich et al. is a spiral ‘worm’ boring hitherto only known from the Upper Jurassic. This stratigraphic gap is likely an artefact; only mouldic preservation of the bored substrate would expose the distinctive Spirichnus boring. These ichnofossils are united in their occurrence in unusual preservational systems.     

Trace Fossils from Lower Cambrian Wusongger Formation in Xiaoerbulake Outcrop,Kalpin Area,Xinjiang

Based on the analysis of trace fossils collected from the typical outcrop of the Lower Cambrian Wusongger Formation in the Kalpin area, ten ichnospecies of six ichnogenus were identified in the upper member of the Wusongger Formation, with most of them being found for the first time. The trace fossils are described seriatim. The ichnospecies are mainly represented by such common trace fossils as Ophiomorpha nodosa, Helminthopsis hieroglyphica, Helminthopsis ichnosp., Planolites beverleyensis, Planolites vulgaris, Planolites montanus, Palaeophycus striatus, Palaeophycus curvatus?, Cochlichnus anguineus and Rituichnus elongatum. The trace fossils are of high diversity and low abundance, and can be diagnosed as the Cruziana ichnofacies. They can be interpreted as having formed in a shallow water environment.     

Trace fossils and their palaeoenvironmental significance in the Lameta Formation of Salbardi and Belkher area, district Amravati, Maharashtra, India

Trace fossils namely, Planolites, Planolites montanus, Thalassinoides and stuff burrows belonging to the Lameta Formation of Central India are reported for the first time from two new localities, i.e., Salbardi and Belkher area. Both the exposures are well represented by the typical marker lithounits as of Jabalpur, Nand-Dongargaon area of central India, i.e., greenish yellow sandstone, clays of various shades, chertified and brecciated limestones, etc. These successions are devoid of megaforal remains, therefore, these biogenic structures play a significant role in the interpretation of palaeoenvironment and palaeoecology during the deposition of the Lameta sediments. The trace fossils assemblage shows that the benthonic community was mainly polychaetes and crustaceans.     

Ichnology of the Callovian-Oxfordian rocks of the Katrol hill range,Kachchh, western India

On the basis of distinct lithologic features such as composition, grain size, bedding characteristics and sedimentary structures, six facies were identified in Callovian to Oxfordian rocks exposed southwest of Bhuj, Kachchh. They are interbedded calcareous shale-siltstone (ICSSF), limestone (LF), ferruginous sandstone (FerSF), felspathic sandstone (FelSF), grey shale (GSF) and oolitic limestone (OLF) facies. The rich and highly diversified trace fossils reveal a wide range of animal behaviours represented by dwelling, feeding, crawling and resting structures. Horizontal feeding structures are found abundantly in all lithofacies indicating low wave and current energy and deposition of poorly sorted muddy to sandy sediments. A few coarse layers containing Arenicolites, Ophiomorpha and Skolithos indicate the presence of opportunistic animals (due to their first appearance under harsh conditions) under -intermittently moderate wave and current energy or storm wave conditions (due to coarse grain size and dominance/presence of only vertical trace fossils) in the shoreface zone. Taenidium occurs mainly in the lower shoreface to transitional zone suggesting low to moderate energy conditions. Thalassinoides occurs in middle to lower shoreface settings under relatively low-energy conditions. Zoophycos represents offshore environment, where it occupies the deepest bioturbation levels.The characteristic lithofacies and assemblages of trace fossils in the rocks of the Chari/Jumara Formation indicate a depositional environment fluctuating from the upper shoreface to offshore zone.     

Trace fossils from Talchir carbonate concretions,Giridih basin,Jharkhand

The carbonate concretions occurring at the bottom of Talchir fissile shale facies preserved signatures of various trace fossils along with a cast of doubtful organisms and cyanobacterial mat structures. The host shale deposited under glacial melt water fed lacustrine condition. The concretions, formed in poorly oxygenated conditions, are either of syndepositional origin and/or deposited a little below the sediment water interface and were later exhumed to the depositional surface due to erosion of soft mud overlying them. The trace fossils are both megascopic and microscopic in nature. The megascopic trace fossils are identified on the basis of their morphology as Monocraterion and Rhizocorallium. Some of the megascopic structures described remain problematic at present. The microscopic trace fossils are formed due to the activity of marine meiofauna (possibly by nematodes), which, although produced morphologically show similar traces of known larger ichnogeneras but much smaller than them. The discovery of these trace fossils apparently indicate the influx of saline water into a lacustrine domain during the Talchir sedimentation at Giridih basin. Moreover, presence of the above two megascopic trace fossils in the marine lacustrine carbonate concretions may lead researchers to consider their much wider environmental significance than hitherto believed.     

First record of an arthropod from the Passaic Formation (Late Triassic), near Milford, New Jersey

Mudstones of the Triassic Passaic Formation, near Milford, New Jersey, have yielded the first evidence of an arthropod impression in that formation. Associated trace fossils include Helminthoidichnites, Lockeia, Scoyenia, Spongeliomorpha, and the reptile footprint Rhynchosauroides, representing the Scoyenia ichnofacies. Associated sedimentary structures include desiccation cracks and raindrop impressions. The Passaic sediments were deposited under shallow water lacustrine shoreline conditions subject to periodic subaerial exposure.     

Geochemical and isotopic characterization of trace fossil infillings: New insights on tracemaker activity after the K/Pg impact event

Geochemical and isotopic analyses of the Cretaceous–Paleogene (K/Pg) boundary deposits were conducted at the Caravaca section (External Subbetic, southeast of Spain) in order to evaluate the recovery of the macrobenthic tracemaker community and the bioturbational disturbance. Samples from the infilling material of several lower Danian dark-colored trace fossils (Chondrites, Planolites, Thalassinoides and Zoophycos) located in the uppermost 8-cm of the light upper Maastrichtian strata, as well as samples from the host sedimentary rock of these trace fossils, were analyzed and compared with data from the lower Danian deposits. The values of element ratios indicative of extraterrestrial contamination (Cr/Al, Co/Al and Ni/Al) are higher in the infilling trace fossil material than in the upper Maastrichtian and lower Danian deposits, which suggests a contribution of the ejecta layer. Regarding the isotope composition, the δ¹³C values are lower in the infilling material than in the Maastrichtian host sedimentary rocks surrounding the traces, while the δ¹⁸O are higher in the infilling material. The geochemical and isotopic compositions of the infilling material evidence the unconsolidated character of the sediment, including the red boundary layer. Softground conditions confirm a relatively rapid recovery by the macrobenthic tracemaker community, starting a few millimeters above the K/Pg boundary layer. The mixture of the infilling material of the trace fossils moreover reveals a significant macrobenthic tracemaker activity affecting K–Pg boundary transition sediments that may have significantly altered original signatures.     

Exceptionally favourable life conditions for macrobenthos during the Late Cenomanian OAE-2 event: Ichnological record from the Bonarelli Level in the Grajcarek Unit,Polish Carpathians

Pelagic and hemipelagic sediments of the Bonarelli Level (uppermost Cenomanian) in the Sztolnia section (Grajcarek Unit, Polish Carpathians) contain trace fossils of the Zoophycos ichnofacies, including (in descending order of abundance): Chondrites (smaller and larger forms), Planolites, Thalassinoides, Palaeophycus, Taenidium, Teichichnus, and Zoophycos. They occur in thick bioturbated layers, which are interbedded with rare, thin layers of unbioturbated black shales. The black shale layers mark the Bonarelli Level and are interpreted as a record of anoxia or dysoxia. Coeval sections in the Western Tethys contain similar trace fossils but they are less abundant and these sections are characterized by thicker unbioturbated black shale layers and thinner bioturbated layers. This confirms the exceptionally favourable life conditions in sediments of the Sztolnia section, which do not record strong global anoxia during the OAE-2 event. Such favourable conditions were probably caused by effective oxygenation of pore waters and deep burial of organic matter, which are a consequence of high rates of accumulation and the palaeogeographical location of the section on a flank of a submarine high, under strong circulation.     

Possible animal body fossils from the Late Neoproterozoic interglacial successions in the Kimberley region, northwestern Australia

New specimens of the enigmatic Ediacara-type fossil Palaeopascichnus have been identified from the upper part of the Neoproterozoic Ranford Formation in the Kimberley region, northwest Australia. New material is morphologically similar to Palaeopascichnus and represents the largest species of this genus. They resemble the present-day xenophyophore protists in chamber morphology and growth patterns, supporting the interpretation that Palaeopascichnus is possibly a xenophyophore body fossil rather than a trace fossil. Stratigraphic correlation reveals that the new Palaeopascichnus specimens are preserved in the interglacial successions between the Landrigan/Marinoan and Egan/Ediacaran glaciations. If correlation with the early Ediacaran formations of South Australia is accepted, this represents the earliest known identifiable member of the Ediacara biota. New fossil record fills the evolutionary gap between the Cryogenian and Ediacaran animal assemblages and well-known Ediacaran biota. The new Palaeopascichnus specimens represent the first record of Ediacara-type fossils in Kimberley, and suggest the probability that additional Ediacaran fossils may be found in northwestern Australia.     

新疆准噶尔盆地西北缘石炭纪浊积岩系中遗迹化石的发现及其意义

准噶尔盆地西北缘石炭纪地层主要由碎屑岩组成,沉积厚度5000—7000m,在下石炭统和中石炭统中首次发现了包括浊流在内的重力流沉积和平流岩,大量的深水相遗迹化石。说明盆地西北缘是深水沉积,而非前人普遍认为的属浅水沉积环境。     

Bathymetric significance of the ichnofossil assemblages of the Kulakkalnattam sandstone, Ariyalur area, Cauvery Basin

Well preserved ichnofossils were found in Kulakkalnattam sandstone exposed at Kulakkalnattam stream in Ariyalur area, Cauvery Basin. It consists of infaunal structures of both suspension and deposit feeders. Five ichnofossils are present in a fine to coarse grain sandstone which includes Ophiomorpha, Palaeophycus, Planolites Skolithos, and Thalassinoides. The study infers that ichnofossils Skolithos and Ophiomorpha are infaunal colonization of the suspension feeders in high energy condition in shifting substrate, whereas Thalassinoides and Planolites-Palaeophycus ichnofossils indicate infaunal deposit feeders living at the sediment-sediment interface in low to moderate energy conditions. Furthermore, the abundance and diversity of the trace fossils indicates there was alternatively fluctuations in energy conditions which lead to development of Skolithos and Cruziana ichnofacies type condition during the deposition of Kulkkallanattam sandstone in foreshore-shoreface environments.     

西南极利文斯顿岛晚三叠世遗迹化石及其环境意义

西南极利文斯顿岛晚三叠世迈尔斯陡崖组形成于海底浊积扇中扇的上、下部分,发育着许多海相遗迹化石.在采集到的样品中鉴定共有15个遗迹属、16个遗迹种,其中有10个可以鉴定到遗迹种、两个比较种,4个只鉴定到遗迹属,未鉴定到遗迹种;并建立了1个新遗迹属及新遗迹种是文献中从未发现的.除新遗迹属种外,其余14个遗迹属,15个遗迹种都曾经在深海相浊积岩内发现过,Belorhaphe、Glockerichnus、Lophoctenium、Rhabdoglyphus、Paleodictyon、Sublorenzinia、Spirophycus、Strobilorhaphe、Tuberculichnus、Cochlichnus等属于浊流前产生在深海相泥岩内的高度分异的雕画迹(Graphoglyptida),它们产于泥岩却保存为上覆砂岩底面的铸型凸起.Fucusopsis和Neonereites却产生在砂岩内代表浊流后形成的沉积后遗迹组合.这些遗迹化石属深水的Nereites遗迹相,为研究区的沉积环境增添了可靠依据.     

Deep-sea trace fossils from the mixed carbonate-siliciclastic flysch of the Monte Antola Formation (Late Campanian-Maastrichtian), North Apennines, Italy

Twenty one ichnogenera and 30 ichnospecies are recognized in the Late Campanian – Maastrichtian Monte Antola Formation of Italy, most of them for the first time. They belong to a specific mixture of the Nereites and Ophiomorpha rudis ichnosubfacies of the Nereites ichnofacies. Composition and abundance of trace fossils vary from outcrop to outcrop. Chondrites intricatus, Ch. targionii, Nereites irregularis, Cladichnus fischeri and Trichichnus linearis are most common. Chondrites patulus, Ophiomorpha recta and “Spirophycus” bicornis trace fossils are less common in outcrops, and the other trace fossils are rare or very rare. Generally, ichnogeneric diversity in Cretaceous–Cainozoic calcareous and mixed siliciclastic-calcareous flysch deposits is distinctly lower than in siliciclastic flysch. Significantly, the foremost pre-depositional forms are distinctly rare, probably due to preservational and depositional factors.     

Early Triassic intertidal/subtidal patterns of sedimentation along the southern margins of the Tethyan seaway,Jordan

The early Triassic Dardur Formation, exposed along the northeastern margin of the Dead Sea area, comprises some 63 m of siliciclastic and carbonate rocks, arranged in two coarsening-upward sequences. Each sequence begins with a heterolithic facies (silty shale dolomite and marlstone) and terminates with a sandstone facies.The occurrence of mixed carbonate-dominated clastic coarsening-upward facies sequences, containing pelecypode and ostracode fossils and trace fossils is attributed to deposition in a tidally-dominated environment. Bedload traction transport was responsible for deposition of the sandstone facies in a permanently submerged, shallow subtidal zone, whereas the presence of small scale interference oscillation ripple marks, tidal rhythmites and mudcracks in the heterolithic facies indicate deposition under more fluctuating intertidal conditions.The organization of heterolithic and sandstone into vertically-stacked, coarsening-upward marine facies sequences is attributed to a deepening-upward trend from intertidal to subtidal conditions, in response to a series of minor local transgressions and regressions along the southern margins of the Tethyan seaway during early Triassic times.