Paleoenvironmental changes at the northern limit of the subantarctic Nothofagus forest,lat 37°S,Argentina

Paleoenvironmental changes dating back to before 10,000 yr B.P. at the northernmost occurrences of Nothofagus forests in Argentina at about latitude 37°S permit the reconstruction of past changes in the intensity of the winter rains, related to the southern westerlies that appear to determine the forest boundary. The paleoenvironmental interpretation is based on changes in the proportions of different Nothofagus species and changes in the ratio betwen forest and steppe taxa. The relatively most diverse and dense Nothofagus forest developed only during the last 4500 yr, prior to human impact during the last 300 yr. Before the middle Holocene, climatic conditions must have been different from the modern ones, with less overall precipitation judging from the overall lower amount of tree pollen and the reduction to primarily Nothfagus pumilio. An interval dated to older than 10,000 yr B.P. is characterized by co-occurrence of Prumnopitys andina, Nothofagus pumilio, and shrub-steppe taxa. Prumnopitys andina is known today only from scattered upper montane forest sites in Chile between 36° and 43°S lat and its ecological requirements are essentially unknown. The taxa combination, however, suggests that late-glacial climate must have been drier, and probably cooler than today. This implies that the winter rains and, consequently, the seasonal shift of the westerly circulation was reduced during the late Pleistocene and did not reach modern levels before 8500 yr B.P.     

A late quaternary pollen record from the Transvaal bushveld, South Africa

Pollen spectra from cores of organic spring deposits from the Transvaal provide evidence for the climatic evolution of the province during the last 35,000 yr B.P. or more. The past climatic phases are derived from palynological reconstructions of past vegetation types by comparison of fossil pollen data with modern surface pollen spectra from various localities. Evidence is provided for an early moist, cool phase with relatively mesic bushveld and expanded montane forest in the central Transvaal, followed by a drier period with drier bushveld which probably lasted until approximately 25,000 yr B.P. During the next phase, which at the latest ended about 11,000 yr B.P., the temperatures were probably 5°–6°C cooler than at present. At that time bushveld vegetation in the central Transvaal was replaced by open grassland with macchia elements. Climatic amelioration came and semiarid savanna returned to the plains, at first gradually and then developing into a warm Kalahari thornveld-type vegetation. After 6000 yr B.P. it apparently became slightly wetter and a more broad-leafed bushveld developed. About 4000 yr B.P. it again became cooler and slightly wetter and the bushveld vegetation on the central and northern plains was comparable to present open upland types. After 2000 yr B.P. conditions gradually became warner until about 1000 yr B.P., when the modern climate of the central Transvaal bushveld originated.     

Late-Glacial Vegetation and Climate Change in Western Oregon

Pollen records from two sites in western Oregon provide information on late-glacial variations in vegetation and climate and on the extent and character of Younger Dryas cooling in the Pacific Northwest. A subalpine forest was present at Little Lake, central Coast Range, between 15,700 and 14,850 cal yr B.P. A warm period between 14,850 and 14,500 cal yr B.P. is suggested by an increase inPseudotsugapollen and charcoal. The recurrence of subalpine forest at 14,500 cal yr B.P. implies a return to cool conditions. Another warming trend is evidenced by the reestablishment ofPseudotsugaforest at 14,250 cal yr B.P. Increased haploxylonPinuspollen between 12,400 and 11,000 cal yr B.P. indicates cooler winters than before. After 11,000 cal yr B.P. warm dry conditions are implied by the expansion ofPseudotsuga.A subalpine parkland occupied Gordon Lake, western Cascade Range, until 14,500 cal yr B.P., when it was replaced during a warming trend by a montane forest. A rise inPinuspollen from 12,800 to 11,000 cal yr B.P. suggests increased summer aridity.Pseudotsugadominated the vegetation after 11,000 cal yr B.P. Other records from the Pacific Northwest show an expansion ofPinusfrom ca. 13,000 to 11,000 cal yr B.P. This expansion may be a response either to submillennial climate changes of Younger Dryas age or to millennial-scale climatic variations.     

Quaternary vegetation history of the Mississippi embayment

Nonconnah Creek, located in the loess-mantled Blufflands along the eastern wall of the Lower Mississippi Alluvial Valley in Tennessee displays a sedimentary sequence representing the Altonian Substage through the Woodfordian Substage of the Wisconsinan Stage. The site has a biostratigraphic record for the Altonian and Farmdalian Substages that documents warm-temperate upland oak-pine forest, prairie, and bottomland forest. At 23,000 yr B.P., white spruce and larch migrated into the Nonconnah Creek watershed and along braided-stream surfaces in the Mississippi Valley as far as southeastern Louisiana. The pollen and plant-macrofossil record from Nonconnah Creek provides the first documentation of a full-glacial locality in eastern North America for beech, yellow poplar, oak, history, black walnut, and other mesic deciduous forest taxa. During the full and late glacial, the Mississippi Valley was a barrier to the migration of pine species, while the adjacent Blufflands provided a refuge for mesic deciduous forest taxa. Regional climatic amelioration, beginning about 16,500 yr B.P., is reflected by increases in pollen percentages of cooltemperate deciduous trees at Nonconnah Creek. The demise of spruce and jack pine occurred 12,500 yr B.P. between 34° and 37° N in eastern North America in response to postglacial warming.     

Sedimentologic and palynologic records of the last deglaciation and Holocene from Ballston Lake, New York

Continuous pollen and sediment records from two ∼8.5-m-long cores document late Pleistocene and Holocene sedimentation and vegetation change in the Ballston Lake basin, eastern New York State. Pebbles at the base of both cores and the geomorphology of the watershed reflect the presence of the Mohawk River in the basin prior to ∼12,900 ± 70 cal yr B.P. Ballston Lake formed at the onset of the Younger Dryas (YD) by an avulsion of the Mohawk River. The transition from clay to gyttja with low magnetic susceptibility (MS), low bulk density, and high organic carbon indicates rapid warming and increased lake productivity beginning 11,020 cal yr B.P. MS measurements reveal that the influx of magnetic particles, associated with pre-Holocene clastic sedimentation, ceased after ∼10,780 cal yr B.P. The pollen record is subdivided into six zones: BL1 (12,920 to 11,020 cal yr B.P.) is dominated by boreal forest pollen; BL2 (11,020 to 10,780 cal yr B.P.) by pine (Pinus) forest pollen; BL3 (10,780 to 5290 cal yr B.P.) by hemlock (Tsuga) and mixed hardwood pollen; BL4 (5290 to 2680 cal yr B.P.) by mixed hardwood pollen; BL5a (2680 cal yr B.P. to 1030 cal yr B.P.) by conifer and mixed hardwood pollen; and BL5b (1030 cal B.P. to present) by increasing ragweed (Ambrosia) pollen. A 62% decrease in spruce (Picea) pollen in <320 cal years during BL1 reflects rapid warming at the end of the YD. Holocene pollen zones record more subtle climatic shifts than occurred at the end of the YD. One of the largest changes in the Holocene pollen spectra began ∼5300 cal yr B.P., and is characterized by a marked decline in hemlock pollen. This has been noted in other pollen records from the region and may record preferential selection of hemlock by a pathogen or parasites.     

Holocene insect fossils from two sites at Ennadai Lake, Keewatin, Northwest Territories, Canada

Ennadai Lake, in the forest-tundra ecotonal region of Keewatin, Northwest Territories, Canada, has been the subject of several paleoecological investigations (palynology, plant macrofossils, fossil soils). This study concerns Holocene insect fossils at Ennadai, a new approach in a region shown to be sensitive to climatic change. The Ennadai I site yielded 53 taxa, representing 13 families of Coleoptera and 7 families of other insects and arachnids, including abundant ants. These fossils range in age from about 6300 to 630 yr B.P. The Ennadai II site produced fossils of 58 taxa, including 13 beetle families and 15 families of other arthropods, ranging in age from 4700 to 870 yr B.P. The insect evidence suggests the presence of trees in the Ennadai region from 6000 to 2200 yr B.P. A conifer pollen decline from 4800 to 4500 yr B.P. at Ennadai has previously been interpreted as an opening up or retreat of forest in response to climatic cooling, but the insect fossils reveal the continued presence of trees during this interval. Both insect assemblages suggest trends of forest retreat and tundra expansion between about 2200 and 1500 yr B.P., presumably due to climatic cooling, with a return of woodland by about 1000 yr B.P.     

Holocene climatic change in the northern Midwest: Pollen-derived estimates

Mapping of Holocene pollen data in the midwestern United States has revealed several broadscale vegetational changes that can be interpreted in climatic terms. These changes include (1) the early Holocene northward movement of the spruce-dominated forest and its later southward movement after 3000 yr B.P. and (2) the eastward movement of the prairie/forest border into southwestern Wisconsin by 8000 yr B.P. and its subsequent westward retreat after 6000 yr B.P. When certain basic assumptions are met, multiple regression models can be derived from modern pollen and climate data and used to transform the pollen record of these vegetational changes into quantitative estimates of temperature or precipitation. To maximize the reliability of the regression equations, we followed a sequence of procedures that minimize violations of the assumptions that underlie regression analysis. Reconstructions of precipitation during the Holocene indicated that from 9000 to 6000 yr B.P. precipitation decreased by 10 to 25% over much of the Midwest, while mean July temperature increased by 0.5° to 2.0°C. At 6000 yr B.P. precipitation was less than 80% of its modern values over parts of Wisconsin and Minnesota. After 6000 yr B.P. precipitation generally increased, while mean July temperature decreased in the north, and increased in the south. The time of the maximum temperature varies within the Midwest and is earlier in the north and later in the south.     

Palynology of cushion bogs of the Cordillera Pelada,Province of Valdivia,Chile

Fossil pollen identified in the earliest sediments of three cushion bogs in the Cordillera Pelada (40°10′S, 73°30′W) dated 10,425 ¹⁴C yr B.P. includes the subantarctic species Dacrydium fonckii, Tetroncium magellanicum, Astelia pumila, Gaimardia australis, Donatia fascicularis, and Drosera uniflora. All grow today in the Cordillera Pelada and range poleward to the southernmost Province of Magallanes; one species, Drapetes muscosa, included with the pollen of these plants in the earliest record, is no longer a constituent of the flora but is limited only to subantarctic Chile. Available evidence indicates that plants survived the last glaciation north of the glacial border with the course of postglacial migration southward following the wastage of the glacier complex. Holocene climatic and vegetational changes in the Cordillera Pelada are interpreted in the context of regional reconstructions which show maximum warmth about 9000 yr ago with a pronounced dry period lasting from 9000 to 6500 yr B.P. Maximum precipitation was later reached around 4000 yr ago but has decreased overall since then. The regional decline of the endemic gymnosperm Fitzroya cupressoides, which today is extensively destroyed in the Cordillera Pelada, follows this decrease in precipitation. These climatic data suggest a net south ward shift in the zone of westerly winds that bring rainfall to the region over the past 4000 yr.     

Late quaternary vegetation of southern Isla Grande de Chiloé, Chile

Late-glacial-Holocene forest history of southern Isla Chiloé (latitude 43°10′ S) was reconstructed on the basis of pollen analysis in three profiles (Laguna Soledad, Laguna Chaiguata, Puerto Carmen). Prior to 12,500 yr B.P. pollen records are dominated by plant taxa characteristic of open habitats (Zone I). From 12,500 yr B.P. to the present, tree species predominate in the pollen records (Zones II–V). Between 12,500 and 9500 yr B.P. ombrophyllous taxa (Nothofagus, Podocarpus nubigena. Myrtaceae, Fitzroya/Pilgerodendron, and Drimys) are frequent in all pollen diagrams, suggesting a wetter and colder climate than the present. Between 9000 and 5500 yr B.P. Valdivian forest elements, such as Nothofagus dombeyi type, Weinmannia, and Eucryphia/Caldcluvia, dominate, indicating a period of drier and warmer climate. From 5500 yr B.P. onward, the expansion of mixed North Patagonian-Subantarctic forest elements and the increased frequence of Tepualia suggest increased rainfall and temperatures oscillating around the modern values.The change from open to forest vegetation (ca. 12,500 yr B.P.) probably represents the most pronounced climatic change in the record and can be interpreted as the glacial-postglacial transition in the study area.     

Quantitative climate reconstruction from pollen data in the Grand Duchy of Luxembourg since 15 000 yr BP

Monthly temperature and precipitation estimates for the period 15000–2000 yr BP are statistically derived from pollen analytical data obtained from a site near Echternach, Luxembourg. A continuous warming trend of over 10°C is recorded for the period 10500—8000 yr BP, along with a rise in precipitation of 500 mm. A fairly constant temperature characterised the period between 8000 and 2000 yr BP, though an episode of slight temperature decrease (-2°C) occurred at about 4500 yr BP. The most humid intervals seem to have been at around 9000–7000 and 5000–3000 yr BP (principally during the warm season: more continental influence). In contrast with results obtained from more southerly sites, the ‘Older Dryas’ climatic reversal is clearly recorded at Echternach slightly before 12000 yr BP. It is marked by a cooling in July temperature of about 4°C and a decrease in monthly precipitation of about 30 mm (40%).     

Holocene vegetation history of the Prince William Sound region, south-central Alaska

Vegetation history during the Holocene is interpreted from the pollen and sedimentary records of nine sections of peat deposits located in sedge tundra at sites in the northern and northwestern parts of the Prince William Sound region. Basal radiocarbon ages of the deposits are between 10,015 and 580 yr B.P. Modern surface pollen data from these and 25 additional sites, ranging from lowlands to an altitude of 675 m in the alpine tundra, were used to aid in the interpretation of the fossil records. Both frequency and influx pollen diagrams of the oldest section disclose a sequence of communities beginning with sedge tundra, containing thickets of willow and alder, followed by alder, which became predominant at about 8300 yr B.P. Later, alder declined, and an inferred growth of sedge tundra and the establishment of colonies of mountain hemlock and Sitka spruce with some western hemlock occurred about 2680 yr B.P. Finally, regrowth of sedge tundra accompanied by the development of forest communites took place over the past 2000 yr. The influence of glacier advances on the vegetation in the fjords occurred during Neoglacial episodes dated at 3200–2500 yr B.P. and during recent centuries. Regional Holocene tectonic activity was also an influential factor, especially at the time of the 1964 earthquake.     

Lateglacial,early and middle holocene environments,human occupation,and resource use in the Atacama (Northern Chile)

Paleoenvironmental data from the Atacama Altiplano (21°–24°S) indicate that water, vegetation and animal resources were more abundant during lateglacial and early Holocene times than today. The rate of precipitation increased above 4000 m elevation to 400–500 mm/yr compared to the present 200 mm/yr. Dry conditions prevailed below 3500 m. Evidence of Paleoindian habitation is still missing, even though there is no evidence for environmental prohibitors during lateglacial time. The early Archaic hunters (10,820 yr B.P.-ca. 8500 yr B.P.) inhabited the Altiplano (high Puna) and its western slope, where water was available due to higher river runoff from the Altiplano, and the resources in different elevation zones were accessible. Natural resources decreased significantly during the middle Archaic period (8500-ca. 5000 yr B.P.). Lakes receded to today's levels, pedogenesis in the Altiplano terminated, and human activities were restricted to the most stable sites in the Río Lao and the Río Purifica catchments north of 23°S. The less stable oases south of 23°S (Salar de Atacama and Punta Negra) were abandoned. The climatic changes are best explained by shifts of the (sub)tropical circulation. © 1994 John Wiley & Sons, Inc.     

Postglacial Vegetation and Climate of the Cascade Range, Central Oregon

Pollen data from two sites provide information on the postglacial vegetation and climate history of the Cascade Range. Indian Prairie in the western Cascade Range was colonized by subalpine forests of Pinus, Picea, and Tsuga and open meadows prior to ca. 12,400 ¹⁴C yr B.P. The treeline lay 500 to 1000 m below its modern elevation and conditions were cooler than at present. From ca. 12,400 to ca. 9950 ¹⁴C yr B.P. Abies became important and the forest resembled that presently found at middle elevations in the western Cascade Range. The pollen record implies a rise in treeline and warmer conditions than before. From ca. 10,000 to 4000-4500 ¹⁴C yr B.P., conditions that were warmer and effectively drier than today led to the establishment of a closed forest composed of Pseudotsuga , Abies, and, at lower elevations, Quercus and Corylus . During this period, Gold Lake Bog in the High Cascades was surrounded by closed forest of Pinus and Abies. The early-Holocene pollen assemblages at both Indian Prairie and Gold Lake Bog lack modern analogues, and it is likely that greater-than-present summer radiation fostered unique climatic conditions and vegetation associations at middle and high elevations. In the late Holocene, beginning ca. 4000-4500 ¹⁴C yr B.P., cooler and more humid conditions prevailed and the modern vegetation was established. A comparison of these sites with others in the Pacific Northwest suggests that major patterns of vegetational change at individual sites were a response to large-scale changes in the climate system that affected the entire region.     

Late pleistocene/early holocene environments and climates,fauna, and human occupation in the argentine altiplano

The Barro Negro site (23°S lat., 65°37′W long.) in the Altiplano (Puna) of northwestern Argentina contains a well stratified sequence of remains of Hippidion, the American extinct horse, camelids, and archaeological materials, which is the focus of this study. In addition to establishing a reliable chronology, paleoenvironmental information was obtained based on analyses of pollen and stable isotopes (oxygen and carbon) from bone and marl. The data indicate that Hippidion was present at the site between 12,000 and 10,000 yr B. P., at a time when Altoandean grasslands had expanded to lower elevations. By 10,000 yr B.P., when modern semi-arid sub-puna scrub had replaced the Altoandean grasslands, only camelids (Lama or Vicugna) were present, simultaneous with the first evidence of local human occupation. This suggests that a climatic shift from cool and moist (winter rain regime) to warm and dry (summer rain regime) conditions took place simultaneously with the disappearance of the American horse and the appearance of camelids and man.     

An 8000-year record of vegetation, climate, and human disturbance from the Sierra de Apaneca, El Salvador

An ∼8000-cal-yr stratigraphic record of vegetation change from the Sierra de Apaneca, El Salvador, documents a mid-Holocene warm phase, followed by late Holocene cooling. Pollen evidence reveals that during the mid-Holocene (∼8000-5500 cal yr B.P.) lowland tropical plant taxa were growing at elevations ∼200-250 m higher than at present, suggesting conditions about 1.0°C warmer than those prevailing today. Cloud forest genera (Liquidambar, Juglans, Alnus, Ulmus) were also more abundant in the mid-Holocene, indicating greater cloud cover during the dry season. A gradual cooling and drying trend began by ∼5500 cal yr B.P., culminating in the modern forest composition by ∼3500 cal yr B.P. A rise in pollen from weedy plant taxa associated with agriculture occurred ∼5000 cal yr B.P., and pollen from Zea first appeared in the record at ∼4440 cal yr B.P. Human impacts on local vegetation remained high throughout the late Holocene, but decreased abruptly following the Tierra Blanca Joven (TBJ) eruption of Volcán Ilopango at ∼1520 cal yr B.P. The past 1500 years are marked by higher lake levels and periodic depositions of exogenous inorganic sediments, perhaps indicating increased climatic variability.     

Late Wisconsin vegetational and climatic history at Kylen Lake, northeastern Minnesota

Kylen Lake, located within the Toimi drumlin field, is critically positioned in relation to Late Wisconsin glacial advances, for it lies between the areas covered by the Superior and St. Louis glacial lobes between 12,000 and 16,000 yr B.P. The pollen and plant-macrofossil record suggests the presence of open species-rich “tundra barrens” from 13,600 to 15,850 yr B.P. Small changes in percentages of Artemisia pollen between 14,300 and 13,600 yr B.P. appear to be artifacts of pollen-percentage data. Shrub-tundra with dwarf birch, willow, and Rhododendron lapponicum developed between 13,600 and 12,000 yr B.P. Black and white spruce and tamarack then expanded to form a vegetation not dissimilar to that of the modern forest-tundra ecotone of northern Canada. At 10,700 B.P. spruce and jack pine increased to form a mosaic dominated by jack pine and white spruce on dry sites and black spruce, tamarack, and deciduous trees such as elm and ash on moist fertile sites. At 9250 yr B.P. red pine and paper birch became dominant to form a vegetation that may have resembled the dry northern forests of Wisconsin today. The diagram terminates at 8410 ± 85 yr B.P. Climatic interpretation of this vegetational succession suggests a progressive increase in temperature since 14,300 yr B.P. This unidirectional trend in climate contrasts with the glacial history of the area. Hypotheses are presented to explain this lack of correspondence between pollen stratigraphy and glacial history. The preferred hypothesis is that the ice-margin fluctuations were controlled primarily by changes in winter snow accumulation in the source area of the glacier, whereas the vegetation and hence the pollen stratigraphy were controlled by climatic changes in front of the ice margin.     

Late Quaternary vegetational and climatic change in the Popayán region,southern Colombian Andes

Late Pleistocene and Holocene vegetational and climatic change have been studied palynologically at a site at 1750 m elevation in the subandean vegetation belt near Popayán, in the southern Colombian Andes. Time control on the pollen record is based on six AMS ¹⁴C ages, ranging from possibly Middle Pleniglacial time (around 50000 yr BP) to 1092 ± 44 yr BP. Because of the presence of two hiatuses only the Middle Pleniglacial and Late Holocene periods (the last 2300 yr BP) are represented. Pollen data indicate the presence of closed subandean forest during glacial time. Changes in the contribution of pollen originating from the uppermost and lowermost subandean forest belts, changes in the contribution of a number of other subandean forest taxa, and changes in species composition between the three pollen zones, suggest that the climate during the Middle Pleniglacial was markedly colder, and perhaps also wetter, than during the Late Holocene. Pollen assemblages from the Late Holocene indicate that the landscape has been affected by deforestation and agriculture since at least 2300 yr BP, but that human impact decreased in the last 780 yr BP. © 1998 John Wiley & Sons, Ltd.     

Holocene climate of New England

Stratigraphic studies of pollen and macrofossils from six sites at different elevations in the White Mountains of New Hampshire demonstrate changes in the distributions of four coniferous tree species during the Holocene. Two species presently confined to low elevations extended farther up the mountain slopes during the early Holocene: white pine grew 350 m above its present limit beginning 9000 yr B.P., while hemlock grew 300–400 m above its present limit soon after the species immigrated to the region 7000 yr. B.P. Hemlock disappeared from the highest sites about 5000 yr B.P., but both species persisted at sites 50–350 m above their present limits until the Little Ice Age began a few centuries ago. The history of the two main high-elevation conifers is more difficult to interpret. Spruce and fir first occur near their present upper limits 9000 or 10,000 yr B.P. Fir persisted in abundance at elevations similar to those where it occurs today throughout the Holocene, while spruce became infrequent at all elevations from the beginning of the Holocene until 2000 yr B.P. These facts suggest a more complex series of changes than a mere upward shift of the modern environmental gradient. Nevertheless, we conclude that the minimum climatic change which would explain the upward extensions of hemlock and white pine is a rise in temperature, perhaps as much as 2°C. The interval of maximum warmth started 9000 yr B.P. and lasted at least until 5000 yr B.P., correlative with the Prairie Period in Minnesota.     

Bronze-early iron age environment of the Iman River Basin,Russian far east

Three archaeological sites of the Bronze and Early Iron Ages were investigated using palynological methods. To interpret the fossil spectra more accurately, recent pollen from alluvium, surface soil and cultural layers within modern villages were also studied in the area. During the time interval when the ancient cultural layers accumulated, thinned-out birch forests were replaced by mixed coniferous and broad-leaved trees. This might have been caused by climatic transition which possibly influenced the changing cultural traditions within the area. The climatic transition also includes a change in humidity. The wettest stages are recorded by pollen data during the transition period between the Bronze and Early Iron Age. This corresponds to the Subboreal/Subatlantic boundary within the European Holocene (around 2500 yr B.P.). © 1994 John Wiley & Sons, Inc.     

Late pleistocene temperature depression and vegetation change in Ecuadorian Amazonia

Paleoecological (pollen, phytolith, and wood) analyses of sediments, radiocarbon dated 33,000 to 26,000 yr B.P., from two sites in Ecuadorian Amazonia provide data that suggest a cooling of ca. 7.5°C below present in equatorial lowlands from 33,000 to 30,000 yr B.P. A period of warning followed in which novel species assemblages, a blend of montane and lowland floral components, persisted for at least 4000 years. These data of forest community change, from sites lying within the postulated glacial rain forest Napo refugium, provide the strongest paleoecological refutation of the refugial hypothesis yet obtained. The large temperature depression at ca. 30,000 yr B.P. allows the possibility that if maximum cooling at the equator was synchronous with the last glacial maximum (LGM) of the northern hemisphere, freezing temperatures would have been experienced in parts of lowland Amazonia between 25,000 and 18,000 B.P.