数据稀缺生态系统中多种类质量谱模型的构建

多种类生态模型已经被广泛应用于渔业活动影响应预测和管理措施效果评估。质量谱模型是一种基于生理过程构建的生态模型,该模型为描述鱼类群落在个体摄食变异和随个体发生的生态位迁移提供了一个可行的方法。尽管生态模型在增进生态系统认识上具有重要意义,其应用在数据稀缺的渔业中受到很大限制。作为实践基于生态系统渔业管理(EBFM)的第一步,本研究构建了海州湾鱼类群落的质量谱模型。本研究详述了数据收集和模型参数化的过程,以促进该模型在数据稀缺的生态系统中未来的应用。作为一个范例,研究展示了不同捕捞努力量对生态系统的影响,并采用一套生态指标监测其动态。群落生物量、多样性指数、W指数,大鱼指数(LFI),平均体重和群落质量谱斜率对捕捞压力的响应呈非线性,最大的捕捞强度并非总是对鱼类群落产生最强的影响。本文强调了构建谱模型在生态研究中的的价值和可行性,并讨论了模型的局限性和改进的可能。本研究旨在促进质量谱模型的广泛应用以更好地支持基于生态系统的渔业管理。     

大西洋热带海域长鳍金枪鱼渔场预报模型的比较

为提高大西洋热带海域长鳍金枪鱼(Thunnus alalunga)渔场预报的准确率,对K最近邻(k nearest neighbor,KNN)、逻辑斯蒂回归(logistic regression,LR)、决策与分类树(classfication and regression tree,CART)、梯度提升决策树(gr...     

基于捕捞努力量的中西太平洋鲣鱼围网渔业入渔预测分析

中西太平洋是全球主要的鲣鱼（Katsuwonus pelamis）围网作业渔场,渔场极易受到海洋环境的影响,但渔场分布在众多岛国的管辖海域,如何科学指导企业准确入渔是重要的研究课题。本文根据1995-2012年中西太平洋鲣鱼围网捕捞生产统计数据,选取产量最高的22个海区（5°×5°）,结合Niño3.4区海表温度距平值（SSTA）和作业海域表温（SST）,研究中西太平洋鲣鱼围网渔场的空间分布规律,同时,以各海区捕捞努力量（作业次数）所占的百分比为入渔指标,建立基于环境因子的入渔决策模型。研究认为,中西太平洋鲣鱼捕捞努力量在纬度方向上主要分布于5°S~5°N,其累计捕捞努力量占所有作业海区的87.4%,其中以130°~140°E经度范围为最高,其捕捞努力量占22个海区的45.08%。入渔指标与Niño3.4区的SSTA、作业海域SST均符合正态模型（P<0.01）,Niño3.4区的SSTA最适值为0.25℃,作业海域SST最适值在29.5℃左右。对预测和实际排名前十的海域进行统计发现,预测值与实际值基本一致。研究认为,所建立的入渔预测模型可有效指导企业的渔业生产,为提高企业生产效率提供支撑。     

基于神经网络的南太平洋长鳍金枪鱼渔场预报

南太平洋长鳍金枪鱼是我国远洋渔业的重点捕捞对象;对南太平洋长鳍金枪鱼进行准确的渔场预报;可以提高捕捞效率;提高渔业的生产能力。本研究根据1993-2010年南太平洋长鳍金枪鱼的延绳钓生产数据以及海洋卫星遥感数据（海水表面温度;SST;海面高度;SSH）和ENSO（El Niño-Southern Oscillation）指标;采用DPS（data processing system）数据处理系统中的BP人工神经网络模型;以渔获产量（单位时间的渔获尾数）和单位捕捞努力量渔获量（CPUE;Catch per unit of effort）分别作为中心渔场的表征因子;并作为BP模型的输出因子;以月、经度、纬度、SST、SSH和ENSO指标等作为输入因子;分别构建4-3-1;5-4-1;5-3-1;6-5-1;6-4-1;6-3-1等BP模型结构;比较渔场预报模型优劣。研究结果表明;以CPUE作为输出因子的BP人工神经网络结构总体上较优;其中以6-4-1模型结构为最优;相对误差只有0.006 41。研究认为;以CPUE为输出因子的6-4-1结构的人工神经网络模型;能够准确预报南太平洋长鳍金枪鱼的渔场位置。     

Introducing a transferable fishing day management regime for Pacific Island countries

On 1 December 2007, eight ‘Small Island Developing States’ in the Western and Central Pacific Ocean implemented a management regime restricting the total number of days fished by tuna purse seine vessels within their waters, commonly referred to as the Vessel Day Scheme (VDS). The VDS is seen as one component of management arrangements to reduce fishing mortality on bigeye and yellowfin tuna, constrain fishing effort, and increase the rate of return from access fees by Distant Water Fishing Nations.     

A spatial ecosystem and populations dynamics model (SEAPODYM) – Modeling of tuna and tuna-like populations

An enhanced version of the spatial ecosystem and population dynamics model SEAPODYM is presented to describe spatial dynamics of tuna and tuna-like species in the Pacific Ocean at monthly resolution over 1° grid-boxes. The simulations are driven by a bio-physical environment predicted from a coupled ocean physical–biogeochemical model. This new version of SEAPODYM includes expanded definitions of habitat indices, movements, and natural mortality based on empirical evidences. A thermal habitat of tuna species is derived from an individual heat budget model. The feeding habitat is computed according to the accessibility of tuna predator cohorts to different vertically migrating and non-migrating micronekton (mid-trophic) functional groups. The spawning habitat is based on temperature and the coincidence of spawning fish with presence or absence of predators and food for larvae. The successful larval recruitment is linked to spawning stock biomass. Larvae drift with currents, while immature and adult tuna can move of their own volition, in addition to being advected by currents. A food requirement index is computed to adjust locally the natural mortality of cohorts based on food demand and accessibility to available forage components. Together these mechanisms induce bottom-up and top-down effects, and intra- (i.e. between cohorts) and inter-species interactions. The model is now fully operational for running multi-species, multi-fisheries simulations, and the structure of the model allows a validation from multiple data sources. An application with two tuna species showing different biological characteristics, skipjack (Katsuwonus pelamis) and bigeye (Thunnus obesus), is presented to illustrate the capacity of the model to capture many important features of spatial dynamics of these two different tuna species in the Pacific Ocean. The actual validation is presented in a companion paper describing the approach to have a rigorous mathematical parameter optimization [Senina, I., Sibert, J., Lehodey, P., 2008. Parameter estimation for basin-scale ecosystem-linked population models of large pelagic predators: application to skipjack tuna. Progress in Oceanography]. Once this evaluation and parameterization is complete, it may be possible to use the model for management of tuna stocks in the context of climate and ecosystem variability, and to investigate potential changes due to anthropogenic activities including global warming and fisheries pressures and management scenarios.     

自然死亡和亲体补充关系对黄鳍金枪鱼资源评估的影响

黄鳍金枪鱼(Thunnus albacares)是全球经济和生态价值最重要的鱼类之一,其资源养护和管理受到各方的高度关注。本文依据年龄结构产量模型研究了印度洋黄鳍金枪鱼的资源状态,着重探讨了其生活史特征的不确定性对资源评估结果的影响。研究结果显示,1960-1985年间印度洋黄鳍金枪鱼资源量保持相对稳定,之后开始逐渐下降,相应的捕捞死亡系数也在2010年之后迅速增加,目前其种群可能存在过度捕捞(F₂₀₂₀/F_MSY>1,SSB₂₀₂₀/SSB_MSY<1)。印度洋黄鳍金枪鱼的资源评估结果对自然死亡系数(M)和亲体-补充量关系陡度参数(h)的改变较为敏感。当h增大时,SSB_MSY和初始SSB(即SSB₀)的变化较大,分别减少了约25.53万t和34.04万t;F₂₀₂₀/F_MSY减小了1.15。当M增大时,F₂₀₂₀/F_MSY、SSB_MSY...     

The past,present and future use of drifting fish aggregating devices (FADs) in the Indian Ocean

The use of drifting fish aggregating devices (FADs) has become the dominant practice in tropical tuna purse seine fishing. However, just as FADs can increase fishing efficiency, their use has been associated with several negative ecosystem impacts, and moves are being made to manage the use of FADs. In the evaluation of potential management options it is important to consider how fishers will respond to the introduction of control measures, which first requires an understanding of fishery and fleet dynamics. This paper addresses this need by characterising the past and present use of FADs in the Indian Ocean tropical tuna purse seine fishery. The paper describes historical trends in fishing practices, summarises spatiotemporal patterns in the use of FADs and establishes and attributes variation in FAD fishing strategies within the fleet. It also provides an overview of current FAD management policies in the Indian Ocean and examines the observed effects of existing measures on the behaviour of the purse seine fleet. Using this comprehensive understanding, the potential impact on the purse seine fleet of a number of plausible FAD management options are discussed and inferences are drawn for the future sustainability of tropical tuna purse seine fishing in the Indian Ocean.     

Ecological Risk Assessment for seabird interactions in Western and Central Pacific longline fisheries

The risk of seabird–fishery interactions in the Western and Central Pacific Ocean (WCPO) was examined by analysing the overlap of seabird distributions with tuna and swordfish pelagic longline fisheries managed by the Western and Central Pacific Fisheries Commission (WCPFC) and its constituent members. The study used spatially-explicit Productivity–Susceptibility Analysis (PSA). Key data inputs were species productivity, fishing effort, likelihood of capture and species density by region. The outputs tailored results to the needs of fisheries- and wildlife-managers, indicating areas of greatest risk of species interactions, species of greatest concern for population impacts, and the flags or fisheries most likely to contribute to the risk. Large albatross species were found to be most likely to suffer population effects when exposed to longline fishing activity, followed by the larger petrels from the genuses Procellaria, Macronectes and Pterodroma. A mixture of coastal states with nesting seabird populations in their Exclusive Economic Zones (New Zealand, Australia and United States of America), distant water fishing nations (Japan, Taiwan) and flags of convenience (Vanuatu) contributed 90% of the risk to seabird populations. Recommendations include enhancing the level of fisheries observer monitoring in areas indicated as high to medium risk for seabird interactions, and consideration of spatial management tools, such as more intensive or more stringent seabird bycatch mitigation requirements in high- to medium-risk areas. The methods used, and similar studies conducted in the Atlantic Ocean could lead to improved targeting of monitoring resources, and greater specificity in the needs for seabird-mitigation measures. This will assist in reducing seabird mortality in longline fishing operations and with more effective use of resources for fishery managers in both domestic fisheries and RFMOs.     

Slow steps towards management of the world's largest tuna fishery

Today, the tuna fishery of the western and central Pacific is one of the world's largest fisheries. Annual catches exceed 2 million mt—approximately 50% of the global tuna catch—with an estimated landed value of USD 3 billion (in 2005). The fishery expanded rapidly from the mid 1980s, following the development of the purse-seine fishery. From the mid 1990s, it was recognised that a Regional Fisheries Management Organisation was required to facilitate cooperation in the management of the resource. Following almost a decade of negotiations and planning, the Western and Central Pacific Fishery Commission (WCPFC) was established in 2004. During the interim period, stock assessments for the key tuna species were developed and revealed that for two species (yellowfin and bigeye tuna) there was concern regarding the increased levels of fishing mortality. However, declarations and resolutions calling for restraint in the expansion of fishing effort were largely unheeded over the last decade and, to date, the WCPFC has been unable to introduce any measures to effectively reduce (or limit) the level of fishing mortality on yellowfin and bigeye tuna. This paper proposes a number of mechanisms for improving the performance of the WCPFC with respect to meeting the conservation and management objectives of the commission and argues that Pacific Island nations need to collectively take the lead to ensure the effective management of the resource.     

基于不同环境因子的中西太平洋鲣鱼资源丰度灰色预测模型构建

鲣Katsuwonus pelamis广泛分布于各大洋热带和亚热带海域,其中以中西太平洋资源量最为丰富。综合评价环境因子对鲣鱼资源量的影响,构建科学的资源预报模型可为我国可持续合理开发该鱼种提供参考。本研究利用1998—2013年中西太平洋渔获量数据,以单位捕捞努力量渔获量(CPUE)为资源相对丰度指标,利用灰色关联方法分析鲣鱼资源相对丰度与环境因子之间的关联度,选取合适的环境因子,并基于不同环境因子构建不同的灰色预测模型对鲣鱼资源相对丰度进行预测,比较选择最优模型。结果表明, 中西太平洋鲣鱼的产量逐年递增,而CPUE在年间有着较大的波动。灰色关联分析认为,海表面温度与CPUE的平均关联度最大,其次为Nino3.4区海表温度距平值,其他的环境因子与CPUE的关联度较小。基于多环境因子的预测模型中,包含所有因子(海表面温度、海表面高度、叶绿素质量浓度a和Nino3.4区海表温度距平值)的模型M1有着最佳的拟合效果,实际值与预测值的相对误差为6.475 2,相关系数为0.687 4;而基于单一环境因子的预测模型中,去除11月SST数据的模型S2有着最佳的拟合效果,实际值与预测值的相对误差为7.419 2,相关系数为0.791 0。相比多环境因子的预测模型,单一环境因子预测模型有着较高的稳定性,实际值与预测值直接相关性也较高,可以作为中西太平洋鲣鱼资源相对丰度预报的最优模型。     

中西太平洋延绳钓黄鳍金枪鱼渔场时空分布与温跃层关系

为了解热带中西太平洋延绳钓黄鳍金枪鱼(Thunnus albacares)适宜的温跃层参数分布区间,采用Argo浮标温度信息和中西太平洋渔业委员会(The Western and Central Pacific Fisheries Commission,WCPFC)的黄鳍金枪鱼延绳钓渔获数据,绘制了热带中西太平洋月平均温跃层特征参数和月平均CPUE的空间叠加图,用于分析热带中西太平洋黄鳍金枪鱼中心渔场时空分布和温跃层特征参数间的关系。分析结果表明:热带中西太平洋温跃层上界深度、温度具有明显的季节性变化,而温跃层下界深度、温度季节性变化不明显,黄鳍金枪鱼中心渔场分布和温跃层季节性变化有关。全年中心渔场的位置分布在温跃层上界深度高值区域,随温跃层上界深度高值区域季节性南北移动。在新几内亚以东纬向区域(5°N~10°S,150°E~170°W)上界深度值全年都在70~100m之间,全年都是延绳钓黄鳍金枪鱼中心渔场。中心渔场上界温度多在26℃以上,但是在上界温度超过30℃区域,CPUE值较小。中心渔场主要分布在温跃层下界深度两条高值带之间区域,在温跃层下界深度超过300m和小于150m区域,CPUE值均偏低。中心渔场主要分布在下界温度低于13℃区域,下界温度超过17℃难以形成中心渔场。频次分析和经验累积分布函数计算其适宜温跃层特征参数分布,得出中西太平洋黄鳍金枪鱼适宜的温跃层上界温度和深度分别是27~29.9℃和70~109m;适宜的温跃层下界温度和深度分别是11~13.9℃和250~299m。文章初步得出中西太平洋黄鳍金枪鱼中心渔场温跃层各特征参数的适宜分布区间及季节变化特征,为我国金枪鱼实际生产作业提供技术支持。     

大眼金枪鱼的资源现状

大眼金枪鱼的分布很广 ,在南、北纬 4 0°以内的太平洋、印度洋和大西洋均有分布 ,是金枪鱼渔业的重要捕捞种类之一。本文分别论述了大眼金枪鱼在大西洋、印度洋和太平洋的渔业概况、生物学特性、资源状况和管理对策。     

The cooperation strategies of fisheries between Taiwanese purse seiners and Pacific Island Countries

The tuna industry in the Western and Central Pacific Ocean (WCPO) is one of the most valuable fisheries in the world. Taiwan has a long fishing history and is one of the major distant water fishing nations in the surrounding area. This study aimed to review the historical cooperation structure between Taiwanese purse seiners and Pacific Island Countries (PICs). Data were collected by literature analysis and semi-standardized in-depth interviews with experts. The results showed that since 1982, the Taiwanese purse seiners have developed and signed bilateral access agreements with seven PICs under a variety of formats. The fisheries access and licensing arrangements have been transformed to a Vessel Day Scheme (VDS); as a result, the access fees have increased rapidly since 2010. Taiwanese industries have invested in facilities and share ownership with some PICs. Moreover, the Taiwanese purse seiner industries have proclaimed their willingness to follow the Western and Central Pacific Fisheries Commission conservation measures and stated that they will cooperate with PICs. They hold conservative views, however, regarding the further transfer of ownership to PICs. For the conservation of tuna resources in the WCPO, distant water fishing nations and PICs should cooperate to limit fishing capacity and ensure that the access fee benefits PICs.     

Comparison of nominal and standardized catch per unit effort data in quantifying habitat suitability of skipjack tuna in the equatorial Pacific Ocean

In the western and central Pacific Ocean, upper strata waters exhibit highly dynamic oceanographic features under ENSO variability. This has been proved to be responsible for the dynamic change of both abundance and zonal distribution of skipjack tuna(Katsuwonus pelamis). Although causality has been suggested by researchers using physical–biological interaction models, cumulative evidence needs to be obtained and the tenability of assertion needs to be tested from an ecological habitat perspecti...     

Parameter estimation for basin-scale ecosystem-linked population models of large pelagic predators: Application to skipjack tuna

A Spatial Ecosystem and Population Dynamic Model (SEAPODYM) is used in a data assimilation study aiming to estimate model parameters that describe dynamics of Pacific skipjack tuna population on ocean-based scale. The model based on advection–diffusion–reaction equations explicitly predicts spatial dynamics of large pelagic predators, while taking into account data on several mid-trophic level components, oceanic primary productivity and physical environment. In order to improve its quantitative ability, the model was parameterized through assimilation with commercial fisheries data, and optimization was carried out using maximum likelihood estimation approach. To address the optimization task we implemented an adjoint technique to obtain an exact, analytical evaluation of the likelihood gradient. We conducted a series of computer experiments in order to (i) determine model sensitivity with respect to variable parameters and, hence, investigate their observability; (ii) estimate observable parameters and their errors; and (iii) justify the reliability of the computed solution. Parameters describing recruitment, movement, habitat preferences, natural and fishing mortality of skipjack population were analysed and estimated. Results of the study suggest that SEAPODYM with achieved parameterization scheme can help to investigate the impact of fishing under various management scenarios, and also conduct forecasts of a given species stock and spatial dynamics in a context of environmental and climate changes.     

基于GAM的西北太平洋日本鲭资源丰度预测模型建立

日本鲭(Scomber japonicus)是西北太平洋重要的鱼类资源之一,科学预测日本鲭的资源丰度有利于其资源的合理开发和利用。本研究依据日本渔业机构提供的1987–2012年日本鲭太平洋群体的资源量数据,结合产卵场和渔场的海洋环境数据以及气候因子,使用广义加性模型对影响日本鲭太平洋群体的海洋环境和气候因子进行分析,筛选出有显著影响的因子并建立该群体的资源量预测模型。结果表明,与该群体资源量有显著关系的影响因子有:北极涛动指数、太平洋年代际振荡指数、渔场海表面高度、渔场海表面盐度和渔场海表面温度。基于赤池信息准则筛选出的4个资源量预测模型分析表明,包含北极涛动指数、渔场海表面高度和渔场海表面温度的模型有较好的预测效果,该模型的验证结果也通过了t检验(P<0.05),可用于日本鲭太平洋群体资源量的预测。     

基于GAM模型研究水温垂直结构对热带中西太平洋黄鳍金枪鱼渔获率的影响

黄鳍金枪鱼索饵水层影响延绳钓捕捞效率,而黄鳍金枪鱼索饵水层分布受水温垂直结构的影响,因此本文采用GAM模型分析次表层环境变量对延绳钓黄鳍金枪鱼渔获率的影响,评估黄鳍金枪鱼垂直水层分布对中西太平洋黄鳍金枪鱼延绳钓单位捕捞努力量渔获量(Catch Per Unite Effort,CPUE)的作用。模型结果表明,环境因子对热带中西太平洋延绳钓黄鳍金枪鱼渔获率空间分布影响明显。黄鳍金枪鱼延绳钓CPUE在2012年之后快速增多,高渔获率月份出现在北半球夏季,空间上在10°S,140°E附近区域。温跃层上界温度和深度、温跃层下界深度、18℃等温线深度、△8℃等温线深度及其和温跃层下界深度的深度差对延绳钓渔获率影响较大,是影响热带中西太平洋黄鳍金枪鱼延绳钓渔获率的关键环境因子。随着温跃层上界温度和深度值变大,延绳钓CPUE逐渐递增,对延绳钓CPUE影响密切的温度和深度分别为27～28℃和70～90 m。温跃层下界深度对延绳钓CPUE影响在250～280 m时最大;之后随着下界深度的变大,CPUE快速下降。18℃等温线深度对延绳钓CPUE影响呈现先震荡后递增的趋势,影响密切的区域在230 m深度上下...     

基于梯度依赖客观分析技术的全球Argo网格化数据集:构建及初步应用

采用梯度依赖相关尺度方法构建了1套2004—2017年间,月平均的全球海洋(0~1 500 m)1°×1°的Argo数据集,并在对该数据集进行对比检验的基础上,将其初步应用于中西太平洋黄鳍金枪鱼的渔场分析研究。结果表明,所构建的Argo数据集与WOA13数据集的温、盐偏差在上表层(150 m)稍大,最大值分别约为0.5 ℃和0.1,且偏差均随深度的增加而逐渐减小;其与TAO浮标时间序列的温度偏差,2004—2017年间均小于1 ℃,最大盐度偏差则小于0.5,且大部分海域接近0。中西太平洋海域,黄鳍金枪鱼中心渔场多集中在 28~29 ℃ 等温线范围内,在 22 ℃以下的海域单位捕捞努力量渔获量(catch per unit effort,CPUE)值极小;中心渔场区温跃层上界深度范围在20~120 m之间,且中心渔场在各个深度上形成的频数大体呈正态分布,温跃层上界深度为90 m时,形成中心渔场的可能性达到最大。研究表明所构建的数据集在水文环境分析及资源评估中有一定的应用价值。     

A limit reference point to prevent recruitment overfishing of Pacific bluefin tuna

Pacific bluefin tuna (PBF) (Thunnus orientalis) is commercially important in the North Pacific Ocean. Although its stock has been relatively low for decades, international discussions on a long-term management framework, including the definition of a limit reference point (LRP), have only recently started. This paper argues that an LRP for PBF could be developed by determining a biomass level that would prevent recruitment overfishing. First, it reviews the development of LRPs for various tuna species and demonstrates that most of these limits are not necessarily based on biological information on the respective species. Then, the current management of PBF is also reviewed as background information for considering an LRP for PBF. Finally, a variety of simple analyses of the stock–recruitment relationship of PBF are conducted to find a biomass level that would prevent recruitment overfishing—i.e. an LRP below which stocks should not fall. It is concluded that, for the first time to our knowledge, defining such an LRP for a tuna species is possible (about 30 thousand tonnes or 5% of estimated unfished spawning stock biomass in our calculation). Not only is the LRP based on actual experience, but also the logic behind it would be easier for stakeholders to understand than the theoretical LRPs used elsewhere. This LRP should be useful in future in more comprehensive management framework, such as one through management strategy evaluations, in which stakeholder involvement in decision-making is crucial.