Towards an Aptian (Lower Cretaceous) ammonite biostratigraphy of the Mina Texali section,Central Atlantic province (Puebla State,Central Mexico)

This work is a study of the ammonite record of a new stratigraphic section of Aptian age, at the Mina Texali (Puebla State, Central Mexico). A detailed biostratigraphic analysis was carried out on 309 specimens systematically sampled on a bed-by-bed basis. An Aptian ammonite zonation is proposed for the Mina Texali (= MT) section with two interval zones, Dufrenoyia justinae and Caseyella sp., and one informal biostratigraphic unit represented by the Huastecoceras trispinosoides beds. We also analyze the taxonomic composition, paleoecology and some systematic issues of the ammonite record of the MT section. The first record in Mexico of the genera Pseudosaynella, Xerticeras and the nautiloid Heminautilus is identified in this section. The ammonite assemblage is assigned to the proximal part of the outer neritic region of the continental shelf. The ammonite record of the studied section is diagnostic in establishing the lower-upper Aptian transition, and the local zonation of the MT provides important data for the development of an Aptian standard ammonite zonation for the Central Atlantic province.     

Lower Aptian ammonite biostratigraphy and potential for further studies of OAE 1a in Bulgaria

Sediments of Early Aptian age in Bulgaria can be assigned to four different facies: platform carbonates (Urgonian complex), shallow-water siliciclastics, hemipelagic and flyschoid siliciclastics. The taxonomic analysis of the ammonite faunas of 18 sections from these four different facies resulted in a revision of the existing ammonite zonation scheme so far applied in Bulgaria and adjoining areas. A new biostratigraphic scheme, which bridges the western and eastern Tethys, is thereby proposed for the Lower Aptian of Bulgaria.The Upper Barremian Martelites sarasini Zone is characterized in its upper part by the Pseudocrioceras waagenoides Subzone in the shallow-water sections and by a horizon with Turkmeniceras turkmenicum in the deep-water settings. The Upper Barremian/Lower Aptian boundary is fixed by the first appearance of Paradeshayesites oglanlensis. For the Lower Aptian the following ammonite zones were established (from bottom to top): The Paradeshayesites oglanlensis Zone, the Deshayesites forbesi Zone (= formerly Paradeshayesites weissi Zone) including the Roloboceras hambrovi Subzone in the upper part, the Deshayesites deshayesi Zone including the Paradeshayesites grandis Subzone in the upper part and the Dufrenoyia furcata Zone. The Lower–Middle Aptian boundary has been defined by the appearance of species belonging to the genera Epicheloniceras and Colombiceras.The Lower Aptian ammonite faunas of Bulgaria, allow an interregional correlation with other areas of the Tethyan Realm. The presence of Turkmeniceras in the Upper Barremian enables a correlation with the Transcaspian region, whereas Roloboceras, Koeneniceras and Volgoceratoides found in the middle part of the Lower Aptian are more typical representatives of the ammonite faunas in northern Europe (England, Germany, Volga region).The analysis of the ammonite successions in combination with sedimentological observations enable us to conclude that the marls and marly limestones of the Lower Aptian studied here also cover the interval of the Oceanic Anoxic Event 1a. An interval of thin-laminated clays, rich in organic matter, was identified in the upper part of the D. forbesi Zone (Roloboceras hambrovi Subzone). This interval is characterized by a total lack of benthic faunas.     

Biostratigraphy and ammonites of the Middle Oxfordian to lowermost Upper Kimmeridgian in northern Central Siberia

The Middle Oxfordian to lowermost Upper Kimmeridgian ammonite faunas from northern Central Siberia (Nordvik, Chernokhrebetnaya, and Levaya Boyarka sections) are discussed, giving the basis for distinguishing the ammonite zones based on cardioceratid ammonites of the genus Amoeboceras (Boreal zonation), and, within the Kimmeridgian Stage, faunas–for distinguishing zones based on the aulacostephanid ammonites (Subboreal zonation). The succession of Boreal ammonites is essentially the same as in other areas of the Arctic and NW Europe, but the Subboreal ammonites differ somewhat from those known from NW Europe and Greenland. The Siberian aulacostephanid zones—the Involuta Zone and the Evoluta Zone—are correlated with the Baylei Zone (without its lowermost portion), and the Cymodoce Zone/lowermost part of the Mutabilis Zone (the Askepta Subzone) from NW Europe. The uniform character of the Boreal ammonite faunas in the Arctic makes possible a discussion on their phylogeny during the Late Oxfordian and Kimmeridgian: the succession of particular groups of Amoeboceras species referred to successive subgenera is revealed by the occurrence of well differentiated assemblages of typical normal-sized macro and microconchs, intermittently marked by the occurrence of assemblages of paedomorphic “small-sized microconchs” appearing at some levels preceeding marked evolutionary modifications. Some comments on the paleontology of separate groups of ammonites are also given. These include a discussion on the occurrence of Middle Oxfordian ammonites of the genus Cardioceras in the Nordvik section in relation to the critical review of the paper of Rogov and Wierzbowski (2009) by Nikitenko et al. (2011). The discussion shows that the oldest deposits in the section belong to the Middle Oxfordian, which results in the necessity for some changes in the foraminiferal zonal scheme of Nikitenko et al. (2011). The ammonites of the Pictonia involuta group are distinguished as the new subgenus Mesezhnikovia Wierzbowski and Rogov.     

Mathesia darderi (Astre) (Bivalvia, Hippuritoidea, Monopleuridae): Morphological, biogeographical and ecological changes in the Mediterranean domain during the late Barremian-Albian

Mathesia darderi, a slender cylindrical monopleurid genus, formerly documented from the late Aptian-Albian of Spain, France, Tunisia, Algeria, Egypt and Turkey, has been discovered in the upper Barremian and the lower Aptian of Bulgaria and Spain. Notwithstanding some morphological changes, Barremian-lower Aptian forms and those of the upper Aptian-Albian possess the same myocardinal organisation and the same microstructural attributes. The inner shell margin of the right valve displays scalloped, festooned, tubular and vermiform microstructures. The most prominent evolutionary trait of M. darderi is the increase in body size through time. A statistical analysis of size distributions show that populations of the late Barremian-early Aptian, and the late Aptian, and those of the early to middle Albian, are significantly different; a pattern which has a biostratigraphic potential. Ecological changes through time are expressed by a displacement of communities from the central/distal part, to the proximal part of carbonate platforms. M. darderi is present locally in the upper Barremian-lower Aptian, and has its major spreading over the European and Arabo-African margins of the Mediterranean Tethys during the Clansayesian-lower to middle Albian. The disappearance of the species at the Middle-Upper Albian boundary, correlates with a critical, spatial reduction of carbonate platforms.     

Barremian–Albian (Early Cretaceous) ammonite faunas of the Katsuuragawa Basin,southwest Japan

The Lower Cretaceous ammonite fauna of Japan was influenced by the Tethyan, Boreal and North Pacific realms with their oceanic current patterns and ammonite distributions. The hypothesis of oceanic circulation can be utilized to interpret the existence of the “Bering Strait” and the changing position of the “Boreal front,” that is the contact region of warm and cold-water masses. To understand such a system fully, a comprehensive understanding of the geographical distribution of ammonite faunas is required. The occurrence of twenty-five ammonite species, belonging to twenty genera, is confirmed in the Barremian to Albian of Japan. Of these, 24 species are described in this paper, including Barremites macroumbilicus sp. nov. The fauna can be divided into three associations, lower, middle, and upper, indicating late Barremian, late Aptian, and late Albian of the European standard zonation. The faunal characters suggest that the habitats of these ammonite faunas may have changed during later Early Cretaceous, with faunas characterizing three different environments, i.e., nearshore, intermediate, and distal shelf to upper slope setting. Ammonites of the lower association (late Barremian) are related to those of the Tethys, Boreal European, and circum-Pacific regions, and suggest that the Early Cretaceous Katsuuragawa Basin was deposited under the influence of currents from both high latitude and equatorial areas. The occurrence of Crioceratites (Paracrioceras) suggests that the Boreal European elements, including Simbirskites and Crioceratites (Paracrioceras), transited between Northwest Europe and Japan through the Arctic Sea, indicating that the Pacific Ocean was connected with the Arctic Sea at that time. The second association (late Aptian) is composed of ammonites of the Tethyan and circum-Pacific regions. As European Boreal ammonite taxa are absent in this association, it is concluded that the Pacific Ocean probably was not connected with the Arctic Sea at the time. Consequently, the “Boreal Front,” marking the contact between warm and cold water masses, was located at mid-latitude in the “Bering Strait” region during Barremian and subsequently moved northward during Albian. The uppermost ammonite association (late Albian) also consists of Tethyan and circum-Pacific taxa. Desmoceras (Pseudouhligella) poronaicum expanded its range with northward and eastward circulation of oceanic currents, suggesting the current must have represented a warm water-mass from the equator. The species subsequently migrated from the southern Katsuuragawa Basin to the Hokkaido area during late to latest Albian.     

Aptian (lower Cretaceous) ammonite biostratigraphy of the Francisco Zarco Dam stratigraphic section (Durango State,northeast Mexico)

This work deals with the study of the ammonoid record of a classical stratigraphic section of Aptian age, the Francisco Zarco Dam (Durango State, northeast Mexico). A detailed biostratigraphical analysis considered 1281 specimens systematically sampled on a bed-by-bed basis. Taphonomy, taxonomic composition and paleocology of the studied assemblage, were also stressed. We formally propose an ammonite zonation for this section, which consists of two interval zones, Dufrenoyia justinae and Gargasiceras? adkinsi, one taxon range zone Caseyella aguilerae, and one biohorizon Huastecoceras trispinosoides. Among those biostratigraphic units, three are of new definition. Another topic discussed herein concerns to the problematic definition of the Lower-Upper Aptian boundary using the first stratigraphical appearance of the genus Epicheloniceras. From a paleoecological point of view, we recognized the ammonoid assemblage of the Francisco Zarco Dam section, to be representative of a distal platform environment. Finally, we compared the zonation built for this section with that considered the standard zonation for the Tethyan Domain, concluding that the differences are so substantial that an independent zonation for the Central Atlantic is necessary.     

Taxonomy and stratigraphy of late Barremian–Albian species of Horiopleura Douvillé (Hippuritida,Polyconitidae) of the Mediterranean and southwestern Asian regions

The genus Horiopleura Douvillé is restricted to the Mediterranean and southwestern Asian regions, and is represented by eight species developed during the late Barremian–Albian. This revision of species contributed to delineate generic and specific characters. The definition of species is based on a set of external and internal qualitative and quantitative characters, the key characters being size, radial bands, ornamentation and the presence of a left posterior canal. Morphometry documents the dimorphism of some species, corroborated by regression analysis. Horiopleura dumortieri is an upper Barremian–lowermost Bedoulian species from SE France, Horiopleura brevis n. sp. is a Bedoulian, Mediterranean species, extending to the lower Gargasian, Horiopleura gigantea (pro Monopleura) is a Bedoulian species, restricted the Ukrainian Carpathes. Horiopleura lamberti and H. almerae are Upper Aptian to lower Albian species, and are known from Southern Europe and North Africa. Horiopleura haydeni is essentially late Aptian and typifies the southwestern Asia “Yasin fauna”. Horiopleura distefanoi and H. gemmellaroi (both formerly placed in Polyconites) are restricted to the Albian of the peri-Adriatic regions and North Africa, the former being present in the Middle East. Multiple correspondence analysis using six categorical variables and their categorical states, applied to the comparison of species, tends to corroborate our overall taxonomic frame-work, and reveals three groups of species with distinctive ages.     

Planktonic foraminiferal biostratigraphy of the Upper Barremian and Aptian of Crimea

Reexamination of the Barremian–Aptian planktonic foraminifers from three sections (Verkhoirechie, mountain Krasnaya, and Marino) allowed the biostratigraphic scheme for Southwest and Central Crimea to be refined and updated. The following standard zones are recognized in the studied sections: Blowiella blowi (upper Barremian), Hedbergella excelsa (upper Barremian–lower Aptian), Leupoldina cabri (lower Aptian), H. luterbacheri, Globigerinelloides ferreolensis, Gl. barri, Gl. algerianus, Hedbergella trocoidea, Paraticinella rohri (upper Aptian). Beds with Hedbergella ruka are recognized in the B. blowi Zone. Foraminifers from the Partizanskoe section, representing the lower Aptian L. cabri and H. luterbacheri zones, are studied. The recognized strata are correlated with ammonite and nannoplankton zones and paleomagnetic data.     

Kimmeridgian (Late Jurassic) ostracods from Highway A16 (NW Switzerland): taxonomy, stratigraphy, ecology, and biogeography

Ostracods are a common microfaunal element of the Kimmeridgian of the Jura Mountains in NW Switzerland. The stratigraphical subdivision within the Kimmeridgian can as clearly be inferred from ostracods as it is the case from the ammonite biozonation. This proves the utiliy of the ostracod biozonation, especially where ammonites are not available or rare. The ostracod-bearing layers of the sequence under study (middle part of the Reuchenette Formation = Banné Member, Courtedoux Member and Lower Virgula Marls) have been deposited in waters with highly brackish to marine salinities (high in the pliohaline range to—predominanttly—brachyhaline according to the Venice System, Oertli 1964). From the base of the section (base of the Banné Member, high brachyhaline in average), salinities slowly decreased, with lowest salinities in the lower dinosaur track levels of Courtedoux Member (high pliohaline on average). They then increased again to higher salinities (high brachyhaline on average) above the upper dinosaur track levels of the Courtedoux Member and the Lower Virgula Marls (lowermost A. eudoxus Zone). These trends perfectly correlate with the increased occurence of ammonites above the upper dinosaur track levels. In terms of Kimmeridgian ostracod palaeobiogeography, the fauna of the NW Swiss Jura Mountains described in this work is most similar to the Aquitan and Paris Basins, a little less to Northern Germany, and even less (with not even half of the species in common) to Southern Germany. The NW Swiss Jura Mountains still belong to a largely boreally influenced “Western and Central European subprovince”, whereas Southern Germany (though located more to the north) was subjected to an enhanced tethyan influence.     

Inoceramid bivalves and biostratigraphy of the upper Albian and lower Cenomanian of the United States Western Interior Basin

Inoceramid bivalves of the upper Albian and lower Cenomanian of the United States Western Interior are revised, Eleven species-level taxa and three genera are described. Two new species, Gnesioceramus mowriensis, characterizing the Mowry Shale of the early, but not the earliest, Cenomanian, and Posidonioceramus merewetheri, of the lower Cenomanian, and on new genus, Posidonioceramus, are recognised. The Western Interior inoceramid species from this interval are strongly endemic and are not good tools for long-distance correlations, although they are very effective in regional dating.In terms of the inoceramid biostratigraphy, middle and upper parts of the upper Albian can be referred to the Gnesioceramus Biozone, represented by G. comancheanus (Cragin) and G. bellvuensis (Reeside). These taxa are endemic to the Western Interior and some adjacent areas (Gulf Coast; Greenland?), but are closely allied to the cosmopolitan species, Gnesioceramus anglicus (Woods). At approximately the Albian-Cenomanian boundary, the endemic clade of ‘Inoceramus’ nahwisi appears, now referred to the newly erected Posidonioceramus, resulting in a distinct P. nahwisi biozone. This zone corresponds to the lower part of the ammonite Neogastroplites’ stratigraphic range. Gnesioceramids re-appear in the early Cenomanian. Close to base of the Cenomanian, for the first in the Western Interior, the genus Inoceramus, represented by Inoceramus irenensis Warren and Stelck, 1958, apparently immigrated into the Western Interior Basin.The Western Interior inoceramids do not allow for direct correlation to chronostratigraphic standard subdivision. The Albian-Cenomanian boundary, as earlier recognized on geochronologic correlations and confirmed, to some extent, based on ammonites, may approximately be located close to the appearance level of the genus Posidonioceramus.     

Integrated stratigraphy across the Aptian/Albian boundary at Col de Pré-Guittard (southeast France): A candidate Global Boundary Stratotype Section

The outcrop of the Marnes Bleues at the Col de Pré-Guittard, 11 km north of the village of Rémuzat in the Départment of Drôme in southeastern France is probably the most intensively studied succession spanning the Aptian/Albian boundary interval. Following the rejection of the proposed GSSP for the base of the Albian Stage (based on the first occurrence of the ammonite Leymeriellla tardefurcata in the section at Le Pillart, Tartonne, Alpes-de-Haute Provence), we re-visit the Pré-Guittard section. A new candidate GSSP defined by the first occurrence of the planktonic foraminifera Microhedbergella renilaevis Huber and Leckie, 2011 is here proposed. This first occurrence is placed in a 100 m section with 28 secondary markers, including calcareous nannofossils, planktonic foraminifera, palynomorphs, an inoceramid bivalve, ammonites, stable carbon isotopes, and local marker beds. The outcrop fulfils most of the physical criteria required of a Global Stratotype Section and Point.     

Report on the 5th International Meeting of the IUGS Lower Cretaceous Ammonite Working Group,the Kilian Group (Ankara,Turkey, 31st August 2013)

The 5th meeting of the IUGS Lower Cretaceous Ammonite Working Group (the Kilian Group) held in Ankara, Turkey, 31st August 2013, discussed the Mediterranean ammonite zonation, and its calibration with different ammonite zonal schemes of the Boreal, Austral and Central Atlantic realms. Concerning the standard zonation, that corresponds to the zonal scheme of the West Mediterranean province, some changes have been made on two stages. For the Valanginian, the Busnardoites campylotoxus Zone was abandoned; the upper part of the lower Valanginian is now characterised by the Neocomites neocomiensiformis and Karakaschiceras inostranzewi zones. For the upper Barremian, the former Imerites giraudi Zone is here subdivided into two zones, a lower I. giraudi Zone and an upper Martellites sarasini Zone. The I. giraudi Zone is now subdivided into the I. giraudi and Heteroceras emerici subzones, previously considered as horizons. The current M. sarasini and Pseudocrioceras waagenoides subzones correspond to the lower and upper parts of the M. sarasini Zone, respectively. The Anglesites puzosianum Horizon is kept. The Berriasian, Hauterivian, Aptian and Albian zonal schemes have been discussed but no change was made. The upper Hauterivian zonal scheme of the Georgian (Caucasus) region (East Mediterranean province) has been compared with the standard zonation. Discussions and some attempts at correlations are presented here between the standard zonation and the zonal schemes of different palaeobiogeographical provinces: the North-West European area for the Valanginian and Hauterivian, the Argentinean region for the Berriasian, Valanginian and Hauterivian, and the Mexican area for the Valanginian–Hauterivian and Aptian–lower Albian. The report concludes with some proposals for future work.     

High resolution ammonite–benthic foraminiferal biostratigraphy across the Aalenian–Bajocian boundary in the Lusitanian Basin (Portugal)

This work describes the ammonite and benthic foraminiferal assemblages recorded across the Aalenian–Bajocian boundary of the Serra da Boa Viagem II section, located about 6 km to the east of the Bajocian GSSP (Murtinheira, Portugal), and calibrated to the standard ammonite zonation previously established for the Lusitanian Basin. A total of 220 ammonite specimens referred to 30 fossiliferous levels were collected and identified throughout the section, enabling the recognition of the Concavum Zone (Concavum and Limitatum subzones) of the upper Aalenian, and the Discites Zone of the lower Bajocian. A total of 2356 foraminifers were obtained from the 16 samples collected along the section, corresponding to 4 suborders, 8 families, 16 genera and 44 species. The occurrence of Lenticulina quenstedti (Gümbel) has enabled the recognition of the Lenticulina quenstedti Zone, ranging from the Bradfordensis Zone (middle Aalenian) to the lower Discites Zone (lower Bajocian). The first record of Ramulina spandeli Paalzow, whose occurrence, up to now, was limited in the Lusitanian Basin to the Murtinheira section (the Bajocian GSSP), highlights the usefulness of the Ramulina spandeli Zone, with its lower boundary referred to the lower Discites Zone (lower Bajocian). Other bioevents displaying local, basinal or regional biostratigraphic interest have also been identified. The benthic foraminiferal record here presented, accurately calibrated with the ammonite record, aims at contributing to support the recognition of the Lenticulina quenstedti Zone and the Ramulina spandeli Zone as formal biostratigraphic units integrating the biostratigraphic scale based on benthic foraminifers for the Aalenian–Bajocian boundary in the Lusitanian Basin (Portugal). Copyright © 2014 John Wiley & Sons, Ltd.     

Stratigraphy and ammonite fauna of the upper Shemshak Formation (Toarcian–Aalenian) at Tazareh,eastern Alborz,Iran

With a thickness of 3900 m, the Tazareh section is one of the thickest developments of the Shemshak Formation in the Alborz range. It overlies with sharp and disconformable contact the limestones and dolomites of the Lower–Middle Triassic Elikah Formation and is topped, again with a disconformable contact, by the marls and limestones of the Middle Jurassic Dalichai Formation. The nearly exclusively siliciclastic succession represents a range of environments, from fluvial channels, flood plains, swamps and lake systems to storm-dominated shelf, and a comparatively deep marine and partly dysoxic basin. The segment of the section between 2300 and 3500 m is exclusively marine and contains a moderately diverse ammonite fauna, ranging from the Middle Toarcian to the Upper Aalenian. The ammonite fauna comprises 21 taxa, among them the new genus Shahrudites with two new species, Shahrudites asseretoi and S. stoecklini from the Middle Aalenian Bradfordensis Zone. The other ammonites from the Shemshak Formation at Tazareh (as elsewhere in North and Central Iran) are exclusively Tethyan in character and closely related to faunas from western and central Europe. An ammonite-based correlation of Toarcian–Aalenian successions of the eastern Alborz with time-equivalent strata of the Lut Block, part of the Central-East Iranian Microcontinent (ca. 500 km to the south), suggests a strong influence of synsedimentary tectonics during the deposition of the upper Shemshak Formation.     

Origin and evolutionary history of Anglonautilus (Nautilida,Cymatoceratidae) and a new species from the lower Aptian of Spain

The cymatoceratid nautilid genus Anglonautilus is distinguished from most other post-Triassic nautiloids by the occurrence of pronounced fold-like undulating ribs on the phragmocone and early body chamber. Anglonautilus praeundulatus n. sp. is described from the lower Aptian of eastern Spain. It is the first record of this genus from Spain and constitutes the oldest definite representative of the genus. This places the new species at the base of the evolutionary history of Anglonautilus. Its ornamental features confirm the previously assumed close relationship between Anglonautilus and Cymatoceras. An analysis of the ornamental pattern of all species hitherto referred to Anglonautilus indicates that there is an evolutionary lineage leading to the type species of the genus, A. undulatus, and a single successor (A. subalbensis). Several Late Cretaceous species hitherto identified as Anglonautilus (A. japonicus, A. mamiyai, A. suciensis) bear fold-like ribs superficially similar to typical representatives of Anglonautilus. The pattern of ornament in these taxa is very different at closer inspection, though. This suggests that these species developed independently from Cymatoceras. The undulations present in these taxa are interpreted here as a result of convergent evolution. They are therefore excluded from the genus Anglonautilus and provisionally referred to as “Anglonautilus” spp.     

Early Berriasian ammonites from Shal, Talesh region (NW Alborz Mountains, Iran)

An early Berriasian (Berriasella jacobi Zone) ammonite fauna is described for the first time from the Alborz Mountains in northwest Iran. It has been collected from a section located near the village of Shal (Talesh region); in addition to rare phylloceratids, lytoceratids and Neolissoceras, the majority of ammonites belong to the neocomitid subfamily Berriasellinae. With the exception of a new genus and species, Taleshites fuersichi, these taxa are common in European and North African Tethyan successions. Associated calpionellids confirm the early Berriasian age of the ammonite-bearing levels.     

Calcareous phytoplankton from the Barremian/Aptian boundary interval in NW Germany

Two cored boreholes in the central part of the North West German Basin recovered a unique section of Upper Barremian to Lower Aptian strata. Calcareous nannofossils show a distinctive shift from boreal endemic assemblages in the Barremian to cosmopolitan ones in the Aptian. This onset of new cosmopolitan species (e.g., Chiastozygus litterarius, Flabellites oblongus, Rhagodiscus angustus, Braarudosphaera sp., Eprolithus sp.) is spread over an interval of 25m, starting well below the early Aptian "Fischschiefer", a dark laminated shale rich in organic matter. These changes in the composition of calcareous nannofossils indicate that major palaeoceanographic changes occurred before the deposition of the Fischschiefer.The distribution pattern of calcispheres allows the differentiation of two sedimentary successions, separated by the Fischschiefer. The lower succession, which includes the sediments below the base of the Fischschiefer, indicates a boreal-pelagic environment. The lower part of the upper successions which includes the Fischschiefer, represents warmer inner shelf conditions. This corresponds to the presence of rich Tethyan-derived nannoconid assemblages and the presence of the planktonic foraminifera Hedbergella in the Fischschiefer. The topmost part of the upper succession (middle Aptian, Hedbergella marl) suggests a change to a pelagic warmer water environment. Two new taxa are introduced: the calcareous dinoflagellate cyst Obliquipithonella laqueata n. sp. and the foraminiferan Choanaella fortunate n. gen. n. sp.     

Aptian综合年代地层事件:定量地层学的运用

以往发表的年代地层数据都是以Ma形式出现,以关键物种的首现面和末现面来界定的。一般而言,这些年龄是在用某种方法确定的某两个层的年龄基础上通过插值获得的,很大程度上仍然是估计值,而不是可以检验的科学结论。我们需要的是一个可以评价的、剖面重现的、标本重现的而且插值方法是经过严格检验的数据库。图形对比技术正是一个有用的方法,它可以综合众多剖面的分散分布的事件,并可以验证这些事件的顺序和年代。图形对比技术是一个定量的、非统计的方法,可以判别两个剖面之间存在的同时代关系。物种时限和非生物事件投点到X/Y图解上,时限可以通过对比线综合到单个的时间尺度上。其他剖面的数据通过重复以上过程把每个剖面的数据综合进来。获得的时限通过评价有这些物种出现的图形解而得到验证。Aptian阶(约124~112 Ma)时期环境发生巨大变化,用来确定Aptian阶的许多关键生物事件和年代地层事件并不都出现在同一个剖面。因此,这些事件的相对年龄始终是不确定的,也不是相关的。两个事件被提出来用于界定Aptian 阶的底界,包括菊石Deshayesites tuarkyricus 和磁极性带CM0。然而这两个事件标准地层剖面相隔600 km。图形对比显示这两次事件出现在7 万年间隔内。通过这一方法,来自23 条Barremian Aptian Albian     

Latest Jurassic to earliest Cretaceous paleoenvironmental changes in the Southern Carpathians, Romania: regional record of the late Valanginian nutrification event

Fluctuation in calpionellid, foraminiferal, and nannofossil diversity and abundance are documented in two successions located in the eastern part of the Upper Jurassic–Lower Cretaceous carbonate platform of the Southern Carpathian area, Romania. The lower part of the studied sections consists of upper Tithonian–upper Berriasian bioclastic limestones. This age is supported by the presence of the calpionellid assemblages assigned to the Crassicollaria, Calpionella, and Calpionellopsis Zones. Based on biostratigraphical data, a gap was identified within the uppermost Berriasian–base of the upper Valanginian (the interval encompasses the Boissieri, Pertransiensis, Campylotoxum, and lower part of the Verrucosum ammonite Zones). Hence, the upper Tithonian–upper Berriasian bioclastic limestones are overlain by upper Valanginian–lower Hauterivian pelagic limestones (the interval covered by the NK3B and NC4A nannofossil Subzones). A detailed qualitative and semiquantitative analysis of the nannoflora was carried out over this interval. To estimate the surface water fertility conditions, the nannoplankton-based nutrient index (NI) was calculated. The fluctuation pattern of NI allow us to recognize four phases in the investigated interval, as follows: (1) phase I (covering the lower part of the NK3B nannofossil Subzone and the upper part of the Verrucosum ammonite Zone, respectively) is characterized by low values of the NI (below 20%), by the dominance of the genus Nannoconus in the nannofloral assemblages (between 60–70%), and moderate abundance of Watznaueria barnesae (up to 23%), while the high-fertility nannofossils constitute a minor component of the assemblages; (2) phase II (placed in the NK3B nannofossil Subzone, extending from the top of Verrucosum ammonite Zone, up to the lower part of the Furcillata ammonite Zone) is characterized by increase of NI above 30%, a decrease of nannoconids (up to 50% at the top), while Watznaueria barnesae increases in abundance up to 27%. The fertility proxies (Diazomatolithus lehmanii, Zeugrhabdotus erectus, Discorhabdus rotatorius, and Biscutum constans) represent again a minor component of the recorded nannofloras (less than 7% in both sections), but they have an ascending trend; (3) phase III (which encompasses the boundary interval of the NK3B and NC4A nannofossil Subzones, corresponding to the upper part of the Furcillata ammonite Zone) contains higher NI values (over 35%, and up 52% towards the base of this phase), an abrupt nannoconid decrease (down to 20%), higher abundance of Watznaueria barnesae (over 30%), while the fertility nannofossils became an important nannofloral component, jointly amounting to almost 20%; (4) phase IV (identified within the NC4A Nannofossil Zone and corresponding to the boundary interval of the Furcillata and Radiatus ammonite Zones) is characterized by a decrease of NI to 25%, a recovery of the nannoconids up to 40%, a decline in abundance of Watznaueria barnesae to 25%, together with a pronounced drop of fertility taxa, which make together no more than 8%. We assume that maximum of eutrophication in the studied interval from the Southern Carpathians was in the Furcillata ammonite Zone. Notably, within the phases 2 and 3, the morphological changes identified in the benthic foraminiferal assemblages (the predominance of flattened morphologies, together with the presence of conical and trochospiral inflated forms), as well as the occurrence of the Zoophycos trace fossils and pyrite framboids, indicate dysaerobic conditions. In the Southern Carpathians, the late Valanginian–early Hauterivian biogeographical changes are coeval with the initiation of the carbonate platform drowning.