A 350,000-year climatic record from the loess sequence of Achenheim, Alsace, France

Up to now the best Quaternary climatic sequences come from oceanic isotope studies, but terrestrial sequences are also well known, usually for pollen or ice core data. A new sequence providing climatic information for the last 500,000 years has been studied in the loess series of Achenheim (Alsace), using the mollusc record in relation to other stratigraphical data. Mollusc assemblages are analysed using a multivariate method. The correspondence analysis used here allows us to explain, in ecological terms, the general variability of the Achenheim set. Most Pleistocene mollusc species have the advantage that modem individuals live in the same assemhlages. So, the known ecology and distribution of the modern molluscs allow us to conclude that the first two factors explain variations in temperature and moisture. Each loading, on a factor, of each association in its stratigraphical level contributes to characterizing the evolution of each climatic parameter through time. For the last five climatic cycles, these evolutions are expressed as a function of the depth within the series. As they are well preserved, the last three glacial cycles are studied in detail. They correspond to the last 350,000 years, are compared with SPECMAP data and particularly show correlations between continental and marine climatic indicators. The mollusc assemblages of the loess sequence also provide information on temperature and moisture conditions. The evolutions of each parameter are not identical during the last three cycles, indicating that the climatic history of one cycle cannot be transferred to the others. The mollusc assemblages also record the occurrence of an oscillatory system, especially during the Weichselian Upper Pleniglacial (isotopic stage 2) when it announces the Late Glacial variations. Similar oscillating excursions seem to have occurred during the older glacial stages.     

Last interglacial-glacial climatic cycle in loess-palaeosol successions of north-western France

The loess series at St. Pierre-les-Elbeuf and St. Sauflieu are key successions for the western European Quaternary stratigraphy. The present study proposes a detailed record of the last interglacial-glacial climatic cycle at St. Pierre and its integration into the synthetic pedosedimentary record of north-western France using detailed correlations with the type sections of St. Sauflieu and Achenheim. Finally, comparisons with the marine isotope, Greenland GRIP chronologies and dust records are proposed. At St. Pierre, the pedostratigraphic and sedimentological analyses (total iron, organic matter, carbonate, grain size), in association with low field magnetic susceptibility measurements, demonstrate that this loess succession records the major climatic events of the Upper Pleistocene. The basal soil complex at St. Pierre is similar to those from the main successions of North (St. Sauflieu) or Northeast France (Achenheim). It shows a Bt horizon of brown leached soil, a deeply reworked grey forest soil and two isohumic steppe soils separated by a non-calcareous loess layer. This loess level corresponds to the first aeolian event clearly observed in the succession and can be correlated with Marker II of the Central European stratigraphy located around the marine isotope stage (MIS) 5/4 boundary. The main aeolian sedimentation starts after the soil complex and ends with the top soil (brown leached soil). Finally, a good parallel is observed between the strongest peaks of the dust records of the ice cores and the main period of loess deposition in St Pierre-lès-Elbeuf occurring during MIS 2.     

Direct evidence of central European forest refugia during the last glacial period based on mollusc fossils

Although there is evidence from molecular studies for the existence of central European last glacial refugia for temperate species, there is still a great lack of direct fossil records to confirm this theory. Here we bring such evidence in the form of fossil shells from twenty strictly forest land snail species, which were recorded in radiocarbon-dated late glacial or older mollusc assemblages of nine non-interrupted mollusc successions situated in the Western Carpathians, and one in the Bohemian Massif. We proposed that molluscs survived the last glacial period in central Europe in isolated small patches of broadleaf forest, which we unequivocally demonstrate for two sites of last glacial maximum age.     

The Buzzard''s Roost and Eustis mollusc sequences: comparison between the paleoenvironments of two sites in the Wisconsinan loess of Nebraska, USA

We investigated the Upper Pleistocene mollusc sequence in the loess deposit of Buzzard's Roost, Nebraska, and compared the results with those obtained for Eustis, 50 km to the west. Both sequences show similarities in: (i) stratigraphy and magnetic susceptibility implying identical fluctuations in loess deposition, (ii) biostratigraphy which indicates a succession of cold-and-moist to cold-and-dry conditions, and (iii) an increasing trend in mollusc abundance and diversity of species characterizing a climatic and/or environmental improvement. We interpret the variations in both sequences as related to the advance of the James and Des Moines ice lobes towards both sites, which was synchronous with the retreat of the Laurentide Ice Sheet. The occurrence of hygrophilous species in the mollusc record, from the upper sequences of Buzzard's Roost, suggests a moister environment than in Eustis. Our preliminary comparison of both malacological sequences suggests that regional or local, rather than global, influence prevailed over the Great Plains between 20 and 12 kyr BP.     

Post‐glacial biota from the inner part of southwest Joseph Bonaparte Gulf

Seventeen vibrocores from the inner part of Joseph Bonaparte Gulf off northwestern Australia penetrate a range of marine and marginal‐marine sediments deposited in the post‐glacial transgression and highstand. Ranging from gravelly sand to fine silt, these sediments contain a diverse fossil biota dominated by molluscs and bryozoans, but also including ostracods and foraminifers. Minor components include solitary corals, echinoids, soft coral and sponge spicules, wood debris and bone fragments. The biota can be divided into five major marine or marginal‐marine environments (intertidal, lagoonal, estuarine, strandline and shelf) and one terrestrial (riverine) environment. The intertidal environment contains four sub‐assemblages (mangroves, salt marsh, mud flat and sand flat) and the shelf environment six sub‐assemblages (hard substrate inner shelf, sandy substrate inner shelf, muddy substrate inner shelf, epiphytic, inshore and oceanic). The most useful organisms for palaeoenvironmental reconstruction are bryozoans for differentiating various shallow‐marine substrates, and foraminifers and ostracods for defining water depths, euryhaline, freshwater and oceanic influences. Palynomorphs were the only microfossils capable of providing control on terrestrial environments. The scarcity of marine plankton and the dominance of terrestrial palynomorphs in these marine sediments provides a salutary warning of the dangers of relying on plant microfossils alone when no independent environmental data are available to test the interpretation. The mollusc and bryozoan biota in the inner part of Joseph Bonaparte Gulf superficially resembles the bryomol assemblage of cool‐water shelves. This biotic assemblage is the result of turbidity rather than water temperature. The turbidity suppresses the photosynthetic, zooxanthellate and hermatypic organisms allowing molluscs, bryozoans and other apparently cool‐water biotic elements to dominate.     

Biogenic Accumulations of Uranium in Recent Seas

The objective of this study is to determine the geochemical role of molluscs in the distribution of uranium in the marine ecosystem. Biogeochemical studies are carried out on recent mollusc shells from the Caspian Sea, Sea of Japan, Sea of Marmara, Aegean Sea, Black Sea, Mediterranean Sea, Baltic Sea, and Indian Ocean, which differ from each other in terms of physical, chemical, geographic, and geochemical characteristics. In this study, nine Gastropoda and fifty-four Pelecypoda shells of different species are analyzed to document variations of uranium in seasonal layers, which were formed by the seasonal carbonate-organic phase of molluscs during their entire lives. Shell used in this study principally comprises three layers: upper (outer) prismatic, middle prismatic, and inner (mother-of-pearl) layers. In addition, when possible, the head, the middle, and the lower parts of the shells are used for analyses separately. Also, the biological accumulation rate values for each different mollusc species rel     

Mollusc assemblages in palaeoecological reconstructions: an investigation of their predictive power using transfer function models

Horsák, M. 2010: Mollusc assemblages in palaeoecological reconstructions: an investigation of their predictive power using transfer function models. Boreas, Vol. 40, pp. 459–467. 10.1111/j.1502‐3885.2010.00195.x. ISSN 0300‐9483. Fossil mollusc assemblages are commonly used to reconstruct past environments, as their shells are abundant in various types of calcium‐rich deposits. However, it is impossible to exactly evaluate estimates derived from fossil data using directly measured factors. To assess the accuracy of environmental variables derived from mollusc species composition, two modern data sets (training and test), each consisting of 73 samples of treeless fen communities, were constructed along with known local and climatic variables. The main predictors of species composition were isolated using canonical correspondence analysis and forward selection with the Monte Carlo permutation test. The accuracy of prediction for those factors that were significant in the forward selection was studied via four commonly used transfer function models. Three independent gradients of species composition driven by calcium content, moisture and temperature were detected. The best predictions were found for variables that correlated with the main changes in species composition. The strongest correlation between the predicted and measured values of the test data set was observed for water conductivity (r=0.86), a good proxy of calcium content. The locally weighted–weighted averaging transfer function model performed best out of all the models for the majority of variables tested. Mollusc assemblages were found to be useful for estimating local environmental variables based on a given species composition. Along with the specific advantages of mollusc fossil material, there is much potential for the use of their fossil assemblages to reconstruct palaeoenvironmental variables using transfer function models calibrated from recent compositional data and directly measured factors.     

Terrestrial molluscan records of Weichselian Lower to Middle Pleniglacial climatic changes from the Nussloch loess series (Rhine Valley, Germany): the impact of local factors

Two malacological sequences sampled in loess sections P1 and P3 of Nussloch (Rhine Valley, Germany) provide the most complete and precise molluscan record of western Europe for the Weichselian Lower and Middle Pleniglacial from about 70 to 34 cal. kyr BP. Qualitative and statistical analyses were performed on 134 mollusc samples. In the most complete Lower Pleniglacial record (P1), malacofauna changes reflect three short phases of vegetation development and climatic improvement related to soils and probably interstadials. A steppe to herb/shrub tundra shift characterizes the Lower-Middle Pleniglacial transition and is followed in both malacological records by the same general environmental trend (decline in vegetation and humidity increase) ending with a new increase in temperature and vegetation cover at the top of P3. In the Middle Pleniglacial, the impact of each shorter climatic change on the malacofauna is less recognizable due to a higher sediment compaction and also to being differently recorded in both sequences as the local topography affects soil water resources, soil and vegetation development and malacofauna adaptation. A comparison shows that the western European biostratigraphical framework can thus be improved by coupling molluscan records from loess sections to pollen sequences.     

黄龙钙华210Pb计年与现代沉积的环境变化研究

本文以黄龙钙华岩芯为主要研究对象, 利用210Pb定年技术在测定年轻碳酸盐样品短时间尺度上的优势建立了研究区近100年来高精度的年代标尺及环境记录; 通过碳氧稳定同位素的分析, 讨论了钙华稳定同位素记录与器测气候数据的相关性特征, 并与其他定年技术所获得的结果进行了相互验证。研究区钙华δ18O在1908~2004年时段内的变化范围为?12.13‰~?10.42‰, 平均值为?11.23‰, 变化幅度达到了1.7‰。同属西南季风区、由于海拔效应和距离水汽来源远近不同等因素的影响, 在同时段内黄龙钙华δ18O的变化范围和变化幅度都比董歌洞和Kahf Defore洞的δ18O记录要大, 说明处于中纬度、高海拔的黄龙钙华相对于低海拔的董歌洞和Kahf Defore洞石笋对气候环境的响应可能更加敏感。     

The Early Pleistocene Praetiglian and Ludhamian pollen stages in the North Sea Basin and their relationship to the marine isotope record

In this paper, the currently accepted correlation of the Early Pleistocene Ludhamian stage of England with the Tiglian‐A sub‐stage of the Netherlands is challenged. Recent investigations of Early Pleistocene marine North Sea deposits from a borehole near Noordwijk (the Netherlands) yielded evidence from molluscs, dinoflagellate cysts and sporomorphs for an alternation of warm‐temperate and arctic intervals within the Praetiglian and Tiglian stages. Marine equivalents of the terrestrial‐based pollen sub‐stages Tiglian A and B have been recognised in the upper part of the sequence. A Praetiglian age can be assigned to the lower part of the sequence on the basis of mollusc analysis. Within the Praetiglian, an alternation of warm and cold phases has been recognised from both the dinoflagellate cyst and molluscan records. Three cold phases within the Praetiglian are tentatively correlated with marine isotope stages (MIS) 96–100. The molluscan assemblages provide evidence for climate forcing of the sea level: highest sea levels are reached in the warm‐temperate intervals. Within the Praetiglian, an interval with an acme zone of the dinoflagellate cyst Impagidinium multiplexum, is correlated with the Ludhamian and tentatively linked to MIS 97 and/or MIS 96. The cold molluscan assemblages from the Noordwijk borehole include an acme zone of Megayoldia thraciaeformis, the first and only occurrence of this North Pacific bivalve in the North Sea Basin. Copyright © 2006 John Wiley & Sons, Ltd.     

A benthic index of environmental condition of Gulf of Mexico estuaries

An index was developed for estuarine macrobenthos in the Gulf of Mexico that discriminated between areas with degraded environmental conditions and areas with undegraded or reference conditions. Test sites were identified as degraded or reference based on criteria for dissolved oxygen levels, sediment toxicity tests, and sediment contamination. Discriminant analysis was used to identify a suite of measures of benthic community composition and diversity that would most successfully distinguish degraded from undegraded sites. The resultant benthic index was composed of a linear combination of three factors: the Shannon-Wiener diversity index, the proportion of total benthic abundance as tubificid oligochaetes, and the proportion of total benthic abundance as bivalve molluscs. This index was used to evaluate the spatial patterns of degraded benthic resources in the Gulf of Mexico.     

Western Palaearctic palaeoenvironmental conditions during the Early and early Middle Pleistocene inferred from large mammal communities,and implications for hominin dispersal in Europe

Large-scale fluctuations in global climate and resulting changes in ecology had a profound effect on human evolution and dispersal. Though hominin remains are scarce, studies focussing on the more abundant records of fossil land mammal communities can contribute greatly to our knowledge of the palaeoenvironmental circumstances that influenced and directed the global spread of hominins. To produce a comprehensive and accurate account of the evolution of western Palaearctic habitat diversity between 2.6 and 0.4 Ma BP, information generated from large mammal communities from 221 key sites has been included in this study.The palaeoecological conditions of the western Palaearctic during the Early and early Middle Pleistocene were principally controlled by the following key factors: (1) a widespread trend of temperature decrease, (2) the periodicity of the global temperature record, (3) the intensity of single climatic stages, (4) the temporal pattern of climatic variation, (5) geographical position, and (6) the distribution of continental water resources. A general picture of the evolution of western Palaearctic habitat diversity saw the replacement of extensive forested terrain by an alternating sequence of varied savannah-like and forested habitats during the 2.6–1.8 Ma span, as well as an alternation between different types of predominantly open habitats between 1.8 and 1.2 Ma. Both of these processes were governed by 41 ka temperature periodicity. During the 1.2–0.9 Ma time span, irregular climatic fluctuations were more common and habitat variability increased. The subsequent 0.9–0.4 Ma interval, a period controlled by 100 ka periodicity, was by comparison more stable, with longer climatic cycles alternating between open and forested landscapes. During the entire Early and early Middle Pleistocene, assemblages of large mammal communities reveal a distinct trend of decreasing continentality between Eastern and South-Eastern Europe on the one hand, and South-Western and North-Western Europe on the other. This trend was due to the effect of the Atlantic Ocean, while in Southern Europe the relatively low continentality was balanced by influences from the Mediterranean Sea.When plotted against evidence of hominin occurrence, the data on western Palaearctic habitat diversity inferred from large mammal communities indicate clear environmental stimuli for the earliest human dispersal in Europe. These are: (1) a wide range of habitats, implying a high diversity of resources; (2) mild climates with low seasonality, implying a lack of strong environmental fluctuations. Around 1.8 Ma at the latest, hominins of African origin entered the western Palaearctic for the first time, taking advantage of the diversity of habitats and resources, particularly along large river systems. Their subsequent westward spread between 1.7 and 1.3 Ma was restricted to Mediterranean-influenced areas, which offered a high variability of habitats and relatively low seasonality. The increase in environmental diversity, which occurred from 1.2 Ma onwards, opened up South-Eastern and Eastern Europe for hominin occupation. According to the available records, North-Western and Central Europe were initially colonized during late Early to early Middle Pleistocene interglacials, when these regions experienced periods of low seasonality and considerable habitat diversity.     

Stable isotope fractionation between mollusc shells and marine waters from Martinique Island

Forty-nine aragonitic and calcitic shells from 14 species of marine tropical molluscs (Bivalvia, Gastropoda, Polyplacophora) and ambient waters from Martinique have been analyzed for their carbon and oxygen isotope compositions. Mineralogy of shells was systematically determined by Raman spectroscopy that reveals composite shell structures and early processes of diagenetic alteration. In mangrove, brackish waters result from the mixing between 89±1% of seawater and 11±1% of freshwater, a hydrological budget quantified by both oxygen isotope and salinity mass balance calculations. Mollusc shells from the mangrove environment (S=31‰; δ¹⁸O=0.5‰) are characterized by mean δ¹³C values (−1.2‰) lower than those (+2.6‰) living in the open sea (S=35‰; δ¹⁸O=1‰). These low carbon isotope compositions result from the oxidation of organic matter into bicarbonate ions used in the building of mollusc shells. The oxygen isotope compositions of the studied mollusc species are mainly controlled by the temperature and composition of seawater whereas the role of the so-called “vital effects” is negligible. Contrasting with carbon isotopes, variability in the δ¹⁸O values among and within species of mollusc shells is very low (1σ=0.15) for a given littoral environment. Using ambient temperatures of seawater (28-30 °C), oxygen isotope fractionations between all studied living species and environmental waters match those extrapolated from the fractionation equation established for molluscs by Grossman and Ku [Chem. Geol., Isot. Geosci. Sect. 59 (1986) 59] in the range 3-20 °C. By analyzing calcite and aragonite layers from the same shell or by comparing shells from different species living in the same environment, there is no evidence that oxygen isotope fractionation between aragonite and water differs from that between calcite and water. On the basis of these results, we conclude that the oxygen isotope compositions of shells from most fossil mollusc species are suitable to estimate past seawater temperatures at any paleolatitude.     

粘土矿物在气候环境变化研究中的应用进展

在古环境和古气候的重建中,粘土矿物和碳-氧同位素、孢粉、树轮、冰芯等代用指标一样具有同等重要的地位。在没有地质构造运动影响时,气候是影响粘土矿物形成的主要因素,沉积物中的粘土矿物可用来分析古气候和古环境。粘土矿物及其组合可以独立或作为主要指标反映气候和环境变化,并已应用于高分辨率的晚第四纪地层的气候环境研究中。粘土矿物在很短的时间内也会发生明显地变化,不仅可以反映千年-百年尺度的气候环境变化,而且也可以反映几十年尺度的古气候环境变化过程。粘土矿物与其它指标对同一气候环境响应的相位差有待深入研究。不同的半定量分析方法测量出的同一剖面中粘土矿物的含量变化趋势和特征具有一致性。     

Evolution of Chinese Neogene Rhinocerotidae and Its Response to Climatic Variations

Chinese Neogene Rhinocerotidae has quite a complete record and its temporal range is the Early Miocene to Late Pliocene. The samples include 25 species in 4 tribes of 2 subfamilies. They are used as a foundation for the study of the evolution of the family and its relation to climatic changes. Taxonomic diversity, new records and extinctions are estimated for each Chinese Neogene mammal faunal unit (NMU). The diversity of Chinese Rhinocerotidae varies noticeably throughout the Neogene and is recognized within five stages: the Middle Miocene and Late Miocene are stages of high diversity and the Early Miocene, early Late Miocene and Pliocene are stages of low diversity. Rhinocerotid diversity and morphology are closely related to environmental factors and particularly sensitive to changes in ambient temperature and humidity. The interpretation of climatic variation reflected in the evolution of Rhinocerotidae corresponds precisely with the conclusions drawn from other workers and provides new evidence fen     

Mollusc and brachiopod skeletal hard parts: Intricate archives of their marine environment

The biogenic carbonate hard parts of fossil bivalves, cephalopods and brachiopods are among the most widely exploited marine archives of Phanerozoic environmental and climate dynamics research. The advent of novel analytical tools has led many workers to explore non‐traditional geochemical and petrographic proxies, and work performed in neighbouring disciplines sheds light on the complex biomineralization strategies applied by these organisms. These considerations form a strong motivation to review the potential and problems related to the compilation and interpretation of proxy data from bivalve, cephalopod and brachiopod hard parts from the viewpoint of the sedimentologist and palaeoceanographer. Specific focus is on the complex biomineralization pathways of a given dissolved ion or food particle from its aquatic environment via the digestion and biomineralization apparatus in molluscs and brachiopods and its incorporation into a biomineral. Given that molluscs and brachiopods do not secrete their hard parts from seawater but rather from their mantle and periostracum, this paper evaluates differences and similarities of seawater versus that of body fluids. Cephalopods, bivalves and brachiopods exert a strong biological control on biomineralization that, to some degree, may buffer their shell geochemistry against secular changes in seawater chemistry. Disordered (amorphous) calcium carbonate precursor phases, later transformed to crystalline biominerals, may be significant in carbonate archive research due to expected geochemical offset relative to the direct precipitation of stable phases. A reasonable level of understanding of the related mechanisms is thus crucial for those who use these skeletal hard parts as archives of the palaeo‐environment. The impact of what is commonly referred to as ‘biological factors’ on the geochemistry of mollusc and brachiopod hard parts is explored for conventional isotope systems such as carbon, oxygen, strontium and traditionally used element to calcium ratios. In particular, the often used δ¹³C_carb or the Mg/Ca and Sr/Ca elemental proxies are fraught with problems. An interesting new research field represents the analysis, calibration and application of non‐traditional proxies to mollusc and brachiopod hard parts. Examples include the carbonate clumped isotope (Δ₄₇) approach and the analysis of the isotopes of Ca, Mg, N, Li, S or element to Ca ratios such as Li/Ca or B/Ca and rare earth elements. Based on considerations discussed here, a series of “do's and don'ts” in mollusc and brachiopod archive research are proposed and suggestions for future work are presented. In essence, the suggestions proposed here include experimental work (also field experiments) making use of recent archive organisms or, where possible, a reasonable recent analogue in the case of extinct groups. Moreover, the detailed understanding of the architecture of mollusc and brachiopod hard parts and their ultra‐structures must guide sampling strategies for geochemical analyses. Where feasible, a detailed understanding of the diagenetic pathways and the application of multi‐proxy and multi‐archive approaches should form the foundation of fossil carbonate archive research. The uncritical compilation of large data sets from various carbonate‐shelled organisms collected at different locations is not encouraged.     

陆生蜗牛化石与中国黄土古环境研究

陆生软体动物蜗牛是黄土地层中重要的环境指标,在黄土地层成因和古环境与古气候研究方面具有独到的优势。文章总结了近10多年来中国黄土陆生蜗牛化石研究的成果,包括黄土蜗牛化石的研究方法、蜗牛典型种属及组合的地理分布与生态环境等,侧重介绍了黄土高原不同时间尺度的蜗牛记录及陆生蜗牛化石在黄土古环境和古气候研究中的新进展。     

Sediment particle size distribution and its environmental significance in Lake Erhai, Yunnan Province

A closed or semi-closed plateau lake, whose sediment records can provide us with plenty of fine and high resolution information, is a sensitive indicator of climatic and environmental changes. During the reconstruction of various short-time-scale climatic and environmental changes, the geochemical records in plateau-lake sediments are superior to other natural files. Based on fine dissection of the vertical profile of sediment particle sizes, this paper reveals the quasi-periodical changes of sediment particle sizes, which indicates the quasi-periodical fluctuations of the regional climate. A synthetic analysis of multiple indexes shows that sediment particle size is a more sensitive and more effective index of climatic and environmental changes than other geochemical indexes. High content of >20μm sediment particles and low content of 2–10μm sediment particles indicate a warm-dry climate and conversely a cold-humid climate, and their ratio can be used as an effective index of climatic changes. The basic climate succession in the region of Lake Erhai is characterized as being alternatively warm-dry and cold-humid and it has been developing into a warm-dry climate as a whole. There exist at least 2 time-scale quasi-periodical fluctuations of the regional climate in Lake Erhai. At present, the region of Lake Erhai is at the end of the warm-dry period and at the beginning of a cold-humid period, so the temperature will go down and the water level will rise. This project was supported by the National Natural Science Foundation of China (Nos. 49894170, 49773207).     

临夏盆地晚新生代哺乳动物群演替与青藏高原隆升背景

临夏盆地的晚新生代沉积中富含哺乳动物化石,以晚渐新世巨犀动物群、中中新世铲齿象动物群、晚中新世三趾马动物群和早更新世真马动物群的化石最为丰富。晚新生代是青藏高原快速隆升的时期,临夏盆地的4个主要哺乳动物群在构造剧烈变化的背景下发生了显著的更替。通过对不同动物群所代表的生态特征的分析,恢复了临夏盆地晚新生代以来的气候环境演变过程:晚渐新世以温暖湿润的森林环境为主,间杂有一些开阔地带;中中新世的森林更加茂密,水体更加丰富;晚中新世演变为炎热半干旱的稀树草原环境,季节性变化加强;早更新世气候寒冷而干燥,并伴有显著的海拔升高。青藏高原在晚渐新世的隆升幅度还不足以阻挡大型哺乳动物在高原南北的交流,但到中中新世已成为明显的障碍,至晚中新世对动物迁徙的阻碍作用更加突出,而临夏盆地在早更新世已经达到相当大的高度,产生了一个高原或高山的动物群     

Climatic conditions recorded by terrestrial mollusc assemblages in the Chinese Loess Plateau during marine Oxygen Isotope Stages 12–10

Past analogs for our present interglacial have been sought for better understanding of our present and future climate. Marine Isotope Stage 11 (MIS 11) has long been considered to be a good candidate and has gained increasing attention in recent studies of marine and terrestrial records. In the Chinese loess–paleosol sequences, S4 soil, interpreted as equivalent to MIS 11, yields a very precise terrestrial paleoclimate record of environmental changes that prevailed during this period. In this study, two high-resolution terrestrial mollusc records from the Loess Plateau have been analyzed to characterize climate variability during the periods of MIS 12–10. The changes in environment and climate, indicated by the variations in abundance of land mollusc species, are thus documented. Our mollusc results show that the L5 loess, equivalent of MIS 12, developed under relatively cold and dry conditions. Climate during this period was not as severe as indicated in the marine isotopic records. An episode of warmer and more humid condition occurred at the middle stage of the MIS 12, reflecting a summer monsoon strengthened during the glacial interval. A similar scenario has been observed in the middle part of MIS 10. In addition, our mollusc records provide insight into the climate conditions over the Loess Plateau during MIS 11, a general warm–humid climate dominated during the formation of S4 soil. But at least four fluctuations occurred at Xifeng region, reflecting unstable climate conditions and regional climate differences within the Loess Plateau during this period. Our study shows that the early part of S4, spanning over 30 ka, was very warm and humid, while the late part was characterized by mild-cool conditions. Comparison of mollusc species compositions of both MIS 11 and Holocene intervals reveals different climate conditions. The early part of MIS 11 was warmer and more humid than the Holocene optimum period, while the late part of MIS 11 was similar or cooler. Our study indicates that the extent of warming during the Holocene in the Loess Plateau might be significantly less than the conditions that prevailed during the MIS 11 interglacial period.