Tethyan-to-boreal correlation in the Kimmeridgian using high-resolution sequence stratigraphy (Vocontian Basin,Swiss Jura,Boulonnais, Dorset)

Ammonite biostratigraphy plays a central role in the definition of Jurassic stratigraphy. Nevertheless, the strong provincialism of European ammonite species during the Kimmeridgian is a long-standing problem in correlation attempts between the boreal and Tethyan faunal realms. Moreover, the sequence-stratigraphic interpretations for northern and southern Europe given in the Jurassic chronostratigraphic chart of Hardenbol et al. in SEPM Publ. 60 (chart) (1998) are different. The present study aims to resolve this correlation problem in order to better understand the connections between the boreal and the Tethyan realms during the Kimmeridgian. A sedimentological and high-resolution sequence-stratigraphic interpretation is presented for two unpublished sections (Cras d’Hermont and Roche de Mars) in the northern Swiss Jura, where recently discovered ammonites display both boreal and Tethyan influences. Then, these sections are correlated with the same time interval in the central Swiss Jura and Vocontian Basin, which belong to the Tethyan realm. Lastly, a long-distance transect is constructed between the Vocontian Basin, Swiss Jura, northern France, and southern England, the last two areas being part of the sub-boreal realm. The main results of this work are that: (1) third-order depositional sequences, and also higher-frequency sequences, can be correlated from the Tethyan to the boreal realm; (2) the sequence-stratigraphic interpretation given by Hardenbol et al. in SEPM Publ 60 (chart) (1998) for northern Europe seems to be accurate and agrees with the sequence-stratigraphic framework established in the Swiss Jura; (3) the Late Kimmeridgian of the Swiss Jura displays boreal influences; (4) integrated high-resolution sequence-stratigraphic and cyclostratigraphic studies are a valuable approach for bridging the correlation gap between northern and southern Europe.     

Facies, cycles, and controls on the evolution of a keep-up carbonate platform (Kimmeridgian, Swiss Jura)

During the Late Jurassic, accelerated ocean-floor spreading and associated sea-level rise were responsible for a worldwide transgression, which reached its maximum in the Late Kimmeridgian. In many Western European basins, this major sea-level rise led to the formation of marly and condensed sections. In the Swiss Jura, however, a shallow carbonate platform kept growing and only subtle changes in the stratigraphic record suggest an increasingly open-marine influence. Field observations and thin-section analyses reveal that the central Swiss Jura was at that time occupied by tidal flats and by more or less open marine lagoons where shoals and bioherms developed. The evolution through time of sedimentary facies and bed thicknesses permits the definition of small-, medium-, and large-scale depositional sequences. The diagnostic features of these sequences are independent of scale and seem largely controlled by the Kimmeridgian second-order transgression. A high-resolution sequence-stratigraphic correlation with biostratigraphically well-dated hemipelagic and pelagic sections in the Vocontian Basin in France reveals that: (i) The most important increase in accommodation recorded in the Kimmeridgian of the central Swiss Jura occurs in the Eudoxus ammonite zone (Late Kimmeridgian) and corresponds to the second-order maximum flooding recognized in many sedimentary basins. (ii) The small- and medium-scale sequences have time durations corresponding to the first and second orbital eccentricity cycle (i.e. 100 and 400 ka, respectively), suggesting that sedimentation on the platform and in the basin was at least partly controlled by cyclic environmental changes induced by insolation variations in the Milankovitch frequency band. The comparison of the high-resolution temporal framework defined in the Swiss Jura and Vocontian Basin with the sequence-stratigraphic interpretation realized in other Western European basins shows that the large-scale sequence boundaries defined in the Kimmeridgian of the Swiss Jura appear in comparable biostratigraphic positions in most Western European basins. Discrepancies that occur are probably because of local or regional tectonics.     

Late Cretaceous ostracod fauna from the Shenjiatun section (Songliao Basin,Northeast China): Biostratigraphic and palaeoecological implications

This work provides the first detailed taxonomic study of ostracod species from the Shenjiatun section (Nenjiang Formation, Songliao Basin, northeast China). Ten species belonging to seven genera are recognized. At the species level, this ostracod fauna shows a high degree of endemicity. The ontogeny of two species is recognized in this study, and sexual dimorphism within Daqingella arca is unequivocally recognized for the first time. The ostracod fauna suggests that this fossil-bearing sedimentary succession should be assigned to members 3–4 (Campanian) of the Nenjiang Formation (mainly Member 3) instead of representing ‘Quaternary sediments’ as previously thought. On the basis of the ostracod fauna, Member 3 of the Nenjiang Formation at Shenjiatun represents a shallow-littoral, lacustrine depositional environment that, at least in the upper part of Member 3, potentially included less stable conditions characterized by smaller, ephemeral waterbodies and increased (non-marine, oligohaline) salinities through evaporation.     

New marine reptile fossils from the Late Jurassic of Poland with implications for vertebrate faunas palaeobiogeography

A new marine vertebrate assemblage from the Late Jurassic (late Kimmeridgian) at Krzy?anowice near I??a in the NE margin of the Holy Cross Mountains in Poland is described. This new locality is rich in fossils of coastal and pelagic reptiles. The most frequent fossils are plesiochelyid turtle shell fragments and pliosaurid skull bones and teeth. The Krzy?anowice vertebrate assemblage is similar to the Late Jurassic Boreal/Sub-Boreal localities of the Kimmeridge Clay in Great Britain and Svalbard Archipelago in the Arctic, in the presence of pliosaurids and long-necked plesiosaurids. However, plesiochelyid turtles and crocodylomorphs are similar to those from the Mediterranean/Sub-Mediterranean sites of the northern border of the Tethys Ocean, as, for example, in the Swiss Jura Mountains and Southern Germany. This unique composition of the Krzy?anowice vertebrate fauna demonstrates that, during the Late Jurassic this new locality was located in the transitional palaeobiogeographic line referred to in this paper as the “Matyja-Wierzbowski Line”. The new palaeobiogeographical reconstructions of Late Jurassic of Europe are based on the composition of the Krzy?anowice locality and other sites with similar turtle-pliosaurid faunas which formed a long-term, stable ecological sympatry in marine ecosystems of the European Archipelago.     

Ostracoda during the Early Aptian (Early Cretaceous) greenhouse period on the Isle of Wight, England

The earliest Aptian marine transgression of the Lower Cretaceous across southern England resulted in the collapse of the generally freshwater Barremian environment and the initiation of marine mileux. Salinities passed from fresh-oligohaline to meso- and pliohaline, reaching fully marine conditions during the obsoletum Subzone (P. fissicostus Zone). Newly formed environmental niches were rapidly occupied by ostracod associations. In the Isle of Wight, freshwater Cypridea-rich assemblages in the lower Shepherd's Chine Member (Vectis Formation) were gradually replaced by faunas dominated by Sternbergella cornigera, Mantelliana mantelli and Theriosynoecum fittoni. Marine taxa recorded from the Atherfield Clay Formation migrated predominantly from the Paris Basin and include Asciocythere albae, Schuleridea derooi, Neocythere gottisi, N. bordeti, Cythereis geometrica, Cytheropteron stchepinskyi and Protocythere croutesensis.     

Biostratigraphy and ammonites of the Middle Oxfordian to lowermost Upper Kimmeridgian in northern Central Siberia

The Middle Oxfordian to lowermost Upper Kimmeridgian ammonite faunas from northern Central Siberia (Nordvik, Chernokhrebetnaya, and Levaya Boyarka sections) are discussed, giving the basis for distinguishing the ammonite zones based on cardioceratid ammonites of the genus Amoeboceras (Boreal zonation), and, within the Kimmeridgian Stage, faunas–for distinguishing zones based on the aulacostephanid ammonites (Subboreal zonation). The succession of Boreal ammonites is essentially the same as in other areas of the Arctic and NW Europe, but the Subboreal ammonites differ somewhat from those known from NW Europe and Greenland. The Siberian aulacostephanid zones—the Involuta Zone and the Evoluta Zone—are correlated with the Baylei Zone (without its lowermost portion), and the Cymodoce Zone/lowermost part of the Mutabilis Zone (the Askepta Subzone) from NW Europe. The uniform character of the Boreal ammonite faunas in the Arctic makes possible a discussion on their phylogeny during the Late Oxfordian and Kimmeridgian: the succession of particular groups of Amoeboceras species referred to successive subgenera is revealed by the occurrence of well differentiated assemblages of typical normal-sized macro and microconchs, intermittently marked by the occurrence of assemblages of paedomorphic “small-sized microconchs” appearing at some levels preceeding marked evolutionary modifications. Some comments on the paleontology of separate groups of ammonites are also given. These include a discussion on the occurrence of Middle Oxfordian ammonites of the genus Cardioceras in the Nordvik section in relation to the critical review of the paper of Rogov and Wierzbowski (2009) by Nikitenko et al. (2011). The discussion shows that the oldest deposits in the section belong to the Middle Oxfordian, which results in the necessity for some changes in the foraminiferal zonal scheme of Nikitenko et al. (2011). The ammonites of the Pictonia involuta group are distinguished as the new subgenus Mesezhnikovia Wierzbowski and Rogov.     

High-frequency palaeoenvironmental changes on a shallow carbonate platform during a marine transgression (Late Oxfordian, Swiss Jura Mountains)

Based on a well-established bio- and sequence-stratigraphic framework, a narrow time window in the Bimammatum ammonite zone (Late Oxfordian) is investigated in six Swiss Jura sections representing a shallow-water carbonate platform. From the detailed facies and microfacies analysis of oncoid-rich (Hauptmumienbank Member) and ooid-rich (Steinebach Member) limestones, a microfacies classification is established, depositional environments are interpreted, and a depositional model for the Swiss Jura platform is proposed. The sequence- and cyclostratigraphic interpretation has been performed for the transgressive part of the medium-scale sequence Ox6+, independently for each section, with a very high time resolution at the scale of elementary depositional sequences. The good correlation of the elementary and small-scale sequences between the six studied sections and the similar number of elementary sequences in all sections strongly suggest that allocyclic processes were involved in their formation. The hierarchically stacked depositional sequences (small-scale and elementary sequences) result from orbitally controlled sea-level changes with periodicities of 100 and 20 kyr, respectively. Thickness variations in the correlated small-scale and elementary sequences imply variable sedimentation rates, probably resulting from differential subsidence due to the activity of tectonic blocks. The tectonically controlled platform morphology contributed significantly to the general pattern of depositional environments and, combined with high-frequency sea-level fluctuations, created a complex facies distribution in time and space on the Swiss Jura carbonate platform.     

Tithonian–Early Valanginian evolution of deposition along the proto-Caribbean margin of North America recorded in Guaniguanico successions (western Cuba)

In the Guaniguanico Mountains of western Cuba, the Late Jurassic–Early Cretaceous limestones occur in three stratigraphic successions, which have accumulated along the proto-Caribbean margin of North America. The Late Jurassic subsidence and shallow-water carbonate deposition of the Guaniguanico successions have no counterpart on the northeastern Maya block, but some distant similarities with the southeastern Gulf of Mexico may exist. Four facies types have been distinguished in the Tithonian–Lower Valanginian deposits of the Guaniguanico tectonic units. Drowning of the Late Jurassic carbonate bank of the Sierra de los Organos occurred at the Kimmeridgian/Tithonian boundary. During this boundary interval, sedimentation in the west Cuban area and southwestern margin of the Maya block (Mexico) has evolved in a similar way in response to a major second-order transgression.The Lower Tithonian ammonite assemblages of the Guaniguanico successions indicate, in general, the neritic zone. Presence of juvenile gastropods and lack of adult specimens suggest unfavorable environment for these molluscs, probably related to low oxygenation levels. The Early Tithonian transgressive phase terminated about the lower boundary of the Chitinoidella Zone. The Late Tithonian “regressive” phase is weakly marked, whereas the latest Tithonian–earliest Berriasian strata were deposited during a deepening phase. The latter transgressive phase has ended in the Late Berriasian Oblonga Subzone. We correlate the bioturbated pelagic biomicrites of the Tumbitas Member of the Guasasa Formation with a significant fall of the sea level during the latest Berriasian–Early Valanginian. The average sedimentation rate for the Tumbitas Member biomicrites was about three times faster than for the Berriasian Tumbadero Member limestones. Sedimentation rates for the Tumbitas Member and the Valanginian limestones at the DSDP Site 535 in the southeastern Gulf of Mexico were similar. In the Los Organos succession, the Late Valanginian transgressive interval is associated with radiolarian limestones and black chert interbeds in the lower part of the Pons Formation. In the Southern Rosario succession, the pelagic limestones pass into the radiolarian cherts of the Santa Teresa Formation indicating a proximity of CCD during Late Valanginian–Hauterivian times.     

Astronomical time scale for the Middle Oxfordian to Late Kimmeridgian in the Swiss and French Jura Mountains

Detailed investigation of facies and sedimentary structures reveals that, during the Middle Oxfordian to Late Kimmeridgian, the shallow carbonate platform of the Swiss and French Jura Mountains recorded high-frequency sea-level fluctuations quite faithfully. The cyclostratigraphic analysis within the established biostratigraphic and sequence-chronostratigraphic framework implies that the resulting hierarchically stacked depositional sequences formed in tune with the orbital cycles of precession (20 kyr) and eccentricity (100 and 400 kyr). The astronomical time scale presented here is based on the correlation of 19 platform sections and 4 hemipelagic sections from south-eastern France where good biostratigraphic control is available. The cyclostratigraphic interpretation suggests that the interval between sequence boundaries Ox4 and Kim1 (early Middle Oxfordian to earliest Kimmeridgian) lasted 3.2 myr and that the Kimmeridgian sensu gallico has a duration of 3.2 to 3.3 myr. The astronomical time scale proposed here is compared to time scales established by other authors in other regions and the discrepancies are discussed. Despite these discrepancies, there is a potential to estimate the durations of ammonite zones and depositional sequences more precisely and to better evaluate the rates of sedimentary, ecological and diagenetic processes. Editorial handling: Hanspeter Funk, Helmut Weissert, Stefan Bucher     

A complementary section for the proposed Toarcian (Lower Jurassic) global stratotype: The Almonacid de la Cuba section (Spain)

We present the biostratigraphy (ammonites, brachiopods, foraminifers, and ostracodes), lithostratigraphy, sedimentology, sequence stratigraphy, magnetostratigraphy, and isotope stratigraphy of the Almonacid de la Cuba section located in the Iberian Range, central-eastern Spain. This section, which contains a continuous and expanded record of the Pliensbachian-Toarcian boundary (Early Jurassic), has been proposed as a complementary section for the Toarcian GSSP. An excellent ammonite-based biozonation has been obtained. Four ammonite assemblages characterized by the presence of Pleuroceras, Canavaria, Dactylioceras (Eodactylites), and Dactylioceras (Orthodactylites) have been distinguished. The base of the Toarcian is located at level CU35.2, based on the first occurrence of Dactylioceras. The occurrence of taxa from the NW European and the Mediterranean provinces is useful to improve the correlation between both provinces. Foraminiferal and ostracode assemblages are rich and diversified and no significant biostratigraphic events take place at the Pliensbachian-Toarcian boundary. The magnetostratigraphic data presented here are the most complete record of reversals of the earth magnetic field for the Pliensbachian-Toarcian boundary. A good record of the onset of the positive δ¹³C excursion reported in the Lower Toarcian of many European sections has been obtained. Average paleotemperatures measured at the latest Pliensbachian Spinatum Biochron of about 12.5°C, recorded a marked increase of the seawater temperature which started during the Toarcian, reaching average temperatures of 16.7°C at the Tenuicostatum Biochron. The obtained ⁸⁷Sr/⁸⁶Sr values fully agree with the LOWESS calibration curve.     

Mapping the rise and demise of Urgonian platforms (Late Hauterivian - Early Aptian) in southeastern France and the Swiss Jura

Lower Cretaceous carbonate deposits historically called “Urgonian limestones” are widely exposed around the margins of the Vocontian basin in southeastern France and in the adjacent Swiss Jura. This paper presents the history of their rise, growth and sudden demise. Eleven maps were constructed for deposits ranging in age from the Late Hauterivian pro parte to the Early Aptian (Bedoulian) pro parte. Based on sequential interpretations, they illustrate the present geographical distribution of the inner platform facies (Urgonian limestones stricto sensu, with rudists), the outer platform facies (essentially bioclastic deposits) and the basinal facies (slope, hemipelagic, pelagic deposits). These maps depict only the final terms of each successive sequence (the late highstand intervals). Chronostratigraphy is constrained by ammonites found mainly in basinal deposits, by echinoids, by rudists and to a lesser extent by dinoflagellates and calcareous nannoplancton. Inner platform, outer platform and slope (talus) deposits are dated by rich assemblages of orbitolinids and dasycladalean algae. Currently 39 species of orbitolinids have been recognized and their ranges collated with those of the ammonites in the area.In the Jura and in Provence the oldest Urgonian deposits are dated early Late Hauterivian, thus showing the synchroneity of the onset of platform carbonates development on both the southern and northern margins of the basin. Thereafter, growth of the platforms led a clearly regressive shallowing-upward trend, resulting from a stepwise progradation toward the center of the Vocontian area, coordinated with cyclical exposures in the inner platform areas. The maximum reduction of the platform deposits occurred early in Late Barremian times, coeval with a noticeable turnover in the orbitolinids assemblages.Thereafter, carbonate platform deposition shifted toward the margins of the Vocontian basin. In Early Aptian time, a well-dated discontinuity of regional extent marks the sudden, almost synchronous disappearance of the Urgonian deposits.     

Magnetostratigraphy of the Maastrichtian continental record in the Upper Aude Valley (northern Pyrenees,France): Placing age constraints on the succession of dinosaur-bearing sites

The first detailed stratigraphic succession of the Upper Cretaceous continental record from the Upper Aude Valley (southern France) is presented together with a magnetostratigraphic study. The combined stratigraphy and magnetostratigraphy of the Marnes rouges inférieures Fm (Lower Red Marls), constrained by biochronological markers such as charophyte occurrence and revised dinosaur eggshells, results in a succession of fluvial red beds dated from chron C32n to the top of chron C31r. It implies an earliest Maastrichtian age close to the C32n.1n-C31r reversal for the majority of the dinosaur sites including Bellevue. In contrast, the upper Maastrichtian is likely represented by a short interval within the lacustrine-palustrine Calcaires et argiles de Vignevieille Fm (Vignevieille Limestones), or it might even not be recorded. The proposed age indicates that the marine to continental transition, as a result of the Late Cretaceous transgression, took place earlier in the north Pyrenean basin than in the southern area.     

Problems of Oxfordian and Kimmeridgian stratigraphy in northern Central Siberia (Nordvik Peninsula section)

The Oxfordian–Lower Hauterivian section of the Nordvik Peninsula (northern Central Siberia) is a reference for developing zonal scales for various fossil groups and improving the Boreal zonal standard. In the middle 1950s–late 1980s, it was studied extensively by geologists, stratigraphers, lithologists, and experts on various fossil groups. These studies yielded rich fossil and microfossil collections and a set of parallel zonal scales for various faunal groups. Recently, a new detailed ammonite zonation of the Oxfordian and Kimmeridgian units of this section has been proposed. These results contradict the previous biostratigraphic data on ammonites, foraminifers, and palynomorphs. In the present paper, all the biostratigraphic data on the Oxfordian and Kimmeridgian units of the Nordvik Peninsula (Cape Urdyuk-Khaya) and northern Central Siberia undergo a comprehensive analysis and comparison with those on the Boreal Realm. The ammonite-constrained stratigraphic position of the lower Upper Jurassic in the Cape Urdyuk-Khaya section is interpreted as Upper Oxfordian or Middle Oxfordian. In our view, this difference in the understanding is due to the misidentification of some Oxfordian ammonite forms. The zones based on other fossil groups (foraminifers, dinocysts) which were distinguished in the Upper Oxfordian and Kimmeridgian sections of the Nordvik Peninsula are well traceable circumarctically. Their stratigraphic position in various regions of the Northern Hemisphere is constrained by ammonites and bivalves. However, if we use the last alternative ammonite zonation of this section part, hardly explicable contradictions will appear in interregional foraminiferal and dinocyst correlations.     

辽宁省四合屯下白垩统义县组的兽脚类足迹化石和可能的造迹者

记述了辽宁省四合屯下白垩统义县组三个半恐龙足迹,归入似鹬龙足迹（亦译为跷脚龙足迹）一未定种（Grallator isp.）。这是义县组恐龙足迹化石的首次描述。该行迹至少由3个造迹者所造。从足迹推断恐龙体长1.51m,属于义县组兽脚类恐龙较为普遍的体长范围。重建了尾羽鸟(Caudipteryx)和中华龙鸟(Sinosauropteryx)的足部,前者的足迹轮廓与似鹬龙足迹未定种的吻合度超过后者。根据化石记录,似鹬龙足迹类型可能广泛存在于义县组的各种中小型兽脚类（驰龙类与伤齿龙类除外）中。     

Morphogenetic impact of microbial mats on surface structures of Kimmeridgian micritic limestones (Cerin, France)

For several years, palaeoecological research has been conducted on micritic limestones of late Kimmeridgian age in the southern Jura Mountains. The sedimentary environment is that of a lagoon with an irregular bottom, which was repeatedly exposed. Between two stages of lime deposition, a microbial mat grew over the muddy surface, giving cohesion to the sediment, restraining erosion and preserving fossil remains and reptile tracks. Various structures at the microbial mat surface can be observed: crescentic wavy, radial wavy, torn, petee and mixed structures. They imply the presence of desiccation periods and a slight bottom slope leading to a downward sliding of the microbial mat. Such features may also be generalized to explain superficial microbial structures of other shallow carbonate-mud environments.     

重庆市綦江县莲花保寨:中国古人与恐龙足迹共存 的直接证据

重庆市綦江县老瀛山莲花保寨除了大量多样化的恐龙足迹之外,还保存了完好的要塞结构及历代题刻。从古迹可知,莲花保寨最早可追溯到南宋宝祐四年(1256年),很可能是当地人躲避蒙元军队对尚属南宋统治下的蜀地的进攻之用。此后,清朝又在此留下多处古迹。这些古迹与大量(至少以5种不同方式保存的)恐龙足迹(尤其是鸭嘴龙类足迹)共存,构成了中国古人在恐龙足迹化石点上长时间生活(可能长达700余年)的直接证据。而莲花保寨的泥裂、鸭嘴龙类足迹和波痕被引申为荷叶叶脉、莲花和水环境,也成为“莲花保寨”得名的证据。这项研究表明恐龙足迹参与了中国部分古地名、民间传说的形成,这些古地名、民间传说或可成为寻找恐龙足迹的重要线索。     

New data on Upper Kimmeridgian-Tithonian biostratigraphy and ammonites of the eastern Crimea

Based on ammonites, Upper Kimmeridgian sediments are first established in the Crimean Mountains. The Kimmeridgian-Tithonian boundary recognizable in a continuous section is placed inside the Dvuyakomaya Formation of uniform largely clayey sediments. Assemblages of Kimmeridgian ammonites Lingulaticeras cf. procurvum (Ziegler), Pseudowaagenia gemmellariana Oloriz, Euvirgalithacoceras cf. tantalus (Herbich), Subplanites sp.) and Tithonian forms (?Lingulaticeras efimovi (Rogov), Phylloceras consaguineum Gemmellaro, Oloriziceras cf. schneidi Tavera, and Paraulacosphinctes cf. transitorius (Oppel) are described. A new biostratigraphic scheme proposed for the upper Tithonian-Berriasian of the Crimean Mountains includes the following new biostratigraphic units: the Euvirgalithacoceras cf. tantalus Beds of the upper Kimmeridgian, ?Lingulaticeras efimovi Beds of the lower Tithonian, and Oloriziceras cf. schneidi and Paraulacosphinctes cf. transitorius beds of the upper Tithonian. The middle Tithonian is proposed to consist of the fallauxi and semiforme (presumably) zones. The ammonities found determine the early Kimmeridgian-Berriasian age of the Dvuyakornaya Formation that is most likely in tectonic contact with the underlying Khutoran Formation.     

The palaeoenvironment associated with a partial Iguanodon skeleton from the Upper Weald Clay (Barremian, Early Cretaceous) at Smokejacks Brickworks (Ockley, Surrey, UK), based on palynomorphs and ostracods

In 2001 a partial skeleton of an Iguanodon was discovered in the Upper Weald Clay (Barremian, Early Cretaceous) at Smokejacks Brickworks near Ockley, Surrey, UK. When the dinosaur was excavated, a detailed stratigraphic section was logged and 25 samples taken for palynological and micropalaeontological (ostracod and megaspore) analysis, including a detailed sample set of the dinosaur bed itself. Qualitative and quantitative analysis of the palynoflora revealed rich and well-preserved non-marine assemblages of pollen and spores, including early angiosperms, and freshwater green algae. Four types of angiosperm pollen are described and assigned to the genus Retimonocolpites Pierce, 1961, but left in open nomenclature. Some marine elements such as dinoflagellate cysts are identified as the result of reworking of Middle and Upper Jurassic sediments. The pollen/spore assemblages depict a vegetational change from principally gymnosperm-dominated assemblages at the base to principally pteridophyte-dominated assemblages at the top of the section. The dinosaur bed shows a pteridophyte-dominated assemblage, with a significantly high amount of the freshwater green alga Scenedesmus novilunaris He Cheng-quan et al., 1992. Samples close to the dinosaur bed yielded the first useful ostracod finds from Smokejacks Brickworks: well-preserved assemblages containing Cypridea clavata (Anderson, 1939), Damonella cf. pygmaea (Anderson, 1941), Stenestroemia cf. cressida Anderson, 1971 and Stenestroemia sp. A, and fragments and damaged valves of a thin-shelled ostracod, possibly belonging to Mantelliana Anderson, 1966. Those identified as Cypridea clavata show a wide range of morphological variety and in opposition to Anderson's (1967, 1985) taxonomic scheme, which would assign them to up to five different taxa, they are considered to be intraspecific variants of a single species. The possibilities and limitations of age determination of the Wealden sediments using palynomorphs and ostracods are discussed; distinct forms of early angiosperm pollen, together with the ostracod fauna, are consistent with an early Barremian age. Pollen and spores are discussed in terms of their parent plants and the reconstruction of vegetation and palaeoclimate. Palynology and ostracods give evidence for temporary freshwater conditions at the time when the Iguanodon died and the carcase was buried.