Factors responsible for the differences in satellite-based chlorophyll a concentration between the major global upwelling areas

The aim of this study was to identify the factors responsible for the differences in chlorophyll a concentration (Chl-a) observed between the California, Canary, Humboldt and Benguela upwelling areas. Monthly climatologic values of Chl-a obtained from satellite images, covering the years 1998–2004, revealed that this pigment was higher in the Benguela system than in the other areas. Upwelling intensity, as derived from offshore Ekman transport computations, was higher in the Benguela and Humboldt regions and, for the same upwelling intensity, Chl-a was higher in Benguela than in the other regions. Upwelling intensity appears to be able to drive Chl-a densities through nutrient supply, as nutrients are correlated to offshore Ekman transport. A linear regression model including the fraction of sea surface over the shelf in each 1° × 1° box, nitrate, silicate, turbulence and variability of offshore Ekman transport explained the 84.8% of the variance in Chl-a among the areas. Differences in offshore Ekman transport explained the lower Chl-a observed in Canary and California and the higher Chl-a observed in Benguela and Peru-Humboldt. A narrow continental shelf and low water column stability also contribute to reducing phytoplankton pigment biomass in the Canary and California areas. The higher Chl-a values observed in Benguela compared to Humboldt-Peru are due to a wider extension of the continental shelf in the Benguela region.     

Production regimes in four Eastern Boundary Current systems

High productivity (maxima 3 gCm⁻² day⁻¹) of the Eastern Boundary Currents (EBCs), i.e. the California, Peru-Humboldt, Canary and Benguela Currents, is driven by a combination of local forcing and large-scale circulation. The characteristics of the deep water brought to the surface by upwelling favorable winds depend on the large-scale circulation patterns. Here we use a new hydrographic and nutrient climatology together with satellite measurements of the wind vector, sea-surface temperature (SST), chlorophyll concentration, and primary production modeled from ocean color to quantify the meridional and seasonal patterns of upwelling dynamics and biological response. The unprecedented combination of data sets allows us to describe objectively the variability for small regions within each current and to characterize the governing factors for biological production. The temporal and spatial environmental variability was due in most regions to large-scale circulation, alone or in combination with offshore transport (local forcing). The observed meridional and seasonal patterns of biomass and primary production were most highly correlated to components representing large-scale circulation. The biomass sustained by a given nutrient concentration in the Atlantic EBCs was twice as large as that of the Pacific EBCs. This apparent greater efficiency may be due to availability of iron, physical retention, or differences in planktonic community structure.     

Observations of a cycle of intense coastal upwelling and downwelling at the research site of the Shirshov Institute of Oceanology in the Black Sea

The paper presents the results of joint analysis of the response of vertical temperature and current velocity profile distributions in the coastal zone of the Gelendzhik region of the Black Sea to strong wind forcing in the third ten-day period of September 2013. This forcing was caused by the propagation of an atmospheric cyclone, which first initiated coastal upwelling that was later replaced by downwelling. We formulate a criterion for the development of full coastal upwelling and demonstrate its efficiency. We assume that frequent events of incomplete coastal upwelling and downwelling are associated with changes in the water dynamics (variations in the intensity and direction of the alongshore current) generally not related to local wind forcing.     

CO2 outgassing off central Chile (31–30°S) and northern Chile (24–23°S) during austral summer 1997: the effect of wind intensity on the upwelling and ventilation of CO2-rich waters

The distribution of pH and alkalinity has been used to calculate the distribution of total inorganic carbon (TC) and fugacity of carbon dioxide (fCO₂) in the upper 200 m of the water column in coastal upwelling areas off northern Chile (23–24°S, near Antofagasta) and central Chile (30–31°S, near Coquimbo) during austral summer 1997. In these upwelling areas, colder surface waters were oxygen poor and strongly CO₂ supersaturated (100% near Antofagasta and 200% near Coquimbo), although below the pycnocline the CO₂ supersaturation invariably exceeded 200% in both areas. The larger surface CO₂ supersaturation and outgassing at 30°S were associated with stronger winds that promoted the upwelling of denser water (richer in CO₂) as well as a higher air–sea CO₂ transfer velocity. The consistent decrease in intensity of the southerly winds (as derived from NSCAT scatterometer data) from 30–31°S to 23–24°S suggests a corresponding decline in the intensity of the CO₂ outgassing due to upwelling. Additionally, we suggest here that the intensity of the local upwelling forcing (i.e. alongshore–equatorward winds) plays a role in determining the water mass composition and phytoplankton biomass of the coastal waters. Thus, while deep upwelling of salty and cold water resulted in high fCO₂ (up to 1000 μatm) and very low phytoplankton biomass (chlorophyll a concentration lower than 0.5 mg m⁻³), the shallow upwelling of less salty (e.g. salinity <34.5) and less CO₂-supersaturated water resulted in a higher phytoplankton biomass, which further reduced surface water fCO₂ by photosynthesis.     

Comparing trophic flows in the southern Benguela to those in other upwelling ecosystems

A balanced trophic flow model of the southern Benguela ecosystem is presented, averaging the period 1980–1989 and emphasizing upper trophic levels. The model is based largely on studies conducted within the framework of the Benguela Ecology Programme and updates the results of an expert workshop held in Cape Town in September 1989. Small pelagic fish other than anchovy Engraulis capensis and sardine Sardinops sagax, mainly round herring Etrumeus whiteheadi and mesopelagic fish, were important components of the food web in the southern Benguela. Severe balancing difficulties were encountered with respect to the semi-pelagic resources (hake Merluccius spp.) and demersal top predators (sharks), indicating the need for further research on the interaction of these groups with their ecosystem. The model is compared to other existing trophic flow models of ecosystems in major upwelling areas, i.e. the northern Humboldt Current (4–14°S), the California Current (28–42°N) and the southern Canary Current (l2–25°N), and to two independently constructed models of the northern Benguela ecosystem. These models are compared using network analysis routines of the ECOPATH software, focusing on the interactions between the five dominant fish species (anchovy, sardine, horse mackerel Trachurus trachurus capensis, chub mackerel Scomber japonicus and hake) that support important fisheries in all systems. The upwelling systems rank by size rather than species dominance. The ratio of catches and primary production differs between systems, partly because of differences in fishing regimes. Predation on the five dominant fish groups by other fish in the system was the most important cause of fish mortality in all models. Fishery catches are generally a larger cause of mortality for these groups than predation by mammals. The ecological cost of fishing appears to be comparatively low in the southern Benguela, because catches are low compared with the primary production, but also because the fishery is relatively low in the foodweb. However, in view of the very tight foodweb demonstrated in the model. it is likely that an increase in fishing pressure would cause severe trade-offs with respect to other components of the southern Benguela ecosystem.     

粤东及闽南近岸上升流对局地风场变化的响应

本文利用2010年6-7月的实测温盐、水位、海流等资料,结合风场数据,讨论了在台风影响较小的情况下,粤东及闽南近岸上升流对局地风场变化的响应特征,主要结论如下:（1）谱分析结果显示,沿岸风、水位、海流、近底层水温均具有3.5～4.0 d、5.0～5.5 d、8.3～9.0 d的波动周期,沿岸风的变化引起上升流强度在3～9 d周期上的波动;（2）上升流对局地风场变化的响应过程如下:利于上升流产生的局地风场发生变化时,沿岸风作用下产生的Ekman输运促使的上升流区水位的下降幅度发生改变,随即向岸方向的压强梯度力也发生变化,进而导致沿岸流及近底层向岸流的增强或减弱,而近底层向岸流强度的改变又会引起近底层水温的变化;（3）相关分析及交叉谱分析的结果表明,沿岸风的变化将在3 d以内影响上升流区近底层水温。以34 m向岸流代表近底层向岸流,则“沿岸风-水位-近底层向岸流-近底层水温”这一过程的响应时间依次为24 h、7 h、27 h左右。     

2006年夏季琼东、粤西沿岸上升流研究

利用2006年夏季广东、海南、广西近海的海洋水文调查资料和卫星遥感QuikSCAT风场资料分析琼东、粤西沿岸上升流的空间结构特征, 探讨风场、风应力旋度对上升流的影响以及上升流区水温、海流、海平面对上升流的响应。结果表明:琼东、粤西沿岸上升流区并非相互独立, 从10 m层以下已经连成一片。琼东沿岸上升流主要由夏季西南季风驱动而产生, 风应力旋度也有一定贡献。琼东沿岸上升流的强度比粤西强。琼东沿岸海域的上层海水(18 m以浅)以离岸运动为主, 中下层海水以向岸运动为主。上层的离岸流速大于中下层的向岸流速。琼东沿岸的上升流现象是间歇性的, 与沿岸风速强弱有关。琼东沿岸海域海平面的升降与上升流的强弱有良好的关系, 上升流的强弱滞后于海平面的升降约1～2 d。     

Upper-ocean thermal structure and heat content off the US West Coast during the 1997-1998 El Niño event based on AXBT and satellite altimetry data

During the 1997/1998 El Niño event, extensive oceanic temperature profiles were taken off the coast of California in January and February 1998 using Airborne Expendable Bathythermographs (AXBTs). These AXBT measurements are compared with altimetry-based upper-ocean temperature estimates using TOPEX and ERS satellite altimetry data. The altimetry-based temperature estimates are well correlated with the AXBT data, in particular when combining the two satellite data sets together to form a blended altimeter temperature estimate. Both the AXBT and altimetry data show that the nearshore coastal El Niño signal differed from that further offshore. The AXBT data show that near shore, the warm anomalies extended to much greater depths and had greater amplitude. A time series of the satellite-derived layer-averaged temperatures, averaged separately over the nearshore and offshore halves of the AXBT analysis domain, also shows a larger El Niño signal in the nearshore half. The role of local atmospheric forcing of the coastal oceanic temperature anomalies is analyzed using NCEP reanalysis and coastal upwelling data sets. The forcing terms include Ekman pumping, radiation, surface heat fluxes, precipitation, and alongshore wind stresses that drive coastal upwelling (expressed as a coastal downwelling index, CDI). The temperature forcing from all of the terms except the CDI anomalies are small. The CDI anomalies can explain most of the slowly varying temperature changes that occur near the coast during a two-year period spanning the El Niño event, as well as some of the larger amplitude, rapid (monthly) warming episodes that appear to be part of the El Niño signal. Several distinct rapid warming episodes, however, are not correlated with the CDI anomalies, and therefore we conclude that the nearshore El Niño signal originates from a combination of both a remote oceanic pathway and local atmospheric forcing.     

Air—sea CO2 fluxes in a coastal embayment affected by upwelling: physical versus biological control

Water column pCO₂ and air-sea CO₂ fluxes were studied during an 18-month period (May 1994–September 1995) in a coastal embayment affected by upwelling, located in the northwestern Iberian Peninsula (Ria de Vigo and adjacent shelf). Overall, the region acted as a net annual atmospheric CO₂ sink, with magnitude ranging from 0.54 mgC m⁻²d⁻¹ in the Ria estuary to 22 mgC m⁻²d⁻¹ offshore. During moderate upwelling and upwelling relaxation conditions the sampling area was a sink for atmospheric CO₂. By contrast, during winter conditions and during intense upwelling the flux reversed towards the atmosphere. The relative influence of physical and biological processes on pCO₂ was evaluated using two different approaches: firstly, statistical analysis of physico-chemical correlations, and secondly, a thermodynamic analysis in the oceanic CO₂ system. Both methods yielded consistent results, showing that the main processes controlling seasonal and spatial pCO₂ variability were the production and remineralization of organic matter, explaining ca. 70 % of the total variability. In the inner part of the embayment, air-sea CO₂ exchange was mainly modulated by CO₂ partial pressure gradient, whereas in the adjacent shelf, wind speed largely contributed to CO₂ fluxes between the ocean and the atmosphere.     

Sea Level Variations Associated with Upwelling/ Downwelling Along the West Coast of India

A three-dimensional numerical model is developed and used to study the coastal upwelling processes and corresponding seasonal changes in the sea level along the west coast of India. The upwelling and associated sea level variations are seen as a response of coastal ocean to pure wind stress forcing. The model is designed to represent coastal ocean physics by resolving surface and bottom Ekman layers as realistically as possible. The prognostic variables are the three components of the velocity field, temperature, salinity and turbulent energy. The governing equations together with their boundary conditions are solved by finite-difference techniques. Experiments are performed to investigate sea level fluctuations associated with the thermal response and alongshore currents of the coastal waters. The model is forced with mean monthly wind stress forcing of January, May, July and September representing northeast monsoon and different phases of the southwest monsoon. It is known from the observational study that the upwelling process reaches to the surface waters by May along the coastal waters of the extreme southwest peninsular region. The process is more intense in July compared to May and September and its strength decreases from south to north. However, during the northeast monsoon season, which is represented by January wind stress forcing in the model, downwelling is simulated along the coast. The model simulations of the coastal response are compared with the observations and are found to be in good agreement. The maximum computed vertical velocity of about 2.0 ×10 -3 cm s -1 is predicted in July in the southern region off the coast.     

Comparison of assemblages and some life-history traits of seabirds in the Humboldt and Benguela systems

There are 21 and 15 species of seabirds that breed in the Humboldt and Benguela upwelling systems respectively. Only two species of gull are common to both systems, one as an endemic subspecies to the Benguela system. Eleven species and two subspecies are endemic (or nearly so) to the Humboldt system; seven species and one subspecies to the Benguela system. Each system has an endemic penguin, sulid, cormorant and tern that feed mainly on anchovy Engraulis spp., sardine Sardinops sagax or both these fish. The Peruvian pelican Pelecanus thagus also feeds primarily on these prey items. A plentiful availability of food has resulted in many of these seabirds attaining high levels of abundance. For the four pairs of species that feed on anchovy and sardine, those in the Humboldt system all have a biology that enables them to increase more rapidly than their Benguela counter-parts. This reflects the higher frequency of environmental perturbations that depress seabird populations in the Humboldt system. In addition, both systems have a small endemic cormorant that feeds near the coast and a small endemic tern that breeds in the adjacent mainland desert and feeds at the sea surface. Several seabirds endemic to a system have no obvious ecological equivalent in the other system: the pelican, a diving-petrel, four storm-petrels and a gull in the Humboldt system; a cormorant and a gull in the Benguela system. Some species with tropical or subantarctic affinities breed at the boundaries of the systems. Others breed also in freshwater systems. The grey gull Larus modestus, which feeds in the Humboldt system, breeds in montane deserts.     

Production characteristics of upwelling systems and the trophodynamic role of hake

The productivities of the Benguela, Peru and California Current upwelling systems and the trophodynamics of the associated hake stocks are compared. Within these large systems, upwelling rate, primary production and fish production are all positively correlated, as would be expected on trophodynamic grounds. Existing measurements suggest that the Benguela and the Peru Current systems are 2–4 times more productive than the California Current in terms of primary production and support 2–10 times more fish per square kilometre. The Benguela and Peru Current systems are 2–3 times more efficient in production of fish and harvestable yield to man as the California Current. The California Current appears to be less efficient primarily because a greater fraction of the annual primary production is transported offshore from the major upwelling sites by the prevailing winds and meandering structure of the currents. The strong seasonal variability in upwelling off California probably also results in poor trophodynamic coupling between the herbivores and phytoplankton. Consequently, less of the phytoplankton is consumed and, potentially, more can be exported out of the system. Hake (Merluccius spp.) are opportunistic, apex predators of euphausiids and pelagic and demersal fish in the three upwelling systems. The striking dominance of euphausiids in the diet of hake in the California Current probably reflects the lesser abundance of demersal fish in that highly advective region.     

Sea Level Variations Associated with Upwelling/ Downwelling Along the West Coast of India

A three-dimensional numerical model is developed and used to study the coastal upwelling processes and corresponding seasonal changes in the sea level along the west coast of India. The upwelling and associated sea level variations are seen as a response of coastal ocean to pure wind stress forcing. The model is designed to represent coastal ocean physics by resolving surface and bottom Ekman layers as realistically as possible. The prognostic variables are the three components of the velocity field, temperature, salinity and turbulent energy. The governing equations together with their boundary conditions are solved by finite-difference techniques. Experiments are performed to investigate sea level fluctuations associated with the thermal response and alongshore currents of the coastal waters. The model is forced with mean monthly wind stress forcing of January, May, July and September representing northeast monsoon and different phases of the southwest monsoon. It is known from the observational study that the upwelling process reaches to the surface waters by May along the coastal waters of the extreme southwest peninsular region. The process is more intense in July compared to May and September and its strength decreases from south to north. However, during the northeast monsoon season, which is represented by January wind stress forcing in the model, downwelling is simulated along the coast. The model simulations of the coastal response are compared with the observations and are found to be in good agreement. The maximum computed vertical velocity of about 2.0 2 10 -3 cm s -1 is predicted in July in the southern region off the coast.     

Annual and interannual variability in the South-East Atlantic during the 20th century

Long time-series of sea surface temperature (SST) and pseudo wind stress (τ) in six areas in the South-East Atlantic are analysed as possible inputs into fisheries models. The areas encompass oceanic and coastal regimes. A clear seasonal signal is evident in all areas, with an amplitude in SST of 3–4°C in the upwelling areas and 5°C farther offshore and on the Agulhas Bank. Warming lags in the north and offshore by 1–2 months. Monthly variability is highest in the upwelling areas. An increasing trend in SST is suggested in all areas, the post-World War II era being about 1,0°C warmer than earlier periods. Some coherence occurs between areas at times, although at other times the anomalies are distinctly out of phase. There is a suggestion of an upward trend in the equatorward wind stress in some offshore areas after 1964, with a sharp change in 1975. Benguela Niños are evident in the environmental record and have a periodicity of around 10 years.     

南海东北部春季海表pCO_2分布及海-气CO_2通量

2013年南海东北部春季共享航次采用走航观测方式,现场测定了表层海水和大气的二氧化碳分压(pCO2)及相应参数。结合水文、化学等同步观测要素资料,对该海域pCO2的分布变化进行了探讨。结果表明,陆架区受珠江冲淡水、沿岸上升流及生物活动的影响,呈现CO2的强汇特征;吕宋海峡附近及吕宋岛西北附近海域受海表高温、黑潮分支"西伸"、吕宋岛西北海域上升流等因素影响,呈现强源特征。根据Wanninkhof的通量模式,春季整个南海东北部海域共向大气释放约4.25×104 t碳。     

琼东上升流的年际变化及长期变化趋势

全球变暖背景下沿岸上升流的年际变化是近年来的研究热点。本文基于1982—2012年的海表面温度和风场资料,分析了琼东上升流的强度和中心位置的年际变化规律以及沿岸风应力及其旋度的作用。结果显示,近30年来,琼东上升流强度总体减弱,相比于沿岸风应力,其变化与减弱的局地风应力旋度相关性更高;琼东上升流强中心位置最大概率发生在19.2°—19.3°N,与最大风应力旋度位置接近,且存在北移趋势。琼东上升流强度和位置的年际变化还存在周期约3年、5年和10年的本征模态,以3年周期变化为主。局地风应力旋度在琼东上升流的年际变化中发挥了重要作用。     

Corrections to ocean surface forcing in the California Current System using 4D variational data assimilation

The option for surface forcing correction, recently developed in the 4D-variational (4DVAR) data assimilation systems of the Regional Ocean Model System (ROMS), is presented. Assimilation of remotely-sensed (satellite sea surface height anomaly and sea surface temperature) and in situ (from mechanical and expendable bathythermographs, Argo floats and CTD profiles) oceanic observations has been applied in a realistic, high resolution configuration of the California Current System (CCS) to sequentially correct model initial conditions and surface forcing, using the Incremental Strong constraint version of ROMS-4DVAR (ROMS-IS4DVAR). Results from both twin and real data experiments are presented where it is demonstrated that ROMS-IS4DVAR always reduces the difference between the model and the observations that are assimilated. However, without corrections to the surface forcing, the assimilation of surface data can degrade the temperature structure at depth. When using surface forcing adjustment in ROMS-IS4DVAR the system does not degrade the temperature structure at depth, because differences between the model and surface observations can be reduced through corrections to surface forcing rather than to temperature at depth. However, corrections to surface forcing can generate abnormal spatial and temporal variability in the structure of the wind stress or surface heat flux fields if not properly constrained. This behavior can be partially controlled via the choice of decorrelation length scales that are assumed for the forcing errors. Abnormal forcing corrections may also arise due to the effects of model error which are not accounted for in IS4DVAR. In particular, data assimilation tends to weaken the alongshore wind stress in an attempt to reduce the rate of coastal upwelling, which seems to be too strong due to other sources of error. However, corrections to wind stress and surface heat flux improve systematically the ocean state analyses. Trends in the correction of surface heat fluxes indicate that, given the ocean model used and its potential limitations, the heat flux data from the Coupled Ocean–Atmosphere Mesoscale Prediction System (COAMPS) used to impose surface conditions in the model are generally too low except in spring-summer, in the upwelling region, where they are too high. Comparisons with independent data provide confidence in the resulting forecast ocean circulation on timescales ～14 days, with less than 1.5 °C, 0.3 psu, and 9 cm RMS error in temperature, salinity and sea surface height anomaly, respectively, compared to observations.     

Landward limit of wind setup on beaches

Open coast storm surge water levels consist of a wind shear forcing component generally referred to as wind setup; a wave setup component caused by wind-induced waves transferring momentum to the water column; an atmospheric pressure head component due to the atmospheric pressure deficit over the spatial extent of the storm system; a Coriolis-forced component due to effects of the rotation of the earth acting on the wind-driven alongshore current at the coast; and, if astronomical tides are present, an astronomical tide component. Astronomical tide is considered to be predictable and, therefore, not a meteorological driven component of storm surge although there may be interaction between the tide and meteorological driven water levels. Typically the most important component of storm surge on the US East Coast and Gulf of Mexico shorelines is the wind setup component. The importance of inland flooding due to the wind setup component of storm surge is considered herein with special reference to the effect of subaerial slope on inland flooding where three different linear slopes are considered and storm surge is calculated for the region above still water level, using an analytic solution. The present study findings show that the inland storm surge from the wind setup component can be of considerable importance and lead to significantly higher storm surges than found for storm surge at the still water level intersection of the beach/land. It is shown that mild slopes can lead to very high water levels at the land–water interface (i.e. above the still water level intersection of the beach).     

Response of the low-frequency fluctuations of sea level to atmospheric forcing along the coasts of the Huanghai Sea and the East China Sea

In this paper, the low-frequency fluctuations of sea level and their relationship to atmospheric forcing along the coasts of the Huanghai Sea and the East China Sea are studied. Spectrum analyses are made for the time series of daily mean sea level, atmospheric pressure and wind stress at seven coastal stations. It is found that at all the stations, the main part of the energy of the sea level fluctuations, within the (2-60)-day period, is concentrated on the (12-60)-day period band and that an obvious spectral peak appears at the 3-day period. Along the coast of the Huanghai Sea, variations in the sea level are greater in winter than in summer. In winter, along the coasts of the Huanghai Sea and the East China Sea there is a kind of sea level fluctuations propagating southwards. Among the many factors causing sea level variation, the most obvious one is atmospheric pressure, followed next by the alongshore wind stress.     

Transport and retention in an upwelling region: The role of across-shelf structure

The paradox of upwelling is the relationship between strong wind forcing, nutrient enrichment, and shelf productivity. Here we investigate how across-shelf structure in velocity and hydrography plays a role in the retention (inshore) and export (offshore) of particles such as nutrients, plankton and larvae. We examine the spatial structure of the coastal currents during wind-driven upwelling and relaxation on the northern Californian Shelf. The field work was conducted as part of the Wind Events and Shelf Transport (WEST) project, a 5-year NSF/CoOP-funded study of the role of wind-driven transport in shelf productivity off Bodega Bay (northern California) from 2000 to 2003. We combine shipboard velocity profiles (ADCP) and water properties from hydrographic surveys during the upwelling season to examine the mean across-shelf structure of the hydrography and velocity fields during three contrasting upwelling seasons, and throughout the upwelling-relaxation cycle. We also present results from two winter seasons that serve as contrast to the upwelling seasons.During all three upwelling seasons clear spatial structure is evident in velocity and hydrography across the shelf, exemplified by current reversals inshore and the presence of a persistent upwelling jet at the shelf break. This jet feature changes in structure and distance from the coast under different wind forcing regimes. The jet also changes from the north of our region, where it is a single narrow jet, adjacent to the coast, and to the south of our region, where it broadens and at times two jets become evident. We present observations of the California Under Current, which was observed at the outer edge of our domain during all three upwelling seasons. The observed across-shelf structure could aid both in the retention of plankton inshore during periods of upwelling followed by relaxation and in the export of plankton offshore in the upwelling jet.