古尔班通古特沙漠生境对植物群落格局的影响

This paper reports a geomorphologic landscape investigation, vegetation survey and soil sampling at 14 sites across the Gurbantunggut Desert between 87°37′09"-88°24′04"E and 44°14′04"-45°41′52′Nl. The study encountered 8 species of low trees and shrubs, 5 of perennial herbs, 8of annual plants and 48 of ephemeral and ephemeroid plants. These species of plants represent one-third of the species found in the Gurbantunggut Desert, and their communities make up a large proportion of desert vegetation with great landscape significance. In the investigation we found that the plant communities are accordingly succeeded with the spatial variation of macro-ecoenvironment.Using Principal Component Analysis (PCA) and Correlation Analysis (CA) we found that the micro-ecoenvironment heterogeneity of aeolian sandy soil′s physical and chemical properties such as soil nutrient, soil moisture, soil salt, pH etc. only impacted the diversity of herb synusia (PIEherb) of the desert, with a negative correlation. Meanwhile, the impact of microhabitat on the plant community pattern with an antagonistic interaction made vegetation′s eco-distribution in a temporary equilibrium.     

Impact of habitat heterogeneity on plant community pattern in Gurbantunggut Desert

1IntroductionStudy on the interrelationship between vegetation patterns and their habitat heterogeneity is important to recover and rehabilitate the desert vegetation, stabilize the desert ecosystem and prevent desert expansion. Domestic and foreign researchers have carried out many studies on this topic (Archer etal., 2002; Bolling etal., 2000; Chen etal., 2003; Dasti etal., 1994; Gu etal., 2002, Li etal., 2001; Li etal., 2001; Liang etal., 2003), which have used and refined statistic tech…     

古尔班通古特沙漠生境对植物群落格局的影响

沿87^o37'09"~88^o24'04"E纵穿古尔班通古特沙漠,对44^o14'04"~45^o41'52"N的范围进行11个样地的地貌景观调查、植被分布参数采集和土壤取样。研究区统计到的植物种数占古尔班通古特沙漠文献资料中植物种数的1/3,出现的植物群落构建了该沙漠大面积成片分布的荒漠植被景观。调查发现,随着宏观生境的空间变化,植物群落作相应的演替。在此基础上,利用因子分析 (PCA)和相关分析 (CA) 查明了沙漠风沙土理化属性,如土壤养分、盐分、水分、碱性 (pH) 等微生境的异质性仅对沙漠植被草本层片的多样性 (PIE草) 产生显著影响。同时,这些微生境因相互拮抗使沙漠环境处于暂时的脆弱平衡之中。     

古尔班通古特沙漠风沙土微量元素对植被格局的影响

横穿古尔班通古特沙漠,测度了沙漠植被盖度、物种多样性和丰富度等格局参数及微地形、土壤理化性质。在此基础上,借助ICP/MS仪测定了沙漠表层（0~10 cm）和次表层（10~30 cm）风沙土中Cu、Zn、Mo、Fe、Co、Ni、Mn、V、Se、B微量元素含量。利用t-检验和CCA排序的方法,分析了微量元素在风沙土中的空间分布特征和对植被格局的影响。结果显示,地势较高的东部Ni和Mn的含量高于其他地段。地势较低、土壤中水分、有机质、全盐含量较高的西部,Mo、Cu、Zn、Fe、Se含量较高。Fe的高含量值主要出现在分选差、有机质较多的风沙土中。另外,绝大多数微量元素在表层风沙土中的平均含量高于浅层。对比不同微量元素成分条件下的乔灌木和草本层片植物组成和结构特征,发现乔灌木层片基本格局所受影响不明显,草本层片则在沙漠生境下已形成的植物群丛内部发生分化,出现多个小的植物组合。     

古尔班通古特沙漠南缘荒漠化过程演变的景观格局特征分析

荒漠化过程是荒漠化研究的核心问题。以1977年MSS,1990年、2010年TM和2001年ETM+四个时期为遥感影像数据源,研究了近40 a古尔班通古特沙漠南缘荒漠化过程演变的景观格局特征。结果表明:(1)近40 a古尔班通古特沙漠南缘荒漠化过程演变经历了微弱逆转-加剧-逆转变化,荒漠化处于正逆交替动态变化中,总体上呈逆转趋势。(2)不同时期荒漠化过程演变格局特征呈波动性变化,以2001年为分水岭。近40 a来轻度和中度始终为主导作用,且有增强趋势。(3)年降水量与归一化植被指数(NDVI)相关系数达0.805,有明显正相关关系。年降水量对荒漠化具有显著作用,且其影响明显大于年均气温。(4)荒漠化过程景观格局特征对年降水量有明显响应。不同时期荒漠化过程演变中斑块密度随着降水量的增加总体呈减小趋势,年降水量达到184.4 mm,斑块密度趋于稳定,景观破碎度和景观异质性趋于稳定。随着降水的增加,轻度荒漠化类型破碎度呈逐渐减小趋势;中度荒漠化土地破碎度呈波动式变化;重度荒漠化破碎度波动下降趋势。(5)近40 a来气候变化是古尔班通古特沙漠的荒漠化与景观格局演化主要因素,但人为因素不容忽视,预示着未来干旱区荒漠化处于继续逆转趋势中。     

Distribution of plant species and species-soil relationship in the east central Gurbantunggut Desert,China

The distribution of plant species and relationships between species and soil factors in the east central part of Gurbantunggut Desert was studied to provide more insight into the flora and determine differences in vegetation across various parts of the desert. Two-way Cluster Analysis showed that the vegetation in the area could be divided into three groups, the first group was dominated by the shrub species, Ephedra przewalskii and the grass species, Carex physodes mainly in areas of flat grounds and gentle slopes; the second group was dominated by C. physodes, Artemisia songorica and A. xerophytica mainly on the slope of sand dunes and the third group was dominated by the shrub species, Haloxylon persicum mainly on the top of sand dunes. There was no difference in plant density between Groups 1 and 2 but there was a significant decrease in Group 3. Soil water under vegetation Group 3 was much lower than that in the other two groups at all soil depths. The EC, organic matter, total P and soluble Na, Ca and Mg varied very similarly with soil water. Canonical correspondence analysis (CCA) satisfactorily assessed the species-soil relations in the area. The distribution of plant species was strongly correlated with the soil factors of water content, organic matter, EC and nutrients. The variations in species occurrence explained by the three CCA axes were about 70%, indicating that some explanatory site variables may exist outside our studied parameters. Soil texture is suggested to be included in future studies to improve the explanation of CCA.     

中国古尔班通古特沙漠东部中心地区植物分布与植物—土壤关系(英文)

The distribution of plant species and relationships between species and soil factors in the east central part of Gurbantunggut Desert was studied to provide more insight into the flora and determine differences in vegetation across various parts of the desert. Two-way Cluster Analysis showed that the vegetation in the area could be divided into three groups, the first group was dominated by the shrub species, Ephedra przewalskii and the grass species, Carex physodes mainly in areas of flat grounds and gentle slopes; the second group was dominated by C. physodes, Artemisia songorica and A. xerophytica mainly on the slope of sand dunes and the third group was dominated by the shrub species, Haloxylon persicum mainly on the top of sand dunes. There was no difference in plant density between Groups 1 and 2 but there was a significant decrease in Group 3. Soil water under vegetation Group 3 was much lower than that in the other two groups at all soil depths. The EC, organic matter, total P and soluble Na, Ca and Mg varied very similarly with soil water. Canonical correspondence analysis(CCA) satisfactorily assessed the species-soil relations in the area. The distribution of plant species was strongly correlated with the soil factors of water content, organic matter, EC and nutrients. The variations in species occurrence explained by the three CCA axes were about 70%, indicating that some explanatory site variables may exist outside our studied parameters. Soil texture is suggested to be included in future studies to improve the explanation of CCA.     

古尔班通古特沙漠88°E沿线风沙土理化性状的纵向分异

纵穿古尔班通古特沙漠中部对风沙土理化性状分析结果表明:沙漠腹地风沙土含水量低于边部。风沙土颗粒物主要由极细沙-细沙-中沙组成,并且北部较中部和南部粗。个体沙垄垄间地风沙土φ1值最高,垄顶的平均粒径最大。北部风沙土分选性差,绝大多数为极正偏分布,峰态很窄。中部、南部风沙土分选性中等和较好,呈窄峰态和近对称的分布形式。沙漠腹地风沙土全盐含量均较低,pH值适中,二者和沙垄固定程度成一定的正相关。风沙土中有机质、N、P、K等营养物质含量较低。相比之下北部风沙土全K和有机质含量高于中、南部,而植被茂盛的垄间地和垄坡有机质和全N含量一般高于固定、半固定垄顶。局地气象条件、微地形地貌以及植被状况的差异是引起沙漠风沙土上述理化性状空间分异的重要因素。     

古尔班通古特沙漠纵向沙垄植被空间异质性

在古尔班通古特沙漠南部纵向沙垄上按5 m×10 m的网格测度植被样方100个,采集浅层风沙土样品184件。调查发现,乔灌木层片和草本层片的分布及相互关系明显受沙垄地形影响,并且在干旱环境下种群间的竞争和空间上的互补使两层片生态优势度具有显著的相关性。地统计分析结果显示,纵向沙垄上植被空间异质性表现为木本层片的物种多样性呈斑块状分布,草本层片物种多样性呈现平行沙垄的带状结构和斑块结构叠加的特点,而植被盖度则在垂直沙垄走向的方向上具有强烈的梯度性变化。和沙垄土壤理化性质空间格局特征相对比,植被盖度及物种多样性与风沙土浅层含水量在自相关空间（变程）及空间异质性程度上比较接近。这是纵向沙垄小生态系统中草本植物和浅层土壤水分耦合关系在空间上的反映,也是木本植物对水分和土壤资源在小尺度（m）上竞争及中小尺度（10～100 m）上互补的结果。     

古尔班通古特沙漠南缘干季土壤水分含量空间异质性

土壤水分含量是荒漠植被发育的主要制约因子。对古尔班通古特沙漠南缘个体、群落、丘间地0~100 cm干季土壤水分含量的空间异质性进行分析。结果表明：（1）同尺度下,土壤水分含量随土层深度增加呈显著升高趋势,且表层土壤水分含量的变异系数高于其他土层。（2）个体尺度,梭梭树干基部周围土壤水分含量在垂直和水平方向存在格局分异。垂直方向,土壤水分含量随土层深度增加呈升高趋势。水平方向,随距树干基部距离增加,坡顶土壤水分含量呈升高趋势,坡中和坡底土壤水分含量呈降低趋势,但没有显著差异。（3）梭梭群落尺度上土壤水分含量异质性较强,且呈斑块状分布。（4）地形是影响丘间地尺度土壤水分含量空间分异的主要因素,形成坡底土壤水分含量最高、坡中次之、坡顶最低的空间分布格局。     

古尔班通古特沙漠地表辐射收支特征

古尔班通古特沙漠是中国最大的固定、半固定沙漠.利用2017年该沙漠克拉美丽站辐射资料,分析了古尔班通古特沙漠不同时间尺度和不同天气条件下的地表辐射变化特征.结果表明:(1)不同月份沙漠辐射收支各分量月平均日变化均呈单峰型,但极值大小及出现时间存在差异.各分量曝辐量季节变化明显:太阳总辐射表现为生长期(4-9月)＞积雪期...     

古尔班通古特沙漠南缘丘间地梭梭群落蒸散特征

根据2016年古尔班通古特沙漠南缘丘间地梭梭生育期定点观测的土壤水分、气象要素等资料,基于水量平衡原理估算了梭梭生育期蒸散量,分析了蒸散变化规律。结果表明：（1）在梭梭生长季,降雨量为206.7 mm,降雨分布不均,梭梭萌发期,降雨量最多;梭梭生长旺盛期,月降雨量逐月减少;梭梭枯落期,降雨量最少。（2）在梭梭生长季,梭梭群落0~400 cm土壤贮水量变化整体呈下降趋势,梭梭萌发期是土壤贮水量盈余期,生长旺盛期和枯落期为土壤贮水量亏损期;梭梭群落发挥土壤水库效应,依靠生长季前土壤蓄水来弥补梭梭群落生长季需水缺额。（3）在梭梭生长季,蒸散量变化特征为多峰曲线,峰值主要出现在降雨集中期,最低值出现在土壤贮水量亏损期。（4）在梭梭生长季,梭梭群落累积蒸散量增幅始终高于累积降雨量增幅,累积蒸散量大于累积降雨量。     

Distribution of ephemeral plants and their significance in dune stabilization in Gurbantunggut Desert

Based on systematically monitoring plants on dune ridges in the southern part of the Gurbantunggut Desert in 2002, this paper, from the angle of dune stabilization by vegetation, describes the temporal and spatial distribution patterns of ephemeral plants on isolated sand dunes, analyses the natural invasion processes of ephemeral plants on human-disturbed sand surface and expounds the importance of ephemeral plants in stabilizing sand dune surface. A total of 45 plant species were identified in the study area, 29 of which are ephemeral plants. Ephemeral plants sprouted in early April and completed their life-circle within about two months. Just as aeolian sand activities came to the strongest stage from April to June in desert regions of northern Xinjiang, the total coverage of trees, shrubs and herbs of long vegetational period on most dune ridges was less than 10%, while the mean coverage of ephemeral plants reached 13.9% in April, 40.2% in May and 14.1% in June. Therefore ephemeral plants acted as the major contributor to dune surface stabilization in the Gurbantunggut Desert. Investigations of vegetation restoration on engineering-disturbed dune surface show that ephemeral plants first recolonized the disturbed dune surface.     

古尔班通古特沙漠短命植物分布及其沙面稳定意义

Based on systematically monitoring plants on dune ridges in the southern part of the Gurbantunggut Desert in 2002, this paper, from the angle of dune stabilization by vegetation,describes the temporal and spatial distribution patterns of ephemeral plants on isolated sand dunes,analyses the natural invasion processes of ephemeral plants on human-disturbed sand surface and expounds the importance of ephemeral plants in stabilizing sand dune surface. A total of 45 plant species were identified in the study area, 29 of which are ephemeral plants. Ephemeral plants sprouted in early April and completed their life-circle within about two months. Just as aeolian sand activities came to the strongest stage from April to June in desert regions of northern Xinjiang, the total coverage of trees, shrubs and herbs of long vegetational period on most dune ridges was less than 10%, while the mean coverage of ephemeral plants reached 13.9% in April, 40.2% in May and 14.1% in June. Therefore ephemeral plants acted as the major contributor to dune surface stabilization in the Gurbantunggut Desert.Investigations of vegetation restoration on engineering-disturbed dune surface show that ephemeral plants first recolonized the disturbed dune surface.     

古尔班通古特沙漠地区短命植物土壤种子库研究

采用野外定点取样和室内萌发相结合的试验方法,研究了短命植物在沙丘不同部位的空间分布、萌发状况、密度、物种多样性及土壤种子库生物多样性与地上植物多样性的关系。结果表明:短命植物种子主要集中分布于地表0~5 cm土壤中,5 cm以下土壤中极少存在有活力的种子;沙丘间及沙丘不同部位的种子库密度大小不同,密度自坡底随高度增加逐渐减少,坡底、坡中部及坡顶部密度分别为124粒/m2、91粒/m2、66粒/m2。迎风坡种子库密度比背风坡面少,迎风坡种子库平均密度为81粒/m2,而背风坡种子库平均密度为100粒/m2;短命植物种子在4月上旬达到萌发高峰,沙丘不同部位物种多样性存在差异,土壤种子库和地上植被存在较高的相似性,此外短命植物在沙丘不同部位的分布存在较高的空间异质性。     

古尔班通古特沙漠南缘不同立地条件植物多样性和优势种群生态位特征研究

以古尔班通古特沙漠南缘不同立地条件植物群落为研究对象,分析了该区不同立地条件植物群落的种类组成、优势种群及优势种群的生态位特征。结果表明:①不同立地条件的植物群落均为灌木-草本模式,优势种群为梭梭、白梭梭、白皮沙拐枣、羽毛针禾、骆驼刺、琵琶柴、狭果鹤虱、沙蓬、倒披针叶虫实;②由于不同立地条件环境因子差异,造成其植物群落的α多样性指数存在差异;③Whittacker指数与Cody指数显示不同立地条件植物群落β多样性随环境变化可分为两个梯度,第一梯度为垄顶、迎风坡与背风坡上部,第二梯度为平坦荒地、丘间平地与背风坡中、下部;④生态位宽度大的物种间生态位重叠大。在古尔班通古特沙漠南缘进行受损群落植被恢复与重建时,可在适宜的立地条件种植梭梭与羽毛针禾。     

古尔班通古特沙漠春季土壤含水量空间格局

春季积雪融水是古尔班通古特沙漠土壤水分的重要补给源之一,为早春植物的萌发和生长创造了有利的条件。为掌握该地区春季沙丘的土壤水分分布状况,采用探地雷达测量技术,于2010年4月进行了多次野外测量,并通过对地面直达波信号的提取和分析,初步获得了一致的测量结果。结果表明：（1）探地雷达地面直达波测量的土壤平均含水量与20 cm深同步TDR的测量结果非常接近,两者总体相差在0.02 cm3/cm3以内,充分反映了GPR地面直达波测量结果的有效性和精确性;（2）春季积雪完全融化后,沙丘/沙垄顶部土壤水分明显低于垄间低地,这与前人的研究一致,反映了积雪和冻土层在消融过程中,沙丘表面融水在坡面重力作用下,沿难以透水的冻土层上界自坡上向坡下迁移,形成了垄间最高、坡部次之和垄顶最少的分布格局;（3）垄间低地的土壤水分明显受到低矮灌草丛的影响进行再分布,这是由于积雪最初沿植丛基部开始消融,并以植丛为中心形成漏斗状融洞,从而形成以植丛为中心汇集较多融雪水的格局,而垄间低地中较为高大的梭梭乔木对融雪水的再分布影响明显不如低矮的灌草丛,这与其地表茎基过少有关。此外,探地雷达测量实验提供了详实可靠的表层土壤含水量分布信息,可为荒漠生态系统的空间尺度研究提供真实可靠的数据基础。     

古尔班通古特沙漠生物结皮小尺度分异的环境特征

古尔班通古特沙漠广泛存在的生物结皮,对地貌部位有极强的选择性分布,其实质是生物对环境条件综合适应的一种生态表现。2002年3~11月在个体沙丘表面初步开展了生物结皮类型小尺度环境分异规律研究。结果表明:苔藓结皮、地衣结皮和藻类结皮依次分布于垄间、沙垄两坡中部和坡上部,从垄间往垄顶,生物结皮盖度逐渐减小,厚度变薄,抗压性减弱。苔藓结皮分布区的物质组成以细沙和极细沙为主,春季表层土壤水分在5%以上,短命植物广泛发育,基质稳定;坡中部的地衣结皮分布区以细沙为主,春季表层土壤水分在4%左右,短命植物亦有广泛分布,地表受风沙活动影响较小;沙垄上部和顶部的藻类结皮分布区,是沙垄表面活动性最强和土壤水分最差的区域,物质组成以中沙为主,高等植物中白梭梭为优势种,可见藻结皮是三类结皮中最能适应恶劣环境的生物结皮类型。     

古尔班通古特沙漠沙丘主要灌木的种群数量动态

种群数量动态的研究是认识种群的生存现状及发展动态的前提。通过对古尔班通古特沙漠固定沙丘、半固定沙丘和流动沙丘上主要固沙灌木的样地调查,分析了固沙灌木的年龄结构,建立了3种沙丘上固沙灌木的静态生命表和存活曲线。结果表明：（1）固沙灌木的存活个体数呈逐级递减的趋势,幼龄个体占有很高的比例;固定沙丘上梭梭（Haloxylon ammodendron）呈现Ⅰ型存活曲线,种群退化,白梭梭（Haloxylon persicum）呈现Ⅱ型存活曲线,种群稳定;（2）半固定沙丘上白梭梭呈Ⅱ型存活曲线,种群相对稳定;沙拐枣（Calligonum mongolicum）和油蒿（Artemisia ordosica）均呈现Ⅰ型存活曲线,种群处于退化状态;（3）流动沙丘上沙拐枣呈现Ⅲ型存活曲线,种群有较大的增长潜力,白梭梭呈现出Ⅱ型存活曲线,种群相对稳定,油蒿呈现出Ⅰ型存活曲线,种群处于不稳定的退化状态。     

古尔班通古特沙漠南缘32种藜科植物种子萌发策略初探

对古尔班通古特沙漠南缘32种藜科植物的萌发特性（萌发率、萌发开始时间、平均萌发时间和萌发持续时间）进行研究,以探讨藜科植物的不同萌发对策及其生态适应意义。结果表明,32种植物萌发率呈双峰型,其中萌发率大于80%,或小于20%的种占大多数;萌发开始时间呈单峰型,28种植物在1~3 d内开始萌发;平均萌发时间也呈单峰型,平均萌发时间小于5 d的植物21种,最大的为杂配藜（23.7 d）;萌发持续时间种间差异较大,其中15种植物萌发持续时间小于15 d,10种植物持续时间在15~30 d,5种植物大于30 d。主成分分析和聚类分析结果表明,32种植物可划分为4种萌发类型:爆发型、过渡型、缓萌型和低萌型。以上结果揭示出32种藜科植物种子萌发策略不尽相同,表现为机会主义萌发策略、下注萌发策略和谨慎萌发策略,这与植物所处土壤盐渍化程度存在差异有关。此外,Log尺度下32种藜科植物种子大小及其变异范围较小（10-2~101）,暗示多数藜科植物多为小种子,其容易沿缝隙进入土壤形成短暂或永久种子库,这也是沙漠植物生存策略的重要一环。