A New Fossil Amiidae (Holostei: Halecomorphi) from the Upper Cretaceous Adamantina Formation,Southeastern Brazil,with comments on western Gondwana amiids

Amiid remains from the Upper Cretaceous (Turonian-Santonian) Adamantina Formation, Bauru Basin, Southeastern Brazil are described. The material is known only from fragmented isolated jaw bones (maxilla and dentary), a vertebral centrum, and isolated teeth and represents the first halecomorph described from this stratigraphical unit. Characters of the maxilla and the shape of the teeth, permit referral to the subfamily Vidalaminae. The Adamantina material differs from material described from the slightly younger Marília Formation (Maastrichthian) of the same Bauru Basin, by the presence of longitudinal ridges on the surface of the maxilla as well as by tooth sockets of equal size. These two records are the unique indisputable occurrences of freshwater vidalamins and confirm the presence of this clade within the incipient Neotropical ichthyofauna.     

Overview of the Late Cretaceous Biota of the western São Paulo State,Brazil, Bauru Group

The Turonian–Maastrichtian beds of the Bauru Group, western São Paulo State (Brazil), have yielded a diverse biota. The nine species of mesoecrocodilians are all mesosuchians. There is a single frog, a Neubatrachia. The vertebrates include, four types of fishes, a neobatrachian frog, an anilioid snake, two lizard, six species of podocnemid turtles, nine species of mesosuchian mesoecrocodilians three theropods, two birds, five titanosaurid sauropods, and one mammalian. The invertebrates include one cyclo, one cyclophoroid and eight pulmonates. Plants are represented by only two carophyte species. The biota from western São Paulo State is one the most diverse of the Late Cretaceous in Brazil. The biota from this area confirms that Bauru Group was connected to Patagonia and other Gondwanan areas during the Late Cretaceous.     

Biogeographical network analysis of Cretaceous Australian dinosaurs

The biogeography of Cretaceous Australian dinosaur fauna has been characterized variously as endemic, cosmopolitan or closely related to other Gondwanan faunas. Over the past decade, a large number of new Australian dinosaur taxa have been described and included in phylogenetic analyses, allowing for new insights into their biogeographical affinities to be obtained. Here, I combine the latest phylogenies of Australian dinosaurs into a supertree that includes as many Australian taxa as possible and apply it to the construction of biogeographical networks of Cretaceous dinosaurs. The results show that the Cretaceous dinosaurs of Australia have a strong connection with South American dinosaurs and to those of other Gondwanan continents via South America. Community detection algorithms indicate that the Gondwanan continents form a community within Cretaceous dinosaur biogeographical networks. The biogeographical affinities between Australia and Laurasia detected in previous studies were largely due to the scarce dinosaur fossil record of Gondwana at that time and to the unstable phylogenetic position of Australian dinosaurs because of the fragmentary nature of their fossils. Continuing discoveries of new Australian taxa and refinement of resulting phylogenetic analyses can further deepen our understanding of the biogeographical history of Australia.     

Cenomanian vertebrate assemblages from southwestern France: a new insight into the European mid-Cretaceous continental fauna

Cenomanian paralic deposits of Charentes (southwestern France) have yielded abundant vertebrate microremains, including rather diversified continental taxa (e.g., frogs, turtles, crocodilians, dinosaurs, pterosaurs, lizards, and mammals). In this short report, the succession of faunal assemblages observed is briefly described in relation to palaeoenvironmental change resulting from the Cenomanian-early Turonian transgressive episode. Continental forms occur commonly in estuarine and shallow marine rocks of the lower part of the Cenomanian stage, but seem to be lacking in younger strata deposited in more open marine settings. Among an unexpected biodiversity, several non-marine groups (e.g., ziphodont crocodilians, carcharodontosaurid and troodontid dinosaurs, marsupial-like mammals) are recorded for the first time in the lower Upper Cretaceous of Europe, thus providing new palaeobiogeographical indications for this poorly documented part of the world.     

Invertebrate traces in pseudo–coprolites from the upper Cretaceous Marília Formation (Bauru Group), Minas Gerais State,Brazil

Pseudo–coprolites are inorganic structures often confused with fossil faeces. The absence of some diagnostic features, such as inclusions, coprofabrics, grain adhesion, and defined shape, suffices to disregard these structures as coprolites. Herein we revise the so–called “coprolites” from the Serra da Galga Member of the Marília Formation (Maastrichtian of Bauru Group, Paraná Basin), at “Ponto 1 do Price” locality near the town of Peirópolis (Uberaba municipality, Minas Gerais State, Brazil) and conclude that they are, in fact, pseudo–coprolites related to calcretes. These data also agree with the geological setting of “Ponto 1 do Price”, composed mainly of coarse sandstones and conglomerates, in which these pseudo–coprolites were found. In addition, some of these specimens exhibit superficial traces, here described as a new boring ichnospecies, Asthenopodichnium fallax isp. nov., produced by invertebrates in Late Cretaceous fresh–water settings of Brazil.     

Morphology and internal structure of two new abelisaurid remains (Theropoda,Dinosauria) from the Adamantina Formation (Turonian – Maastrichtian), Bauru Group,Paraná Basin,Brazil

The knowledge on the Brazilian theropod fauna is hampered by the limited number of specimens unearthed so far. The most potential deposits for the finding of those dinosaurs are the layers of Bauru Group, which comprises several different formations of Late Cretaceous age. Most of those remains are referred to Abelisauroidea, a clade that is particularly well represented in Gondwana. Here we report two new abelisaurid specimens comprising a left ilium (DGM 927-R) and the distal articulation of a right femur (MCT 1857-R) unearthed from the outcrops of Adamantina Formation (Turonian – Maastrichtian) in the locality known as Santo Anastácio, that comprises an abandoned quarry located in the outskirts of the homonymous city in São Paulo State. The ilium has a preacetabular process horizontally oriented and ventrally deflected, as well as an acute angle between this process and the pubic peduncle. The femur shows a well-developed anteromedial crest, but not to the same degree as in noasaurids, and ossified ridges between condyles and tibiofibular crest. Those features allow us to assign both specimens to Abelisauridae. CT-scans of those specimens shows that the ilium has pneumaticities similar to the camellate pattern previously recorded in neosauropods and reveals the diverticula invasion of abdominal air-sacs. The femur presents a developed anisotropic trabecular system, which suggests that the main muscular strain is located at the distal portion of this bone. These features have not been previously reported in Abelisauridae, indicating that the internal organization of the postcranial skeleton in the group is rather complex and quite variable.     

Evolution and palaeoenvironment of the Bauru Basin (Upper Cretaceous,Brazil)

The Bauru Basin was one of the great Cretaceous desert basins of the world, evolved in arid zone called Southern Hot Arid Belt. Its paleobiological record consists mainly of dinosaurs, crocodiles and turtles. The Bauru Basin is an extensive region of the South American continent that includes parts of the southeast and south of Brazil, covering an area of 370,000 km². It is an interior continental basin that developed as a result of subsidence of the central-southern part of the South-American Platform during the Late Cretaceous (Coniacian–Maastrichtian). This sag basin is filled by a sandy siliciclastic sequence with a preserved maximum thickness of 480 m, deposited in semiarid to desert conditions. Its basement consists of volcanic rocks (mainly basalts) of the Lower Cretaceous (Hauterivian) Serra Geral basalt flows, of the Paraná-Etendeka Continental Flood Basalt Province. The sag basin was filled by an essentially siliciclastic psammitic sequence. In lithostratigraphic terms the sequence consists of the Caiuá and Bauru groups. The northern and northeastern edges of the basin provide a record of more proximal original deposits, such as associations of conglomeratic sand facies from alluvial fans, lakes, and intertwined distributary river systems. The progressive basin filling led to the burial of the basaltic substrate by extensive blanket sand sheets, associated with deposits of small dunes and small shallow lakes that retained mud (such as loess). Also in this intermediate context between the edges (more humid) and the interior (dry), wide sand sheet areas crossed by unconfined desert rivers (wadis) occurred. In the central axis of the elliptical basin a regional drainage system formed, flowing from northeast to southwest between the edges of the basin and the hot and dry inner periphery of the Caiuá desert (southwest). Life in the Bauru Basin flourished most in the areas with the greatest water availability, in which dinosaurs, crocodiles, turtles, fish, amphibians, molluscs, crustaceans, and charophyte algae lived. The fossil record mainly consists of transported bones and other skeletal fragments. In the northeastern and eastern marginal regions fossils are found in marginal alluvial fan deposits, broad plains of braided streams and ephemeral alkaline water lakes. In the basin interior the fossil record is related to deposits in sand sheets with braided streams, small dunes, and shallow lakes. In the great Caiuá inner desert a few smaller animals could survive (small reptiles and early mammals), sometimes leaving their footprints in dune foreset deposits. The aim of this article is to present and link the basin sedimentary evolution, palaeoecological features and palaeontological record.     

Taphonomy of a Baurusuchus (Crocodyliformes,Baurusuchidae) from the Adamantina Formation (Upper Cretaceous,Bauru Basin), Brazil: Implications for preservational modes,time resolution and paleoecology

Upper Cretaceous vertebrate accumulations from the Adamantina Formation are known due to their high taxonomic diversity. On the other hand, taphonomic analyses still are rare, limiting the understanding of processes related to the biostratinomic and fossildiagenetic histories of this lithostratigraphic unit. In 2005, fossils were collected from an outcrop located at Jales municipality, state of São Paulo, southeastern Brazil. From this outcrop, a well-preserved Baurusuchus was recovered, which displays a peculiar set of taphonomic signatures. This paper identifies and interprets taphonomic features of a specimen of Baurusuchus (Crocodyliformes, Baurusuchidae; UFRJ DG 418-R) from the Adamantina Formation (Upper Cretaceous of the Bauru Basin), in Jales municipality, state of São Paulo. Brief taphonomic comparisons with other specimens previously studied (crocodiles and dinosaurs) and a lithofaciological analysis of the outcrop were undertaken in order to enhance the knowledge of the stratigraphy and paleoenvironment and improve the time resolution for the Adamantina Formation in the region of Jales. Furthermore, paleoecological data were interpreted based on the taphonomic analysis.The fossil is composed of an articulated segment of vertebral column, ribs, part of the pelvic girdle and gastralia. There is no hydraulic equivalence between both the specimen of Baurusuchus and the size of quartz grain predominant in the fossiliferous layer, suggesting death in situ or short transport as a “water carcass”. Teeth marks identified on the pubes were assigned to a small/juvenile baurusuchid crocodyliform or a theropod dinosaur. The repositioning of some elements (ribs and dorsal osteoderms) is suggestive of mummification. Desiccation marks were observed and attributed to the stage 1 of weathering. These features suggest subaerial exposure of the carcass prior to burial, however, probably after the mummification. On the other hand, the subaerial exposure was short, because the individual was not fully disarticulated. Furthermore, the degrees of articulation and preservation of the specimen nullify the hypothesis of reworking.Lithofaciological and taphonomic analyses suggest a well-drained floodplain as the burial environment and an arid or semi-arid climate in the region of Jales outcrop. In addition, the taphonomic signatures seem to indicate a time resolution about 10⁰–10⁴ years for the layer where the crocodyliforms were collected, but it seems to have, within this time limit, temporal-mixing among terrestrial crocodiles and dinosaurs collected from the same layer, suggesting time-averaging in this outcrop.     

Description of a juvenile titanosaurian dinosaur from the Upper Cretaceous of Brazil

We describe a juvenile specimen of a titanosaurian sauropod dinosaur consisting of two dorsal and three caudal vertebral centra, an ilium fragment, and an ischium unearthed in 1991 from Site Km 153.5 at BR-050 highway in the Serra da Galga region, municipality of Uberaba, state of Minas Gerais, Brazil. The outcrop at the site is assigned to the Serra da Galga Member of the Marília Formation (Bauru Basin: Bauru Group; Upper Cretaceous: Maastrichtian). Although the material is very incomplete, features such as strongly procoelous caudal centra suggest an affinity with the titanosaurian clade Lithostrotia. The extensive vertebral pneumaticity with deep pleurocoels and well-developed camerae supports the hypothesis that, in titanosaurs, the air sac system was already present and fully developed even at early ontogenetic stages.     

浙江白垩纪恐龙与恐龙蛋化石的时空分布

浙江省恐龙化石地质遗迹调查表明,全省有9个白垩纪陆相红层盆地产恐龙和恐龙蛋化石,至今已发现有4科5属5种恐龙骨骼和7科11属24种恐龙蛋化石。浙江省恐龙和恐龙蛋化石主要分布在天台盆地和金衢盆地的东阳、衢州、龙游、金华、浦江、兰溪一带,主要产于天台群的赖家组,衢江群的金华组、衢县组。永康、南马、新昌、嵊州、仙居、诸暨、湖山等地永康群朝川组、方岩组中也有较丰富的恐龙化石。依据恐龙和恐龙蛋化石在盆地内地层中的分布、地层的同位素年龄和相伴生的其他生物群特征等分析表明,恐龙和恐龙蛋化石的赋存地质时代以晚白垩世为主。此外,恐龙和恐龙蛋化石在浙江的广泛分布表明该地区地质历史时期气候温暖,植被茂盛,水资源丰富,是恐龙及其他生物生存繁衍的理想场所。     

Osteoderms of Montealtosuchus arrudacamposi (Crocodyliformes,Peirosauridae) from the Turonian-Santonian (Upper Cretaceous) of Bauru Basin,Brazil

The skin of Crocodyliformes is characterized by osteoderms of various sizes and shapes. It plays roles in defence, thermoregulation, sexual attraction, calcium reserves, and locomotion. This study presents the morphometric characteristics of osteoderms preserved in the nuchal, dorsal, ventral and appendicular shield of Montealtosuchus arrudacamposi, a Peirosauridae crocodyliform from the Turonian-Santonian (Upper Cretaceous) of Adamantina Formation, Bauru Basin, Brazil. The results of the analysis show that its dermal shield had protective and thermoregulatory functions and a well-defined mechanical function allowing terrestrial locomotion and enabling good agility and capacity for movement.     

中国东北中生代地层划分对比之新见

在总结近二十年来有关我国东北地区中生代地层研究诸多新发现与新进展的基础上 ,以生物化石为主要依据 ,以 "海相检验法”及国际性生物地层对比为标准 ,结合同位素年代地层学新资料 ,对东北地区中生代地层的划分及对比 ,提出了新的方案。文中还讨论了三叠系 -侏罗系界线、侏罗系 -白垩系界线及下白垩统 -上白垩统界线的划分以及辽西义县组的时代。同时 ,还首次报道了笔者等对黑龙江东部龙爪沟群研究的最新进展 ,并提出了对龙爪沟群时代及与鸡西群对比的新认识。     

The turtles from the uppermost Jurassic and Early Cretaceous of Galve (Iberian Range,Spain): Anatomical,systematic, biostratigraphic and palaeobiogeographical implications

The faunal associations of the uppermost Jurassic and Early Cretaceous Villar del Arzobispo, El Castellar, and Camarillas formations of the Spanish town of Galve (Maestrazgo Basin of the Iberian Range) are composed of more than fifty taxa belonging to Chondrichthyes, bony fishes, lissamphibians, mammals, and reptiles. Although the presence of turtles has been recognized in these three formations, being very abundant, the published information is limited. The detailed study of these turtles allows us to recognize at least nine taxa, several of them previously not identified in the Spanish record. These taxa correspond to stem Testudines (Solemydidae), members of Paracryptodira (Pleurosternidae), several representatives of Plesiochelyidae and taxa closely related with this clade, representatives of the stem group of Cryptodira (Xinjiangchelyidae) and members of crown Cryptodira. The chelonian fauna of Galve increases the systematic, anatomical, biostratigraphic, and paleobiogeographic knowledge on several clades, represented by taxa shared with other European regions, clades also present in Early Cretaceous sites of North America or Asia, as well as taxa exclusive of the Iberian record.     

浙江恐龙和蛋化石的时代

在浙江省内,有12个白垩纪红层盆地中发现恐龙化石和蛋化石,且多个盆地龙蛋共生。经初步调查,除金衢盆地衢江群下部中戴组发现恐龙化石外,恐龙化石和蛋化石均产于盆地内的中上部地层:即永康群朝川组、方岩组,衢江群的金华组和衢县组,天台群两头塘组和赤城山组。从相关地层的古生物组合时代和火山岩同位素年龄为101～90Ma以及地层的古地磁均为正向极性特征分析,浙江绝大部分恐龙蛋的时代应为晚白垩世,而蜂窝蛋类化石时代则很可能是早白垩世晚期或未期。     

Evolution of the carnivorous dinosaurs during the Cretaceous: The evidence from Patagonia

Patagonia has yielded the most comprehensive fossil record of Cretaceous theropods from Gondwana, consisting of 31 nominal species belonging to singleton taxa and six families: Abelisauridae, Noasauridae, Carcharodontosauridae, Megaraptoridae nov. fam., Alvarezsauridae, and Unenlagiidae. They provide anatomical information that allows improved interpretation of theropods discovered in other regions of Gondwana. Abelisauroids are the best represented theropods in Patagonia. They underwent an evolutionary radiation documented from the Early Cretaceous through to the latest Cretaceous, and are represented by the clades Abelisauridae and Noasauridae. Patagonian carcharodontosaurids are known from three taxa (Tyrannotitan, Giganotosaurus and Mapusaurus), as well as from isolated teeth, collected from Aptian to Cenomanian beds. These allosauroids constituted the top predators during the mid-Cretaceous, during which gigantic titanosaur sauropods were the largest herbivores. Megaraptorans have become better documented in recent years with the discovery of more complete remains. Megaraptor, Aerosteon and Orkoraptor have been described from Cretaceous beds from Argentina, and these taxa exhibit close relationships with the Aptian genera Australovenator, from Australia, and Fukuiraptor, from Japan. The Gondwanan megaraptorans are gathered into the new family Megaraptoridae, and the Asiatic Fukuiraptor is recovered as the immediate sister taxon of this clade. Although megaraptorans have been recently interpreted as members of Allosauroidea, we present evidence that they are deeply nested within Coelurosauria. Moreover, anatomical information supports Megaraptora as more closely related to the Asiamerican Tyrannosauridae than thought. Megaraptorans improve our knowledge about the scarcely documented basal radiation of Gondwanan coelurosaurs and tyrannosauroids as a whole. Information at hand indicates that South America was a cradle for the evolutionary radiation for different coelurosaurian lineages, including some basal forms (e.g., Bicentenaria, Aniksosaurus), megaraptorans, alvarezsaurids less derived than those of Laurasia, and unenlagiids, revealing that Gondwanan coelurosaurs played sharply differing ecological roles, and that they were taxonomically as diverse as in the northern continents. The unenlagiids represent an endemic South American clade that has been recently found to be more closely related to birds than to dromaeosaurid theropods. Analysis of the theropod fossil record from Gondwana shows the highest peak of origination index occurred during the Aptian–Albian and a less intense one in the Campanian time spans. Additionally, peaks of extinction index are recognized for the Cenomanian and Turonian–Coniacian time spans. In comparison, the Laurasian pattern differs from that of Gondwana in the presence of an older extinction event during the Aptian–Albian time-span and a high origination rate during the Cenomanian time-bin. Both Laurasian and Gondwanan theropod records show a peak of origination rates during the Campanian.     

Gondwanasuchus scabrosus gen. et sp. nov., a new terrestrial predatory crocodyliform (Mesoeucrocodylia: Baurusuchidae) from the Late Cretaceous Bauru Basin of Brazil

Baurusuchids are among the most common and diverse crocodyliform fossils from the Late Cretaceous Bauru Basin of Brazil. This inland continental basin was the habitat of a rich crocodyliform fauna containing five mesoeucrocodylian families, of which the Baurusuchidae represents highly specialized predatory crocodyliforms of terrestrial habits as indicated by their dental, cranial, and postcranial features. The large size they achieved, together with likely predatory adaptations, would suggest they competed and occupied theropod ecological niches in the Bauru Basin. Here we describe Gondwanasuchus scabrosus gen. et sp. nov., a medium-sized baurusuchid with a strongly laterally compressed skull, bearing unique dentition with deep apicobasal sulci and probably well-developed binocular vision. The cranial and dental features in Gondwanasuchus suggest that this active predator would have fed on small vertebrates and took the role of small theropods in terrestrial guild. Gondwanasuchus is the most distinctive baurusuchid known to date and enriches the knowledge on these important Gondwanan terrestrial predatory crocodyliforms.     

U–Pb geochronology of the southern Brasília belt (SE-Brazil): sedimentary provenance, Neoproterozoic orogeny and assembly of West Gondwana

The Brasília belt borders the western margin of the São Francisco Craton and records the history of ocean opening and closing related to the formation of West Gondwana. This study reports new U–Pb data from the southern sector of the belt in order to provide temporal limits for the deposition and ages of provenance of sediments accumulated in passive margin successions around the south and southwestern margins of the São Francisco Craton, and date the orogenic events leading to the amalgamation of West Gondwana.Ages of detrital zircons (by ID–TIMS and LA-MC-ICPMS) were obtained from metasedimentary units of the passive margin of the São Francisco Craton from the main tectonic domains of the belt: the internal allochthons (Araxá Group in the Áraxá and Passos Nappes), the external allochthons (Canastra Group, Serra da Boa Esperança Metasedimentary Sequence and Andrelândia Group) and the autochthonous or Cratonic Domain (Andrelândia Group). The patterns of provenance ages for these units are uniform and are characterised as follows: Archean–Paleoproterozoic ages (3.4–3.3, 3.1–2.7, and 2.5–2.4 Ga); Paleoproterozoic ages attributed to the Transamazonian event (2.3–1.9 Ga, with a peak at ca. 2.15 Ga) and to the ca. 1.75 Ga Espinhaço rifting of the São Francisco Craton; ages between 1.6 and 1.2 Ga, with a peak at 1.3 Ga, revealing an unexpected variety of Mesoproterozoic sources, still undetected in the São Francisco Craton; and ages between 0.9 and 1.0 Ga related to the rifting event that led to the individualisation of the São Francisco paleo-continent and formation of its passive margins. An amphibolite intercalation in the Araxá Group yields a rutile age of ca. 0.9 Ga and documents the occurrence of mafic magmatism coeval with sedimentation in the marginal basin.Detrital zircons from the autochthonous and parautochthonous Andrelândia Group, deposited on the southern margin of the São Francisco Craton, yielded a provenance pattern similar to that of the allochthonous units. This result implies that 1.6–1.2 Ga source rocks must be present in the São Francisco Craton. They could be located either in the cratonic area, which is mostly covered by the Neoproterozoic epicontinental deposits of the Bambuí Group, or in the outer paleo-continental margin, buried under the allochthonous units of the Brasília belt.Crustal melting and generation of syntectonic crustal granites and migmatisation at ca. 630 Ma mark the orogenic event that started with westward subduction of the São Francisco plate and ended with continental collision against the Paraná block (and Goiás terrane). Continuing collision led to the exhumation and cooling of the Araxá and Passos metamorphic nappes, as indicated by monazite ages of ca. 605 Ma and mark the final stages of tectonometamorphic activity in the southern Brasília belt.Whilst continent–continent collision was proceeding on the western margin of the São Francisco Craton along the southern Brasília belt, eastward subduction in the East was generating the 634–599 Ma Rio Negro magmatic arc which collided with the eastern São Francisco margin at 595–560 Ma, much later than in the Brasília belt. Thus, the tectonic effects of the Ribeira belt reached the southernmost sector of the Brasília belt creating a zone of superposition. The thermal front of this event affected the proximal Andrelândia Group at ca. 588 Ma, as indicated by monazite age.The participation of the Amazonian craton in the assembly of western Gondwana occurred at 545–500 Ma in the Paraguay belt and ca. 500 Ma in the Araguaia belt. This, together with the results presented in this work lead to the conclusion that the collision between the Paraná block and Goiás terrane with the São Francisco Craton along the Brasília belt preceded the accretion of the Amazonian craton by 50–100 million years.     

Adamantinasuchus navae: A new Gondwanan Crocodylomorpha (Mesoeucrocodylia) from the Late Cretaceous of Brazil

A new Late Cretaceous Mesoeucrocodylian from the Adamantina Formation (Bauru Basin), São Paulo State, Brazil is described. The main features of this new species are the short, high oreinirostral rostrum, the large laterally positioned orbital notches and external nares in the anteriormost portion of the rostrum. The mandible is robust and concave-shaped in relation to the skull. The dentition is highly specialized, with two prominent incisiform teeth, a hypertrophied caniniform, and seven molariform teeth. The molariform teeth are ornamented with denticles in their lingual surface and are smooth on their labial surface. The molariforms are elliptical in cross-section, presenting the largest axis in the labial-lingual direction. Such dental characteristics are unique among the terrestrial crocodylomorphs of the Gondwana.     

A review of the upper Campanian vertebrate site of Armuña (Segovia Province,Spain)

The Upper Cretaceous outcrops of Armuña (Segovia Province, Spain) yielded relatively abundant material of vertebrates during prospection and excavation in the second half of the 1980s. However, little has been published on these remains. A new analysis of the specimens from this upper Campanian site reveals the presence of some clades in the site for the first time (e.g., Dortokidae, Anguimorpha, Mosasauroidea). Furthermore, the material of the clades previously recognized there has been reviewed and described in more detail, with some previous systematic attributions confirmed and others refuted. Consequently, a relatively high local diversity has been identified. New taxa (i.e., a member of Anguimorpha and a eusuchian crocodyliform) are identified in Armuña, coexisting with other taxa previously described in other sites from the Iberoarmorican Realm. The vertebrates from Armuña confirm that the fauna from the Upper Cretaceous of the Iberian Peninsula is composed of a mixture of European endemic clades and lineages shared with other continents such as North America (e.g., anguimorphs) and Africa (e.g., bothremydids).